Electronic Supplementary Material 6 (A). Relationships between descriptors of song complexity and environmental, social and community
variables GLMMs were performed, and transect and site identity were entered as random factors (transects nested in sites). Only those models
separated by less than 4 AIC points from the model with the lowest AIC are shown. ·P<0.1, *P<0.5, **P<0.01, ***P<0.001
Species and variable
Model
Z values
AIC
Crested lark
Mimicry diversity Habitat diversity + Irrigated land cover
(2.3)*+(-2.1)*+(2.6)**+(3.0)** +
(Landscape)+α species richness+ γ species
(-2.7)**+(1.4)+(3.2)**
richness+ α species richness× γ species richness+
Number of transects+ Number of strophes recorded
Habitat diversity + α species richness+ γ species
(1.4)+(1.7)· +(2.1)*+
richness+ α species richness× γ species richness+
(-1.8)·+(-1.0)+(3.1)**
Number of transects+ Number of strophes recorded
Mimicry prevalence α species richness+ γ species richness+ α species
(1.7)·+(1.8)·+(-1.7)·+
richness× γ species richness+ Number of transects+ (-1.9)·+(-0.4)
Number of strophes recorded
Landscape diversity + α species richness+ γ species (-0.9)·+ (1.8)·+(2.0)* +
richness+ α species richness× γ species richness+
(-1.9)·+(-1.8)·+(-0.3)
Number of transects+ Number of strophes recorded
Overall song complexity Irrigated land cover (Landscape)+ Habitat
diversity+α species richness+ γ species richness+ α
species richness×γ species richness+ Number of
transects+ Number of strophes recorded
(-0.8)+(1.1)+(2.0)*+(1.6)· +
(-1.9)· +(-0.4)+ (0.8)
Irrigated land cover (Landscape)+ Agricultural land (-1.5)+(-2.5)*+(-1.7)+
cover (Habitat)+Natural vegetation
(0.6)+(2.1)*·+(1.8)·+
(Habitat)+Habitat diversity+α species richness+ γ
(-2.0)*+(-0.1)+ (0.8)
species richness+ α species richness×γ species
richness+ Number of transects+ Number of
strophes recorded
Homospecific song complexity Fallowland cover (Habitat)
(2.4)*
159.1
160.6
73.9
76.5
416.4
419.4
388.6
1
CV duration Irrigated land cover+ Agricultural mosaic cover
(Landscape)+Crested lark density
CV mimimum frequency Crested lark density+ Habitat diversity
Crested lark density
CV frequency range Crested lark density+ γ species richness+ Natural
vegetation cover (Habitat)+Landscape diversity
(-2.0)·+(-2.1)*+(2.3)*
-41.2
(3.6)***+(-2.2)*
-143.6
(2.8)**
-146.2
(1.9)·+(2.9)**+(-2.5)**+(-1.3)
-92.2
(2.6)**·+(2.2)*·+(1.7)·+(2.0)*+(1.0)
+(-2.1)*+(-0.7)+(4.0)***
122.7
Thekla lark
Mimicry diversity Irrigated land cover (Landscape)+Steppeland cover
(Landscape)+Agricultural mosaic cover
(Landscape)+ α species richness+ γ species
richness+ α species richness× γ species richness+
+Number of transects+ Number of strophes
Mimicry prevalence Fallowland cover (Habitat)+γ species
(3.8)***·+(2.7)**+(-1.6)·+(3.2)***
richness+Number of transects+ Number of strophes
Fallowland cover (Habitat)+ α species
(2.8)**·+(1.3)+(-0.1)·+(2.8)**
richness+Number of transects+ Number of strophes
35.1
38.6
Overall song complexity No significant relationship
Homospecific song complexity Habitat diversity + γ species richness+ Thekla lark
density+ Thekla lark density× γ species
richness+Number of transects
CV duration γ species richness+ + Thekla lark density+ Thekla
lark density× γ species richness+Number of
transects
(-1.7)·+(0.7)+(2.0)*+(-2.4)*+(-0.1)
(2.0)*·+(1.7)·+(1.7)·+(-1.6)·
CV mimimum frequency No significant relationship
CV frequency range No significant relationship
2
314.4
48.9
Electronic Supplementary Material 6 (B). Relationships between Crested and Thekla lark densities in transects and environmental and
community variables. GLMMs were performed, and site identity was entered as a random factor. Only those models separated by less than 4 AIC
points from the model with the lowest AIC are shown. ·P<0.1, *P<0.5, **P<0.01, ***P<0.001
Variable
Crested lark
density
Thekla lark
density
Model
Z-values
AIC
Landscape diversity+ Habitat diversity+
α species richness
(2.2)*+(2.7)**+(2.7)**
276.1
Landscape diversity+ Habitat diversity
(1.9)·+(3.3)***
278.3
Habitat diversity+ Landscape diversity+ α species
richness+ α species richness2
(-5.1)***+(0.4) +
(3.4)**+(-3.7)***
245.5
Habitat diversity+ α species richness+ α species
richness2
(-5.2)***+(3.5)**+
(--3.7)***
246.9
3