SPECIES 384 BLUE-WINGED WARBLER Vermivora pinus The Blue-winged Warbler is usually recognized by its buzzy "bee-buzz" songs. This attractive warbler with yellow head and body and bluish wings is found through the southern half of the LP and locally in the northern part of the LP. The breeding range for the species includes most of North America east of the Great Plains, including the northern states and extreme southern Ontario, and the southern states as far south as northern Georgia. Its wintering range extends from Mexico and Central America south to Panama (AOU 1983). HABITAT The Blue-winged Warbler breeds in open shrubby habitats, woodland openings, edges of streams, wil low swamps, and old fields with shrubs and small deciduous trees. Vegetation commonly consists of aspen, willow, maple, hawthorn, black locust, gray dogwood, and alder buckthorn, in early stages of succession. The habitat often includes low, dense shrubs bordered by an open grassy area (Ficken and Ficken 1968a, 1968b, Gill and Murray 1972a, Ewert 1981). The nest is built on the ground under a tussock of grass. SEASONAL OCCURRENCE Blue-winged Warblers are summer residents in Michigan. They arrive from early to mid May. Males are most conspicuous while they sing during the second half of May and early June, and they become more quiet when nesting is finished in June. Nesting begins with egg laying dates around 28 May, and the species has only one brood in a season (Will 1986). Most individuals leave for their wintering grounds around the middle of August (Wood 1951). STATUS Blue-winged Warblers are now common in the southern third of Michigan but were not known in the state during the middle 1800s. The first record was in Kalamazoo in 1879; the first pair, suggesting breeding, was seen in Wayne Co. in 1902. A Blue-winged Warbler was seen in 1896 in Ann Arbor, Washtenaw Co., but they were not regular there until 20 years later (Barrows 1912, Wood 1951, W. Koelz pers. comm.). As in the eastern part of their range in North America, Blue-winged Warblers in Michigan have expanded northward during the past 80 years and the range ACCOUNTS extension has coincided with a northward shift of Goldenwinged Warblers (Gill 1980). By the 1930s Blue-wings were fairly common in south western Michigan and throughout the southern three tiers of counties. By the 1950s they were more common than the Golden-winged Warbler in southwestern Michigan (Wood 1951, Berger 1958), and by the late 1960s they were more common in southeastern Michigan (Gill and Murray 1972a, 1972b, Murray and Gill 1976). In Isabella Co., halfway up the LP, the species appeared in 1979 and 1980 in sapling aspen and paper birch where Golden-winged Warblers were also seen (Ewert 1981). The Atlas observations provide good documentation of this warbler's present-day distribution, and are consistent with a continuing expansion in Michigan. Blue-wings are now com mon in much of the southern LP. Overall, they are more com mon in the southwestern part of the state. They were found in every township in Barry and Kalamazoo counties. In the southeast, they breed in several areas in Livingston, Oakland, Lapeer, St. Clair, and Tuscola counties. The northernmost breeding, in Benzie Co., involved a hybrid "Brewster's Warbler." The well-concealed nests are difficult to find. Most confirmed breeding records were of adults attending young and fledglings after they had departed the nest. Difficulties in censusing the species include the similarity of the songs of Blue-winged and Golden-winged warblers, though usually the songs are distinct. Some individuals of each species give the song of the other species (Ficken and Ficken 1968b, 1968c, 1969, Gill and Murray 1972a, 1972b). Also, the two sometimes associate in mixed pairs, interbreed, and produce hybrid offspring. "Brewster's Warbler," once named as a distinct species, is the most common form of hybrid, the male white on the throat and belly, and the breast, wing bars, and cap bright yellow. This hybrid was observed in 27 Atlas blocks. "Lawrence's Warbler," the hybrid form with yellow underparts and a black throat and face pattern, is less common and was seen in 8 Atlas blocks. Both hybrid forms were observed throughout the range of overlap of Blue-winged and Goldenwinged warblers in the Atlas observations. One "Brewster's" was found in Dickinson Co. in the UP, observed with a Golden-winged Warbler. Two color-banded male "Brewster's Warblers" survived for four years in the E. S. George Reserve in Livingston Co. One mated with a female Blue-winged Warbler; the other mated with a female Golden-winged Warbler and fledged a brood of six young. In each case the female mated with a male "Brewster's Warbler" that sang the song of her own species, and in each case the fledged brood were yellow in juvenile plumage, like young Blue-winged Warblers (orig. obs.). Hybrids may have the song of either parent species, perhaps depending on which species was the father. Females of both species of warblers tend to mate with a like male, but where they both occur in southern Michigan, the inci dence of mixed matings and hybridization is about 20% (Gill and Murray 1972a). Playback of tape-recorded songs usually attracts the singing male into sight (Ficken and Ficken 1969, Gill and Murray 1972b). The species is more likely to respond selectively to song of its own species in an area where the two have lived together for many years (Crook 1984). Because the song may not match the plumage identity of the singing bird, and its mate may not be the same species as the sing ing male, censusing efforts should include visual identifica tion of the warbler and its mate. Robert B. Payne BLUE-WINGED WARBLER BREEDING EVIDENCE Number of townships reported ■ Possible • Probable ■ Confirmed REGION TOTALS UP NLP SLP 0 0 37 171 TOTALS 208 16 116 132 0 4 57 61 0 57 344 401 Percentage of total townships 0.0 (of 610) BBS Abundance Number of birds per route 0 40 Kilometers 80 9.7 (of 590) 49.4 (of 696) 21.1 (of 1896)