Biodiversity in the Library Simpson's Diversity Index is a measure of diversity. In ecology, it is often used to quantify the biodiversity of a habitat. It takes into account the number of species present, as well as the abundance of each species. Biological Diversity - the great variety of life. Biological diversity can be quantified in many different ways. The two main factors taken into account when measuring diversity are richness and evenness. 1. Richness - Richness is a measure of the number of different kinds of organisms present in a particular area. For example, species richness is the total number of different species present in a community. Some communities may be simple enough to allow complete species counts to determine species richness. However, this is often impossible, especially when dealing with insects and other invertebrates, in which case some form of sampling has to be used to estimate species richness. 2. Evenness - Evenness is a measure of the relative abundance of the different species making up the richness of an area. To give an example, we might have sampled two different fields for wildflowers. Both samples have the same number of species (3) and the same total number of individuals (1000). The total number of individuals in sample 1 is quite evenly distributed between the three species. In the second sample, most of the individuals are buttercups, with only a few daisies and dandelions present. Sample 2 is therefore considered to be less diverse than sample 1. Flower Species Daisy Dandelion Buttercup Total Table 1: Flowers Numbers of individuals Sample 1 Sample 2 300 20 335 49 365 931 1000 1000 1. What two main factors are taken into account when measuring diversity? 2. What is species richness? Which sample above is more “rich”? 3. What is species evenness? Which sample above is more “even”? 4. In this lab we are looking at the different books found in the library. Therefore, we will use which index (richness or evenness) to determine diversity? Which would you consider to show greater diversity, a small island with 100 lizards that belong to 5 species or a larger island that has 2500 lizards belonging to 5 species? Both islands have the same number of species, therefore that number alone cannot properly show biodiversity. For that reason, diversity is measured in different ways: Method One: D = (S-1)/(Log N) D = diversity – the larger the number, the more diverse S = number of species N = total number of individuals in the sample. Determine the diversity of lizards on the islands and circle the most diverse Small Island diversity = _____________ Large Island diversity = _____________ Method Two: Some measures of diversity include a way of counting the number of individuals of each species as well as the total number of species. One commonly used measure of species diversity that includes proportions of individuals is represented by the Shannon-Weaver equation, which is: -sum (pI ln pI) H = diversity- the larger the number, the more diverse Pi = relative abundance = ni/N ni = number of individuals of species “i ” N= total number of individuals of all species Small Island Diversity ni pi Species i Lizard Species 1 1 (species “#1”) 22 individuals 2 20 3 25 4 15 5 18 Lizard Species 2 Lizard Species 3 Lizard Species 4 Lizard Species 5 S= 5 Total number of species pi ln pi ni/N= 22/100 N=100 1.00 Total individuals ln pi - H = sum (Pi ln(Pi)) H= Large Island Diversity ni pi Species i Lizard Species 1 1 (species “#1”) 635 individuals 2 561 3 664 4 391 5 249 Lizard Species 2 Lizard Species 3 Lizard Species 4 Lizard Species 5 S= 5 Total number of species ln pi pi ln pi ni/N= - N= 100 H = sum (Pi ln(Pi)) 1.00 Total individuals H= Determine the H value (diversity) of lizards on the islands and circle the most diverse: Small Island diversity = _____________ Large Island diversity = _____________ Did both methods produce the same answer? Which do you feel is more accurate? Why? Procedure: You will determine the diversity of books in the library using the S-W Index. In groups you will record the number of individuals based on the your assigned species (color) of book in the section. Please remember to look for the predominant color on the spine of the book. Remember that members of the same species may come in various sizes. Record your data on the table on the next page Library location: Spine Color Black Number Spine Color Yellow Brown White Blue Red Green Purple Orange Totals of all colors: Number Library Diversity Location: Species i Black Brown Blue Green Orange Purple Red White Yellow Non-Fiction 500-700 Group 4 Non-Fiction 700-800 Group 5 Non-Fiction 800-900 pi ln pi ni/N= 4 5 6 7 8 9 N= Total individuals Number of individuals Non-Fiction 300-500 Group 3 ln pi 3 Total number of species Non-Fiction 000-300 Group 2 pi 2 S= 5 Group 1 ni 1 (species “#1”) 1.00 - H = sum (Pi ln(Pi)) H= H Value Rank of diversity (1 being most diverse)