THE THEORY OF EVOLUTION NATURAL SELECTION AND EVIDENCE OF EVOLUTION CHARLES DARWIN AND NATURAL SELECTION Has been said that its impossible to understand any field of biology without understanding evolution. The modern theory of evolution is perhaps the most fundamental concept in Biology. A rich fossil record has been important to biological sciences since the 18th century. It formed the basis of early evolutionary concepts Scientist wondered where these species came from, why some no longer existed, and what relationship, if any, they have to modern species. Became clear that Earth was much older than expected Early 19th Century scientist were convinced that life slowly changed over time, or evolved. Charles Darwin (1809-1882) English scientist Father of modern evolutionary theory 1831 at age of 21 recommended for position of naturalist on HMS Beagle Ship chartered for a 5 year collecting and mapping expedition to South America and South Pacific. A voyage that would forever change the science of Biology Darwin’s job was to collect, study, and store specimens discovered The voyage took him through a number of environments with some of the World’s greatest biodiversity Galapagos Islands were among the most important Group of small islands about 1000km off west coast of South America Support a great diversity of animal and plant life Darwin studied and compared the anatomy of many species of reptiles, insects, birds, and flowering plants unique to the islands. Similar species seen in other parts of the world. By the end of the trip he was convinced that evolution occurs- species change over time. Darwin returned from his voyage in 1836 Spent next 22 years studying his collection and conducting experiments Darwin was also intrigued by an essay written at the time that suggested human population was growing faster than the food supply on Earth. He knew many species produce large numbers of offspring, and since Earth was not covered with such species, their must be a struggle for existence among individuals. He envisioned many kinds of struggles = comp for food and space, escape from predators, and need to find shelter. Only some of individual in population survive the struggles to produce offspring. His answer came from his own pigeon breeding experiments Individual pigeons have different variations of traits that can be inherited. Breed pigeon with desirable variations, he was able to produce offspring with these same features. Artificial Selection = a breeder selects particular traits Same force in nature? Using data he gathered from natural world he began his well known idea of evolution by natural selection. Natural Selection = a mechanism for change in populations that occurs when organisms with favorable variations for a particular environment survive, reproduce, and pass these variations on to the next generation. Organisms with less favorable variations are less likely to survive and pass on traits to the next generation. “Survival of the Fittest” Therefore, each new generation is made up largely of offspring from parents with the most favorable variations. Fig 18.3 On the Origin of Species by Natural Selection: 1859 Convinced many scientist that evolution does occur through natural selection Unifying theory for all biology (today) NATURAL SELECTION AND ADAPTATION Why do some plants have thorns? Why some animals have distinct colors? Adaptation = any trait that aids the chances of survival and reproduction of an organism. Mole-rat possesses a number of features that adapt them to life underground Ancestors lived above ground and didn’t have these characteristics Fig 18.4 Structural Adaptation: = changes in the structure of body parts Some obvious ones are used for defense against predators Thorns on rosebushes Spines on sea urchins Mimicry: = structural adaptation that provides protection for an organism by enabling it to copy the appearance of another species Fig 18.5 Camouflage: = a structural adaptation that enables an organism to blend in with its surroundings A defense adaptation that involves changes to the color of organism More likely to escape predators and survive to reproduce Structural adaptations may take millions of years to develop Depends on type and rate of reproduction and environmental factors Some evolve relatively quickly Camouflage adaptations in moth’s evolve in 100 years Physiological Adaptation: = changes in an organisms metabolic process Can develop rapidly Some medicines developed during this century are no longer useful 50 years ago antibiotic drug penicillin “wonder drug” Could kill many types of disease-causing bacteria Not as effective today Many species of bacteria have evolved physiological adaptations make them resistant to penicillin. “Multi-drug resistance” Rapid changes in pest organisms such as insects and weeds Resistant to chemicals after being exposed to pesticides EVIDENCE OF EVOLUTION Most of the evidence of evolution comes from indirect sources Ancestors of whales once land-dwelling, wolf-like animals? Study their fossils. Although the fossil record in incomplete, it does provide evidence that evolution has occurred. Provides a record of earlier life. “Completing a jigsaw puzzle with missing pieces” May still be able to understand the overall picture Same with fossils Can’t trace each and every step because intermediate forms cannot be found. Still understand the general pathway of evolution Can show a step-by-step sequence of evolution Fig 18.7: Camal Used skulls, teeth, and limb bones 40-50 mya camals were a small, rabbit sized animal Homologous Structures: = modified structure that is seen among different groups of descendants Forelimbs of animals look the same in Fig 18.8 From outside, they would look different and vary in function, but they are similar in skeletal features. Evolved from a common ancestor with the same basic limb structures Over the course of evolution, vertebrates moved into different environments. Within each environment, animals faced very different needs for survival Limbs more useful in environment = survival Became adapted to different ways of life. Modified structures are still the same even though they aren’t used for same function. Often similar in structure, in function, or both However, similarity of function doesn’t always mean that the 2 organisms are closely related. Homologous structures probably had a common evolutionary origin. Analogous Structures: = any body part that is similar in function but different in structure Butterfly wing and Bird wing Fig 18.9 Evolved independently in 2 distantly related groups of ancestors Can’t be used to indicate evolutionary relationships, but they do provide evidence of evolution. Insect and bird wings most likely evolved when their different ancestors independently adapted to similar ways of life. Vestigial Structures: = any body structure that is reduced in function in a living organism, but may have been used in an ancestor. Functionless structures indicate evolutionary pathways. What is appendix used for? Evolutionary biologist can tell you what an appendix WAS for. Organisms contain structures that have reduced functions but were once used by ancestral organisms. Appendix = intestine, different diets, secrete hormones/enzymes/bile? Particular structures become vestigial as species change in form and behavior The structure, although it may serve no function, continues to be inherited as part of the body plan for that species. The eyes of sightless species, common Mole-Rat and Cave Fish, may be considered vestigial because eyes were most likely functional in ancestors. Fig 18.10 = Ostrich wings Embryonic development shows evolution from a common ancestor. Look at embryos Fig 18.11: Vertebrates Tail, gills slits, can be seen in all species As development continues, become more distinct Common Ancestors Genetic comparisons may reveal hidden relationships Help tell us how different species are related Comparison of DNA and RNA within a taxonomical group More reliable that anatomical relationships Use nucleotide sequencing to indicate levels of relationships Since the 1970’s research has shown that by comparing nucleotide sequence in the DNA and RNA of species, its possible to constrict hypothetical evolutionary trees showing levels of relationships Fig 18.12