Module 2 THE ECOLOGICAL HISTORY OF THE MELVILLE KOPPIES NATURE RESERVE Although Melville Koppies is a tiny reserve in the heart of a city, it presents an extraordinary variety of habitats generated by its topology and geology. Consisting of three east-west quartzite, fissured, ridges, dipping southwards, separated by shallow shale-based valleys and all resting on ancient granite and greenstone it has an interesting wlidlife that has survived the vicissitudes of its history. There are specific fauna and flora communities associated with the different habitats,- (1) the ridge summits, (2) the thin acid soils on the exposed southern slopes, (3) the deeper and slightly less acid soils of the more sheltered intervening valleys, (4) the sandy, wetter, but well-drained alluvial soils in the southward oriented valleys in geological faults, (5) the warmer, drier, rocky, quartzite-based soils on the talus below the northernmost ridges, (6) the drainage basins in northern-oriented faults, and further out the richer granite-based soils of the northern slopes and even here there is a perceptible difference between the communities on (7) ancient granite and on (8) greenstone-based soils, (9) a riverine community along the Westdene Spruit and finally (10) a moist bush above the Old Road. These natural differences have been compounded by man. As there is little iron in the Melville Koppies rocks, there must have been extensive woodland (possibly similar to that on the southern slopes of the Magaliesberg and the intervening Bankenveld) on the northern slopes of the ridge, along the Westdene and Braamfontein Spruits and in the valleys of Melville to have attracted an extensive Iron Age industry. The iron makers would probably have taken out all the large trees as appears to have happened around the Great Lakes centuries before.This process would have continued through grazing, burning and the gathering of firewood and plants for gardens or muti when the area was a farm and later when it was simply open space. Even since it has been protected different parts have been subjected to varied frequencies of burning. Man has a long history of contact with the area. Up to 1000 years ago the reserve was a camping ground for San family groups. They were displaced by permanent settlements of Iron Age farmers needing fresh pasture for their cattle and wood for their furnaces. At the beginning of the 19 th century the area was visited with increasing frequency by hunter/explorers who have left descriptions of the herds of elephants and other game on the rolling grasslands stretching to the north. In the 1860’s Melville Koppies became farmland and in the 1880’s was prospected for gold The encircling of Melville Koppies began early in the 1900’s when the southern and and eastern parts of the farm-now Melville and Greenside- became small-holdings. Melville became a township in 1900 , Greenside in 1937 Emmarentia in 1939 and Emmarentia Extention which included the most eastern part of the ridge, in 1945.The City Council bought what was left of the the western half of the farm in 1932.In the 1960’s the Linden Road which bisected the ridge was replaced in the 1960’s by the larger and very busy D.F.Malan Drive, the kloof through which runs the Westdene Spruit was extensively damaged by the laying of sewage and stormwater drains and an adjacent wetland was filled in to make a sportsfield for RAU. For 30 years the ridge was completely neglected. It was pillaged for firewood, stone, soil and plants, used for dumping, the wildlife was poached or hunted by cats and dogs and alien plants were allowed to establish themselves. Older Melville residents rember dassies and a troop of baboons on the ridge and vultures on Northcliff, all of which have long disappeared. Aerial photographs of the ridge in 1960 show only a sprinkling of bush along the ridge. In 1959 the part of the ridge between Emmarentia township and D.F.Malan Drive was proclaimed a nature reserve. It was fenced, policed by security guards, provided with 2 waterpoints, paths, and essential buildings by the Council. The Johannesburg Council for Natural History, formed to advise the Council, had inventories made of the fauna and flora, removed alien vegetation, constructed a nature trail, published a handbook for it and encouraged publications on the reserve. Management policy was the preservation of the ridge, fires were as far as possible excluded and entry was restricted.After some 25 years the Council found it increasinglydifficult to fund the maintenance of its nature reserves. The guards were withdrawn and another period of neglect followed. In 1992 the Council asked local communities to take over the management of its nature reserves and the present management committee was formed. This is reelected annually at a public meeting. Current management policy is to make the reserve as useful to the community as far as is possible without affecting the intrinsic nature of the reserve and to develop community support to the point where it does not need Council funding. Policy changes were ;-(a) as the exclusion of fire had led to bush encroachment planned veld-burning was started, and (b) to increase awareness of Melville Koppies , entry, although still restricted, was made easier, Open Days were increased to 3 and widely publicised. Another change was the addition of the still undeveloped parts of the ridge east and west to the nature reserve . In their case entry has remained unrestricted even though it results in frequent unplanned fires and damage and littering by vagrants. Since the 1950’s the ridge has been increasingly isolated from open country. The only immigration corridor left is along the Braamfontein Spruit which is steadily becoming longer and narrower. as Sandton develops northwards. An even more serious barrier is the replacement of suburban gardens by high density housing surrounded by high walls. Thus replenishment is becoming increasingly difficult for creatures that cannot fly or pass undetected through suburban gardens and a population collapse of any species can mean extinction within the reserve. On the other hand the afforestation of the northern suburbs has facilitated the immigration of other kinds of forest and bush wildlife which has produced a change in the population composition of the reserve,patricularly of the birdlife. The flora has also been affected by encirclement by suburban gardens. The seeds of alien plants are constantly introduced by birds and wind and dumping and pose a very serious threat to the indigenous vegetation. Other detrimental effects of contact with man are the destruction caused by vagrants camping in the reserve, vandalism, littering and erosion on heavily used paths.. Fire has had an important role in shaping Melville Koppies. For thousands of years the northern grassland of the Closed Section as part of the highveld grassland would have experienced lightning generated fires that would have burnt up to the forest margins. The Iron Age farmers who arrived 1000 years ago burnt the dry grass annually to bring on fresh green grass for their cattle and the felling of the forests by the iron makers. brought grassland still closer to the ridges, a practice that was continued by the later white farmers. As the city grew fires in the northern grassland became rarer then virtually ceased. Now a small part is burnt every 4 or 5 years as part of a planned programme to preserve biodiversity. This is not the case in the grassland on the southern sides of the ridges as the LGVS, being open to the public, is a source of fires caused by carelessnes or vandalism. These fires burn across the back of the reserve sometimes as far as D F Malan Drive.They seldom burn into the bush on the northern slopes due to the updraught behind the steep rock faces. The part of the ridge west of D F Malan Drive has again a different experience as it is burnt from end to end every year by carelessnes, vandals or deliberately and fires started on the northern slopes burn up into the bush which is noticeably thinner than that in the Closed Section.