Evolution Evolution is a process that affects living things. It is the alteration of life through time. Evolution – the Janet and John version Living things have a set of characteristics… All living organisms possess a set of characteristics, for a human this means, two arms, two legs, a flat face with nose in the middle, two forward directed eyes and a mouth etc. There are thousands of these characteristics in an individual, not just external ones, but others that are not so obvious (e.g. The use of iron in the blood, the production of enzymes to digest food.) …………organisms in the same species may differ; this is variation. Even though all of the organisms belonging to the same species may possess the same set of characteristics, there will be a range of values for each characteristic within the species. Think about height, some people will be very tall and others will be very short, but most people will have a height close to the average. Other characteristics do not have a continuous spread of values, instead characteristics such as eye colour are discrete; they can be one thing or another (brown, blue, green). This difference between the characteristics in individuals is known as variation. Characteristics are controlled by genes. The characteristics possessed by an individual are controlled by genes. Genes are lengths of a molecule called D.N.A., which are found on strands called chromosomes present in the nuclei of cells. Genes are inherited from the parents……… During reproduction, the genes from the parents are ‘copied’, ‘mixed’ and passed on to the offspring, so the young inherit characteristics from the previous generation. ………but there may be changes, called mutations. Occasionally, during the ‘copying’ process mistakes are made, this is called a genetic mutation. This results in genes with a different set of instructions. Often this has no effect on the characteristics but sometimes this is not the case. The genetic mutation may result in a change in the characteristics. 1 Characteristics may give organisms an advantage, this is called natural selection. Some characteristics will give the individual an advantage in the struggle to survive. For example a rabbit with long legs may be able to run a little faster than the other rabbits. It is more likely to survive if the rabbits are chased by a fox, so we say it has been selected for survival. This is an example of natural selection. Advantageous characteristics may be inherited by the offspring. An organism that has survived can pass its genes onto its offspring. There is a good chance that the offspring will possess some of the advantageous characteristics of the parent, so they too will have an advantage in the struggle to survive. New species arise through this process of selection and inheritance. Over a very long period of time, through generation after generation, many new characteristics can become common in the population. Eventually, the individuals become so different from the originals, that a new species has been formed. This is called speciation. In this way new species develop from older species and this is evolution. The pattern of evolution Darwin thought that evolution was a slow and gradual process of change. He envisaged that the changes built up slowly in small steps, generation after generation; this hypothesis is called gradualism. The alternative model is known as punctuated equilibrium, in this hypothesis it is thought that the changes are sudden and dramatic. M odels for evolution Gradualism Punctuated equilibrium From our knowledge of genetics we know that both of these models are possible. The only way to determine which is correct is to look at the evidence – the fossil record. One particular group of trilobite fossils seems to show that gradualism is the mechanism. Successive trilobite fossils show a gradual increase in the number of segments. But most 2 of the fossil record does not show the same. We see particular organisms present in some rocks, then younger rocks show startlingly different fossils. Is this punctuated equilibrium or simply due to the many big gaps in the fossil record? Major turning points in the history of life 1. Life in the Precambrian. The Precambrian is the most mysterious time in the history of the Earth. It represents the time from the formation of the planet 4.6 billion years ago to the start of the Cambrian 600 million years ago. Initially the Earth could not support life; its surface was too violent and bombarded by harmful radiation from the sun. Eventually the oceans condensed and provided an environment in which life could begin. The origin of life is a mystery. There are two main schools of thought. The most popular theory is that life began from the combination of amino acids found in the early seas. A scientist (Stanley Miller) has shown that complex chemicals could form from simple building blocks under the right conditions (electrical discharges, such as in a storm). An alternative theory is called the Panspermia hypothesis. This suggests that the early Earth was “seeded” by biological material from space, delivered in the dust from passing comets. For a long time this idea was not accepted because it sounded rather like science fiction, but one of the most influential believers in this hypothesis (a Professor of Astronomy in Cardiff) recently found