CH. 22 Origin of Species (Chapter 22.1-22.5) By: Jose Solarzano (Chief), Jose Arambulo, Daniel Guevarra,Eric Galicia, and Jarod Smith. Period 4 Biology 22.1 The Nature of species and the Biological Species Concept Reproductive Isolating Mechanisms Mechanism Description Prezygotic Isolating Mechanism Prezygotic Isolating Mechanism Geographic Isolation Species occur in different areas, which ae often separated by a physical barrier, such as a river or mountain range. Ecological Isolation Species occur in the same area, but they occupy different habitats and rarely encounter eachother. Behavioral Isolation Species differ in their mating rituals Temporal Isolation Species reproduce in different seasons or at different times of the day. Mechanical Isolation Structural differences between species preventing mating. Prevention of Gamete Fusion Different species/animals cannot successfully sexually reproduce. Postzygotic Isolating Mechanism Postzygotic Isolating Mechanism Hybrid inviability or Infertility Hybrid Embryos do not develop properly, hybrid adults do not survive in nature, or hybrid adults are sterile or have reduced fertility. 22.2 Natural Selection and Reproductive Isolation -Selection may reinforce isolating mechanisms. Populations may evolve complete reproductive isolation allopatry. If populations that have evolved only partial reproductive isolation come into contact, natural selection can lead to increased reproductive isolation, a process termed “reinforcement.” Alternatively, genetic exchange between the populations can lead to homogenization and thus prevent speciation from occurring. CH. 22 Origin of Species (Chapter 22.1-22.5) By: Jose Solarzano (Chief), Jose Arambulo, Daniel Guevarra,Eric Galicia, and Jarod Smith. Period 4 Biology -22.3 Natural Selection and Reproductive Isolation -The Role of Genetic Drift and Natural Selection in Speciation ● Genetic Drift: randomly generated changes in a population’s genetic ● Random changes may cause reproductive isolation- In small populations, genetic drift may cause populations to diverge; the differences that evolve may cause the populations to become reproductively isolated ● Adaptation can lead to speciation- Adaptation to different situations/environments may incidentally lead to reproductive isolation. In contrast to reinforcement, these differences are not directly favored by natural selection because they prevent hybridization Natural Selection can directly select for traits that increase reproductive isolation -22.4 The Geography of Speciation Speciation: A two-part process, initially identical populations must diverge, and second, reproductive isolation must evolve to maintain differences Problem: Homogenizing effect of gene flow between populations is constantly acting to erase any arising differences by genetic drift or natural selection Gene flow occurs only between populations that are in contact, and geographical isolation occurs in populations for many reasons Ernst Mayr: First biologist to demonstrate that geographically separated (allopatric) populations appear more likely to have evolved differences leading to speciation (ex.) Little paradise kingfishers vary little throughout the wide range in New Guinea, despite great variation in in topography and climate. allopatric speciation: takes place when populations are geographically separated sympatric speciation: takes place when populations are not geographically separated Sympatric speciation can occur either instantaneously or over the course of multiple generations, without geographic separation Instantaneous sympatric speciation occurs when an individual is born that is reproductively isolated from all other members of its species CH. 22 Origin of Species (Chapter 22.1-22.5) By: Jose Solarzano (Chief), Jose Arambulo, Daniel Guevarra,Eric Galicia, and Jarod Smith. Period 4 Biology Polyploidy: produces individuals that have more than two sets of chromosomes, often seen in plants This polyploid factor can arise in two ways; autopolyploidy or allopolyploidy Autopolyploidy: all of the chromosomes may arise from a single species, which might happen due to an error in cell division that causes a doubling of chromosomes, those of which that contract autopolyploidy this way are referred to as tetraploids because they have four sets of chromosomes Allopolyploidy: A more common type of polyploid speciation, which may happen when two species hybridize, resulting in offspring with one copy of the chromosomes of each species, and are usually infertile due to incorrect chromosome pairing, but some are otherwise healthy, can reproduce asexually, and can become fertile through certain events About half of the approximately 260,000 species of plants have a polyploid episode in their history, including wheat, cotton, tobacco, sugarcane, bananas, and potatoes Speciation by polyploidy is known to occur in a variety of animals, such as insects, fish, and salamanders, but more rarely than plants Sympatric speciation can occur over the course of multiple generations through the process of disruptive selection which can cause a population to contain individuals exhibiting two different phenotypes Before two phenotypes could become different species, they would have to evolve reproductive isolating mechanisms Sympatric speciation by disruptive selection is a rare event, yet a number of cases have appeared that are difficult to perceive in any way other than sympatric speciation -22. 5 Adaptive Radiation a Biological Diversity Adaptive radiations- occurs when a species finds itself in a new or suddenly changed environment with many resources and few competing species Key innovation-the evolution of a new trait that allows individuals to use previously inaccessible parts of the environment; may also trigger an adaptive radiation Character displacement- A process in which natural selection favors individuals in a species that use resources not used by other species. This results in evolutionary change leading to species dissimilar in resource use Ground Finches- adapted the size of their bills to the size of the seeds that they feed on/ six species exist CH. 22 Origin of Species (Chapter 22.1-22.5) By: Jose Solarzano (Chief), Jose Arambulo, Daniel Guevarra,Eric Galicia, and Jarod Smith. Period 4 Biology Tree finches- Five species of this finch exist. Four of them have bills suitable for feeding on insects, but one (the woodpecker) has a chisel-like beak. It usually carries a twig around so it can probe crevices for insects. Vegetarian finch- Have very heavy bills to wretch buds from branches Warbler finch- Are unusual birds with slender, warblerlike beaks that continuously search the mainland for leaves and branches for insects.