References
Arthur, DK., 1976. Food and feeding of larvae of three fishes occurring in the California Current,
Sardinops sagax, Engraulis mordax, and Tachurus symmetricus. Fish. Bull 74:517-30.
Barnes, J.T., L.D. Jacobsen, A.D. MacCall, and P. Wolf, 1992. Recent population trends and
abundance estimates for the Pacific sardine (Sardinops sagax). CalCOFI Reports, 33:60-75.
Baumgartner, T. R., A. Soutar, and V. Ferreira, 1992. Reconstruction of the history of Pacific
sardine and Northern anchovy populations over the past 2 millenia from sediments of the
Santa Barbara Basin, California. Calif. Coop. Oceanic Fish Invest. Rep., 33:24-40.
Baumgartner T. R., G. Auad, and A. J. Miller, 2007. Regulation of sardine productivity in the
California Current. In preparation for Progress in Oceanography.
Butler, J., Fuller, D. and Yaremko, M., 1999. Age and growth of market squid (Loligo
opalescens) off California during 1998. Calif. Coop. Oceanic Fish Invest. Rep. 40, 191-195.
Borcard, D.; Legendre, P., 2002. All-scale spatial analysis of ecological data by means of
principal coordinates of neighbour matrices. Ecological Modeling 153: 51-68.
Borcard, D.; Legendre, P.; Avois-Jacquet, C.; Tuomisto, H.. 2004. Dissecting the spatial
structure of ecological data at multiple scales. Ecology 85(7): 1826-1832.
Chavez, FP, J Ryan, SE Lluch-Cota, and M Ñiquen. 2003. From anchovies to sardines and back:
multidecadal change in the Pacific Ocean. Science 299:217-21.
Chua, B.S., and A.F. Bennett, 2001. An inverse ocean modeling system, Ocean Modeling, 3,
137-165.
Di Lorenzo, E., A. J. Miller, D. J. Neilson, B. D. Cornuelle and J. R. Moisan, 2004: Modeling
observed California Current mesoscale eddies and the ecosystem response. International
Journal of Remote Sensing, 25, 1307-1312.
Di Lorenzo, E., A. M. Moore, H. Arango, B. Chua, B. D. Cornuelle, A. J. Miller and A. Bennett,
2007. Ocean Modelling, 16, 160-187.
Fleminger, A, Isaacs, J.D., Wyllie, J.G., 1974. Zooplankton biomass measurement from
CalCOFI cruises of July 1955 to 1959. CalCOFI Atlas Number 21, State of California
Marine Research Committee, 118pp.
Foote, Kenneth G., Roger T. Hanlon, Pat J. Iampietro, Rikk G. Kvitek., 2006. Acoustic detection
and quantification of benthic egg beds of the squid Loligo opalescens in Monterey Bay,
California. J. Acoust. Soc. Am. 119: 844-856.
1
Francis, R.I.C.C., 2006. Measuring the strength of environment-recruitment relationships: the
importance of including predictor screening within cross-validations. ICES Journal of
Marine Science 63: 594-599 .
Haidvogel, D.B., H. Arango, K. Hedstrom, A. Beckmann, P. Rizzoli and A. Shchepetkin, 2000.
Model evaluation experiments in the North Atlantic Basin: Simulations in non-linear terrainfollowing coordinates, Dyn. Atm. Oceans, 32, 239-281.
Haidvogel, D. B., H. Arango, W. P. Budgell, B. D. Cornuelle, E. Curchitser, E. Di Lorenzo, K.
Fennel, W. R. Geyer, A. J. Hermann, L. Lanerolle, J. Levin, J. C. McWilliams, A. J. Miller,
A. M. Moore, T. M. Powell, A. F. Shchepetkin, C. R. Sherwood, R. P. Signell, John C.
Warner, and J. Wilkin, 2007: Regional ocean forecasting in terrain-following coordinates:
model formulation and skill assessment. Journal of Computational Physics, accepted pending
minor revisions.
Hickey, B. M., 1998. Coastal oceanography of western North America from the tip of Baja to
Vancouver Island, in The Sea, The Global Coastal Ocean: Regional Studies and Syntheses,
vol. 11, edited by A. R. Robinson and K. H. Brink, Wiley, New York, 345-393.
Hill, K., N.C.H. Lo, B.J. Macewicz, and R. Felix-Uraga, 2006. Assessment of the Pacific
sardine (Sardinops sagax caerulea) for U.S. Management in 2006. NOAA-TM-NMFSSWFSC-386.