evidence of bacteria in the upper atmosphere which may have come from space. There are a number of problems with finding fossils from the Precambrian. The great age of the rocks means that there is a very high probability that they will have been destroyed at some point in the past. The other problem is that the organisms living at this time were small and soft bodied. The oldest fossils found are around 3.6 – 3.8 million years old, they belong to the super-kingdom called the Prokaryotes. These are single –celled organisms with no nucleus including bacteria and blue-green algae. The earliest prokaryotes would have simply consumed the amino acids in the water around them, later the blue-green algae evolved, which were capable of photosynthesis. Fossils from the Precambrian include stromatolites. These are sedimentary rocks that seem to contain narrow bands, building up to form mounds. We can see modern stromatolites forming in a place called Shark Bay in Australia. They are formed by a mat of photosynthetic bacteria, sticking the sediment together with some form of mucus. If the bacteria become covered by more sediment (such as when the tide deposits sand), the bacteria work their way up to the surface so they are exposed to the sunlight. A new layer of stuck sediment forms and the process is repeated many times. The lack of oxygen in the early atmosphere meant that the bacteria were anaerobic, gaining energy from chemical reactions that did not require the use of oxygen (e.g. sulphur reduction). Some of these anaerobic bacteria still live in anoxic environments (such as deep sediments) today. As time went on the simple prokaryotes evolved into the more complex eukaryotes. These are organisms that have distinct organelles such as 3 nucleus, mitochondria, chloroplasts etc. This evolutionary step may have happened when one type of prokaryotes incorporated another type into its body. Instead of providing food, they may have worked together, forming a more complex organism. The simplest animals are the protozoans. These are single-celled creatures that do not photosynthesise. Fossils from Zambia suggest that these gave rise to the multicellular animals such as sponges sponges about 1000 million years ago and by about 700 million years ago many invertebrates including creatures like jelly fish lived in the sea. A famous group of fossils from Precambrian rocks are called the Ediacaran fauna. These include a group called medusoids, rather like jellyfish. One set of these fossils, which are about 700 million years ago, is found in Charnwood Forest in England. 2. Life in the Cambrian – advanced invertebrates, development of hard parts. The Cambrian period is a very important time in the history of life. The fossil record shows that between 600 and 500 million years ago, there was a sudden and rapid increase in the number of species. This is called the Cambrian explosion. One of the most important developments was the evolution of organisms with the ability to extract dissolved chemicals from the surrounding seawater and to use them to form hard outer coverings called exoskeletons. Probably the best examples are the trilobites, a large group of arthropods with a jointed exoskeleton. They were particularly abundant during the Cambrian. One very important group of fossils come from a quarry in Canada containing a rock called the Burgess Shale. This is an example of exceptional preservation because the fine-grained sediment was able to preserve the organisms living on the seabed in incredible detail. Even soft tissue was preserved. Many of the organisms present in the Burgess Shale were unfamiliar to scientists, and they were so strange that it is quite difficult to interpret how they lived. One particular fossil called Hallucigenia was actually originally interpreted upside down; scientist thought that spines on its back were used for walking and the tentacles waved around in the water. The diagram shows the correct (?) interpretation for the animal. 3. The evolution of the vertebrates. One of the animals found as a fossil in Cambrian rocks possesses a flexible cord running along its back. This animal is the most primitive chordate – the group containing the vertebrates. Vertebrates are animals with a brain in a skull and a series of bones forming 4 the backbone. The backbone provides protection for a spinal cord, which connects the brain to the other parts of the body. The first of the five groups of vertebrates to evolve were the fishes. Fish are coldblooded animals that live in water extracting oxygen from the water using gills. The most primitive fish had no jaws, just a permanently open mouth, some of these fish still survive today, such as the lamprey, which attaches itself to larger fish by means of the mouth. Around 400 million years ago, fish with jaws evolved and during the Devonian there were many species of fish filling the world’s oceans and lakes; one genus, called Dunkleosteus grew to 9 metres in length and by examining the massive armoured jaws we can interpret its mode of life as a formidable predator. One group of fish did not evolve to use a skeleton of bone, instead their skeletons are composed of the softer, lighter material called cartilage. These are the sharks and rays, a group that has successfully populated the seas since the Devonian. Most modern fish belong to the