12. Edwards CA. 2004 Earthworm Ecology. Boca Raton: CRC Press.

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Text S1: Trait data
We included traits which are commonly measured and shown to be key traits in affecting
the organisms’ response to environmental change and/or to have functional significance: for
carabid beetles [1 - 5], pollinators [6 - 8], earthworms [9 - 13], nematodes [14] and dung beetles
[15 - 18]. Ecosystem functions are determined not only by the specimens per capita efficiency,
but also by the specimens numerical abundance. Hence, we argue that in observational studies
traits that affect community response to environmental change (response traits) and traits that
affect ecosystem processes (effect traits) are both important for predicting ecosystem
functioning, because species composition (with regard to species richness and abundances in a
community) can be expected to be affected by species response traits while the efficiency of
resource exploitation will be determined by species effect traits as well [19]. An example would
be a community in which the dominant predator, though not possessing optimal traits for prey
exploitation, nevertheless may reach high abundances – and exert high pressure on the prey
community - due to being resilient to disturbance. Moreover, only few animal traits have been
experimentally tested for their effect on ecosystem functioning and it is often hard to distinguish
between response and effect traits because some traits may be both or response and effect traits
can be strongly correlated [20 - 22].
Trait data for carabids included: body size (continuous), time of reproduction (spring,
summer or inconclusive), activity period (diurnal, nocturnal or inconclusive), wing morphology
(macropterous, brachipterous or dimorphic), overwintering strategy (adult, imago or
inconclusive), trophic level (mainly herbivore, mainly predator or omnivore), humidity
preference (dry, medium-dry, medium, medium-wet, wet or all), light preference (non-shaded,
partly shaded, shaded) and habitat preferences (open, wooded or inconclusive). Trait data for
pollinators included: nest place and construction (carder, cleptoparasite, excavator:ground,
renter:existing cavities, renter:snail shells, social parasite), mean intertegular distance (ITD) of
females (continuous), habitat specialism (number of main habitat use), diet specialization
(oligolectic, polilectic), buzz pollinator (yes, no), tongue length (long, short), sociality (eusocial,
solitary), voltinism (univoltine, univoltine or bivoltine, bivoltine, univoltine to multivoltine,
multivoltine), total months active (continuous), emergence time (6 levels: Jan to Jun). Trait data
for earthworms included: ecological group (edogeic, epigeic,anecic), weight of adults and
juveniles (continuous), width of adults (continuous), number of segments (continuous),
reproductive staregy (obligatory biparental, parthenogenetic), hatchling peaks (April to May,
May to June), number of cocoons per worm (continuous), days to hatching (continuous), months
to sexual maturity (continuous), dry weight of gut content (continuous), gut passage time of food
in h (continuous), length of adults and juveniles (continuous), ph (continuous). Traits for
nematodes included: length of adults and juveniles (continuous), width of adults and juveniles
(continuous), mass of adults and juveniles (continuous), type of feeding interaction
(bacterivorous, fungivorous, fungivorous & herbivorous, herbivorous, omnivorous, omnivorous),
trophic level (2 levels: first and second). Traits for dung beetles included activity period (diurnal,
nocturnal), nesting strategy (roller, tunneller), length (continuous), mass (continuous) and body
size (large, small).
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