Hunt, J. C., 1996. The behavior and ecology of midwater cephalopods from Monterey Bay:
submersible and laboratory investigations. PhD thesis, University of California, Los Angeles,
USA.
Hurley AC, 1978. School structure of the squid Loligo opalescens. Fish Bull 76:433–442.
Isaacs, J.D., 1965: Larval sardine and anchovy interrelationships. CalCOFI Rep., 10, 102-138.
Jackson, G. D., and M. L. Domeier. 2003. The effects of an extraordinary El Niño/La Niña event on
the size and growth of the squid Loligo opalescens off Southern California. Mar. Biol.
142:925−935.
Jacobson, L. and A.D. MacCall, 1995. Stock-recruitment models for Pacific sardine (Sardinops
sagax). Can. J. Fish. Aquat. Sci., 52:566-577.
Kanamaru H, and Kanamitsu M, 2007: Scale-selective bias correction in a downscaling of global
analysis using a regional model. Mon. Wea. Rev., 135, 334-350.
King, DPF and PR Macleod. 1976. Comparison of the food and filtering mechanism of pilchard
Sardinops ocellata and anchovy Engraulis capensis off South West Africa, 1971-1972.
Invest. Rep. Sea Fish. Brch S. Afr. 111:1-29.
Landry, MR. 1977. Review of important concepts in trophic organization of pelagic ecosystems.
Helgol. Wiss. Meeresunters. 30:8-17.
2
Lasker, R., and P. E. Smith. 1977. Estimation of the effects of environmental variations on the
eggs and larvae of the northern anchovy. California Cooperative Fisheries Investigations
Reports 19:128–137.
Lavaniegos BE, Ohman MD, 2003: Long-term changes in pelagic tunicates of the California
Current. Deep-Sea Res., 50, 2473-2498
Lluch-Belda, D, RJM Crawford, T Kawasaki, AD MaCcall, RH Parrish, RA Schwartzlose, PE
Smith. 1989. Worldwide fluctuations of sardine and anchovy stocks: the regime problem. S.
Afr. J. Mar. Sci. 8:195-205.
Lo, N.C.H., D.A. Grith, and B.J. Macewicz, 2005. Spawning biomass of Pacific sardine
(Sardinops sagax) from 1994-2004 off California. CalCOFI Reports, 46:93-112.
Logerwell, E.A.& Smith, P.E. 2001. Mesoscale eddies and survival of late stage Pacific sardine
(Sardinops sagax). Fisheries Oceanography 10(1): 13-25.
Mais, K. F., 1974. Pelagic fish surveys in the California current. Fish Bull. 162, 1-79.
Mantua, N.J. and S.R. Hare, Y. Zhang, J.M. Wallace, and R.C. Francis, 1997: A Pacific
interdecadal climate oscillation with impacts on salmon production. Bulletin of the American
Meteorological Society, 78, 1069-1079.
McGowan, J.A., 1954: Observations on the sexual behavior and spawning of the squid, Loligo
opalescens, at La Jolla, CA. Cal. Fish Game 40:47−54.
McGowan, J.A., and C.B. Miller, 1980: Larval Fish and Zooplankton Community Structure.
CalCOFI Rep., 21, 29-35
McGowan JA, Cayan DR, Dorman LM, 1988: Climate-ocean variability and ecosystem response
in the northeast Pacific. Science, 281: 210-217.
Mendelsohn, R., Schwing, F.B. & S.J. Bograd, 2004: Nonstationary seasonality of upper ocean
temperature in the California Current. Journal of Geophysical Research 109: Art. No
C10015.
Miller, A. J., J. C. McWilliams, N. Schneider, J. S. Allen, J. A. Barth, R. C. Beardsley, F. P.
Chavez, T. K. Chereskin, C. A. Edwards, R. L. Haney, K. A. Kelly, J. C. Kindle, L. N. Ly, J.
R. Moisan, M. A. Noble, P. P. Niiler, L. Y. Oey, F. B. Schwing, R. K. Shearman, and M. S.
Swenson, 1999: Observing and modeling the California Current System. Eos, Transactions,
American Geophysical Union, 80, 533-539.
Moore, A.M., H.G. Arango, E. DiLorenzo, B.D. Cornuelle, A.J. Miller, and D.J. Neilsen, 2004.
A comprehensive ocean prediction and analysis system based on the tangent linear and
adjoint of a regional ocean model, Ocean Modeling, 7, 227.