group known as the bony fishes whose fins are supported by bones but another group of bony fishes had fins that were mainly flesh. They evolved about 390 million years ago and they are though to be the ancestors of the land vertebrates. Lungfishes belong to the fleshy finned fishes. They are able to move onto the land and to gulp air if hot dry weather dries up pools and lakes. About 350 million years ago these fishes evolved into the first amphibians. The fleshy fins developed into legs and the gulping breathing technique of their ancestors developed into proper lungs. Some of the amphibians grew to very large sizes (up to 9 metres) and until the evolution of the reptiles; they were the dominant land animals. Ichtyostega – an early amphibian 350 million years ago the amphibians gave rise to the reptiles, the first group of vertebrates able to live permanently on land. Reptiles evolved to occupy many ecological niches; their dominance of the Earth reached a peak during the Mesozoic era. The terrestrial reptiles belong to the group known as the dinosaurs but other very large reptiles 5 dominated the oceans (the plesiosaurs and ichthyosaurs) and even conquered the air (the pterosaurs). Other reptile groups that survive today include the crocodiles, the tortoises and turtles, the snakes and lizards. The four main groups of reptiles evolved around 270 million years ago. The Reptiles' Family Tree Dinosaurs, crocodiles, & pterosaurs. Snakes & lizards Mammal-like reptiles. Tortoises & turtles Sea reptiles. Ancient reptiles Amphibians Many scientists now believe that the dinosaurs were actually warm blooded and a recent discovery in China suggests that some possessed feathers, probably as a form of insulation. This is the point when birds are thought to have evolved from the reptiles. The earliest fossil of a true bird is Archaeopteryx from the late Jurassic. It is a remarkable fossil because it shows many of the characteristics of both groups; the feathered wings of a bird and the bony tail, teeth and clawed fingers of a reptile. This fossil is another example of exceptional preservation, found from a quarry of very finegrained limestone in Germany. The mammal-like reptiles evolved around the start of the Permian period and some of them developed into massive carnivores like Dimetrodon. This animal had a huge “sail” of skin supported by vertical bones, running down its back. This may have been an adaptation for controlling its body temperature. The first mammals evolved from the mammal-like reptiles about 190 million years ago. During the Mesozoic, the mammals were small rodent-like animals but once the dinosaurs became extinct, they evolved to fill many of the niches previously occupied by the dinosaurs. They are now the dominant group of vertebrates. 6 4. The evolution of land plants. Plants initially evolved in the Precambrian as tiny unicellular algae, capable of photosynthesis. They produced oxygen as a waste product and this gas built up in the atmosphere. Eventually the oxygen built up to become a significant proportion of the atmosphere. Oxygen molecules (O2) reacted together to form molecules of another gas called ozone (O3). Ozone high in the atmosphere has the ability to prevent harmful ultraviolet radiation from reaching the Earth’s surface. The presence of an ozone layer was crucial to the colonisation of the land. True plants (those organisms with a cell surrounded by a cellulose cell wall) include the simplest group; the algae. They evolved from blue-green algae about 1000 million years ago. The first land plants to evolve were mosses, which appeared about 400 million years ago. Algae are adapted to live in water, obtaining all their needs directly from the water, mosses live in cushion-like clumps close to the ground so they remain wet, but to colonise the land, plants must be able to transport materials around the body. Vascular plants possess a transport system, several groups of vascular plants evolved during the Palaeozoic including the ferns, horsetails and gymnosperms (seed ferns and conifers). All of these groups reproduced using spores. Until 120 million years ago there were no flowers. Flowering plants (the angiosperms) evolved during the Mesozoic. The flowering plants are the most important land plants today; they exist in many different environments reproducing by means of pollination. This involves transferring pollen from one flower to another, this can occur by means of the wind, or commonly using insects. The evolution of insect pollinated plants and the evolution of insects is an interesting example of how two entirely different groups evolve to benefit each other. The plants evolved flowers that were more attractive to insects (e.g. colours, nectar, scent) and the insects evolved to become more efficient at obtaining nectar (e.g. the coiled proboscis of a butterfly). Pollen is so small and produced in such quantities that it is very widely spread by currents in the atmosphere. Pollen is composed of a very tough material, resistant to destruction, and is one of the most common microfossils found in sedimentary rocks. They are excellent as environmental and climatic indicators. During the Tertiary period, the grasses evolved. ESTA GEOTREX The Geology Teachers Resource Exchange Contributor: Owain Thomas School Date: 22:04:05 Establishment: Amman Valley 7