New species of Franchia and Protozigzagiceras (Ammonoidea, Middle Jurassic): the
phyletic origin of Zigzagiceratinae
by Sixto R. Fernandez-Lopez1* and Giulio Pavia2
1
Departamento de Paleontología, Facultad de Ciencias Geológicas, calle José Antonio Novais
2, Universidad Complutense de Madrid, 28040-Madrid, Spain; e-mail: sixto@geo.ucm.es
2
Dipartimento di Scienze della Terra, via Valperga Caluso 35, 10125-Torino, Italy; e-mail:
giulio.pavia@unito.it
*Corresponding author
Measurements
A. Measurements for the specimens of Franchia arkelli Sturani, 1967
Specimens
Figures
PU31689
Figs. 4A–B
holotype
[M]
PU31690
Figs. 5E–F
[M]
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
220.0
60.0
27.3%
61.2
27.8%
101.6
46.2%
1.02
14
35
2.5
170.0
53.8
31.6%
52.0
30.6%
78.2
46.0%
0.97
15
–
–
147.0
43.5
29.6%
40.0
27.2%
71.8
48.8%
0.92
–
–
–
114.0
34.5
30.3%
34.7
30.4%
54.8
48.1%
1.01
20
45
2.3
16
–
–
PU112317
–
[M]
92.0
31.9
34.7%
29.4
32.0%
40.2
43.7%
0.92
PU31691
Figs. 5A–C
[M]
70.0
22.5
32.1%
22.0
31.4%
33.0
47.1%
0.98
–
–
–
53.0
13.7
25.8%
16.8
31.7%
23.9
45.1%
1.23
22
–
–
PU111325
Fig. 6D
[M]
75.0
23.8
31.7%
23.3
31.1%
31.9
42.5%
0.98
17
–
–
PU112318
Figs. 4D, 7A
[m]
31.5
9.3
29.5%
9.8
31.1%
14.6
46.3%
1.05
–
–
–
B. Measurements for the specimens of Franchia subalpina sp. nov.
Specimens
Figures
PU111399
Fig. 6A
holotype
PU111578
PU111573
–
Figs.6B–C, 7B–
C
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
140.0
46.5
33.2%
43.0
30.7%
57.7
41.2%
0.92
–
45
–
75.0
26.0
34.7%
30.0
40.0%
33.0
44.0%
1.15
16
–
–
[M]
45.0
12.5
27.8%
20.1
44.7%
19.2
42.7%
1.61
16
–
–
[m]
32.0
10.5
32.8%
9.8
30.6%
15.3
47.8%
0.93
19
32
1.7
[M]
C. Measurements for the specimens of Protozigzagiceras torrensi (Sturani)
Specimens
Figures
PU31725
Torrens 1987,
pl. 9, fig. 3
PU31717
Figs. 11A–B
PU31711
Torrens 1987,
pl. 8, fig. 2
PU31719
Sturani 1967,
pl. 15, fig. 1
PU31720
Figs. 12A–B
[M]
[M]
[M]
[M]
[M]
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
115.6
41.0
35.5%
45.1
39.0%
43.8
37.9%
1.10
21
44
2.1
94.4
33.7
35.7%
38.0
40.3%
37.7
39.9%
1.13
20
44
2.2
105.7
46.2
43.7%
48.2
45.6%
30.5
28.9%
1.04
17
48
2.8
79.0
30.0
38.0%
31.5
39.9%
26.5
33.5%
1.05
–
–
–
98.9
38.0
38.4%
38.0
38.4%
36.0
36.4%
1.00
20
39
2.0
75.9
27.4
36.1%
29.6
39.0%
29.2
38.5%
1.08
24
42
1.8
93.3
42.8
45.9%
39.4
42.2%
28.6
30.7%
0.92
21
46
2.2
77.9
31.8
40.8%
31.6
40.6%
24.6
31.6%
0.99
19
44
2.3
67.4
23.5
34.9%
25.9
38.4%
28.7
42.6%
1.10
22
36
1.6
52.0
18.1
34.8%
20.6
39.6%
22.9
44.0%
1.14
20
34
1.7
PU31680
Figs. 10G–H
[m]
61.0
21.5
35.2%
21.8
35.7%
25.0
41.0%
1.01
20
–
–
PU31677
Figs. 10I–J
[M?]
55.0
20.1
36.5%
23.5
42.7%
23.0
41.8%
1.17
26
–
–
PU31681
Figs. 10E–F
[m]
52.0
18.0
34.6%
19.5
37.5%
22.8
43.8%
1.08
21
36
1.7
38.0
13.0
34.2%
14.2
37.4%
18.1
47.6%
1.09
20
31
1.6
PU31676
Figs. 10A–D
holotype
[m]
49.0
15.6
31.8%
20.5
41.8%
21.2
43.3%
1.31
24
–
–
PU31682
Figs. 10K–M
[m]
44.0
15.0
34.1%
16.7
38.0%
20.5
46.6%
1.11
20
33
1.7
33.0
10.0
30.3%
12.5
37.9%
16.1
48.8%
1.25
23
34
1.5
PU111577
Pavia et al. 2008,
pl. 3, figs. 9–10
39.2
13.0
33.2%
17.0
43.4%
18.0
45.9%
1.31
22
–
–
[M]
D. Measurements for the specimens of Protozigzagiceras tethycum sp. nov.
Specimens
Figures
PU31694
Figs. 16H–I
PU111299
[M]
Figs. 16E–G
[m]
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
51.0
17.0
33.3%
17.0
33.3%
20.1
39.4%
1.00
20
39
2.0
37.0
14.0
37.8%
14.0
37.8%
15.9
43.0%
1.00
21
–
–
40.0
14.6
36.5%
11.6
29.0%
17.1
42.8%
0.79
23
46
2.0
28.0
9.0
32.1%
8.1
28.9%
12.9
46.1%
0.90
19
–
–
PU111574
Figs. 16J–L
holotype
[m]
36.0
10.0
27.8%
12.4
34.4%
17.4
48.3%
1.24
19
33
1.7
PU31695
Figs. 16A–D
[m]
30.0
9.7
32.3%
10.5
35.0%
13.6
45.3%
1.08
19
36
1.9
33.0
10.0
30.3%
12.5
37.9%
16.1
48.8%
1.25
23
34
1.5
E. Measurements for the specimens of Protozigzagiceras flexum sp. nov.
Specimens
Figures
PU112319
Figs. 17E–F
PU111326
Figs. 17A–D
holotype
[M]
[m]
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
51.3
16.3
31.8%
14.9
29.0%
21.6
42.1%
0.91
–
–
–
41.5
13.7
33.0%
14.1
34.0%
17.4
41.9%
1.03
24
46
1.9
41.0
13.1
32.0%
15.0
36.6%
19.0
46.3%
1.15
–
–
–
30.0
9.2
30.7%
11.0
36.7%
14.6
48.7%
1.20
24
38
1.6
F. Measurements for the specimens of Protozigzagiceras densum sp. nov.
Specimens
Figures
PU111322
Figs. 18A–D
holotype
[m]
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
35.6
11.5
32.3%
13.8
38.8%
16.1
45.2%
1.20
18
46
2.6
G. Measurements for the specimens of Megazigzagiceras subarabicum sp. nov.
Specimen
FSL 177 639
Figures
Figs. 19A–B
holotype
[m]
D
H
H/D
W
W/D
U
U/D
W/H
Ni/2
Ne/2
i
100.0
22.3
22.3%
25.0
25.0%
60.0
60.0%
1.12
16
–
–