3
Moore, A. M., H. G. Arango, E. Di Lorenzo, A. J. Miller and Bruce D. Cornuelle, 2007: An
adjoint sensitivity analysis of the Southern California Current circulation and ecosystem. Part
I: The physical circulation. Journal of Physical Oceanography, sub judice
Morel, FMM, RJM Hudson, and NM Price. 1991. Limitation of productivity by trace-metals in
the sea. Limnol. Oceanogr. 44:382-92.
Muccino, J. C., H. G. Arango, A. B. Bennett, B. S. Chua, B. D. Cornuelle, E. Di Lorenzo, G. D.
Egbert, D. Haivogel, L. Hao, J. C. Levin, A. J. Miller, A. M. Moore and E. D. Zaron, 2007:
The Inverse Ocean Modeling System. II: Applications. Journal of Atmospheric and Oceanic
Technology, sub judice.
Myers, R.A. 1998. When do environment-recruitment correlations work? Reviews in Fish
Biology and Fisheries 8: 285-305.
Okutani T., and J. A. McGowan. 1969. Systematics, distribution and abundance of the
epiplanktonic squid (Cephalopoda, Decapoda) larvae of the California Current April,
1954−March, 1957. Bull. Scripps Inst. Oceanog. 14:1−90.
Platt, T. and K Denman. 1978. The structure of pelagic marine ecosystems. Rapp. P.-V. Réun.
Cons. Int. Explor. Mer. 173:60-65.
Reiss, Christian S., Michael R. Maxwell, John R. Butler, and Annette Henry. 2004. Investigating
environmental effects on population dynamics of Loligo opalescens in the Southern
California Bight. CalCOFI Reports 45: 87-97.
Reiss, C., 2007: Re-analysis of the stock-recruitment relationship for Pacific Sardine (Sardinops
sagax) during a population growth phase. Sub judice.
Roemmich, D., and J. A. McGowan, 1995. Climatic warming and the decline of zooplankton in
the California Current, Science, 267,1324-1326.
Roughan, M.; Mace, A.J.; Largier, J.L.; Morgan, S.G.; Fisher, J.L.; Carter, M.L. 2005.
Subsurface recirculation and larval retention in the lee of a small headland: A variation on
the upwelling shadow theme. Journal of Geophysical Research Oceans
110(C10027):doi:10.1029/2005JC002898.
Roughan, M.; Terrill, E.J.; Largier, J.L.; Otero, M.P. 2005b. Observations of divergence and
upwelling around Pt. Loma, California. Journal of Geophysical Research 110 (C04011):
doi:10.1029/2004JC002662.
Rykaczewski, RR, and DM Checkley, Jr., 2007. Influence of ocean winds on the pelagic
ecosystem in upwelling regions. Proc. Natl. Acad. Sci., sub judice.
Shchepetkin, A.F., and J.C. McWilliams, 2004. The Regional Ocean Modeling System: A splitexplicit, free-surface, topography-following coordinate ocean model, Ocean Modeling, 4,
347-404.
4
Smith, PE, JK Horne, and DC Schneider. 2001. Spatial dynamics of anchovy, sardine, and hake
pre-recruit stages in the California Current. ICES J. Mar. Sci. 58:1063–1071.
van der Lingen CD, L Hutchings, and JG Field. 2006. Comparative trophodynamics of anchovy
Engraulis encrasicolus and sardine Sardinops sagax in the southern Benguela: are species
alternations between small pelagic fish trophodynamically mediated? Afr. J. Mar. Sci.
28:465-477.
Zeidberg, Louis D. 2004. Allometry measurements can determine the size and swimming speeds
of juvenile and adult Loligo opalescens (Cephalopoda: Myopsida) squid from in situ video.
Journal of Experimental Biology 207: 4195-4203.
Zeidberg, L. D., and W. M. Hamner. 2002. Distribution of squid paralarvae, Loligo opalescens
(Cephalopoda: Myopsida), in the Southern California Bight in the three years following the
1997 El Niño. Mar. Biol. 141:111−122.
Zeidberg, L. D., W. Hamner, K. Moorehead, and E. Kristof. 2004. Egg masses of Loligo
opalescens (Cephalopoda: Myopsida) in Monterey Bay, California following the El Niño
event of 1997−1998. Bull. Mar. Sci. 74:129−141.
Zeidberg, L. D., W. M. Hamner, N. P. Nezlin, and A. Henry. 2006. The fishery of the California
market squid, Loligo opalescens (Cephalopoda, myopsida), from 1981-2003. Fishery Bulletin
104: 46-59.
5