Research Interests
Biological oceanography with emphasis on primary production as a function of nutrient supply,
light and variations in marine climate. Pioneered integration of marine laboratory and field
investigations in Norway, including studies of chemical composition of phytoplankton using
unialgal cultures. Also specialised in the photobiology and bio-optics of marine phytoplankton,
with emphasis on photo-acclimation and P vs E curves. The field activity has, besides local
waters been concentrated on the ecology of the Barents Sea and the Southern Ocean.
Forskningsprofil
Biologisk oseanografi med vekt på primærproduksjon som funksjon av tilførselen av
næringssalter, lys og klimavariasjoner i havet. Var pioner for å integrere laboratorie- og
feltarbeid. Kjemisk sammensetning av planktonalger som funksjon av nærings- og
lysbegrensning (på artsnivå). Bruk av enalgekulturer i laboratoriet for å oppnå målet, inkludert
tolkning av fotsyntese-mot-lys data for anvendelse på feltdata. Initierte fotobiologi som en
spesialitet ved TBS. Feltinnsatsen har, foruten lokale farvann vært konsentrert om polarstrøk
(Barentshavet, Sørishavet).
Dr. students: Torunn Beate Hancke (fluorescens-kinetikk og fotobiologi hos giftige alger
Erlend Kjeldsberg Hovland:Bio-optikk hos marine kalkflagellater under
blomstringer
M.Sc.student: Raluca Rogin (Romania; akvakultur av stør for kaviarproduksjon, tilhørende
miljøproblemer)
Teaching in 2010:
BI 2060: Marine Ecology
BI 2061: Physical Oceanography
BI 2098: Experts in Team (Underwater Robotics)
BI 3061: Biological Oceanography (masters level)
C.V.
Name/Position:
Egil Sakshaug, professor in marine ecology at NTNU (The Norwegian University of Science
and Technology), Trondheim, Norway
Education:
1978: Dr. Philos., University of Trondheim
1967 Cand. real., University of Oslo
Positions:
1982-
Professor, tenured at the Biological station, Department of Biology, NTNU
1970-1982
1969-1970:
1996-
Associate professor, tenured at the Biological Station, NTNU
Grantee, International Biological Programme (IBP), NTNU
Courtesy professor, University of South Florida, College of Marine Sciences, St.
Petersburg, Florida
Sabbaticals:
1973
Graduate School of Oceanography, University of Rhode Island
1973-74, 1986
Food Chain Research Group, Scripps Institution of Oceanography, La
Jolla, California, USA
1985
Bigelow Laboratory for Ocean Sciences, West Boothbay Harbor, Maine, USA
1996
Department of Marine Science, University of South Florida, St. Petersburg
2005
Institut de Recherche pour Développement, Nouméa, Nouvelle Calédonie
2007
University of Nagoya, Japan, visiting professor
Awards and memberships:
2006199619941994
199419931993
19901976-
Honorary member, Norwegian Association of Oceanographers, member since 1970
Norwegian Academy of Science and Letters, Oslo (elected member)
Norwegian Academy of Technological Sciences, Trondheim (elected member)
Nansen-Prize of the "Fram" committee for merits in polar research
American Geophysical Union (AGU)
Royal Norwegian Society of Science and Letters, Trondheim (elected member)
Antarctic Service Medal; US Coast Guard
Norwegian Society of Photobiology and Photomedicine,
American Society of Limnology and Oceanography (ASLO)
Cruises and field work:
Trondheimsfjord, Gulf of California, Norwegian Coastal Current, Lindåspollene (Norway),
Scotia Sea (Southern Ocean), North Sea, Litoralstation Sylt (Germany), Antarctic
circumnavigation cruise, Barents Sea, section from Capte-Town to Dronning ;aud Land
(Antarctica)
Teaching and tutoring:
Since 1971, lectures on algal systematics (seaweeds), physical oceanography for undergraduates,
courses in methods and phytoplankton taxonomy, lectures on marine ecology and biological
oceanography (master level); 2 summer courses (Nordic Council and EU); tutoring of 22 master
students and 8 Ph.D students; 2 summer courses (Nordic Council and EU)
Grants:
Norwegian Research Council (18), EU (7);
New in 2010: HYDRALAB (EU). Research infrastructure, initiator in 1998, manager 19982004; activity still running, now under HYDRALAB
Administrative (partial):
2004-06
2004
2001-04
2000-04
2000-04
2002-09
2002
1999-2003
1998
1996-2004
1995
1994-2000
1994
1993-2003
1993-2001
1991-92
1989-92
1988-90
19871987-99
1987-94
1985-89
1984-2010:
1984
Norwegian Research Council, Section of Environment, member
IPCC, Chapter on Polar Climate, referee
Arctic Systems Inventory Analysis (ACIA), contributing author
Scientific advisory board, Inst. Meereskunde, Univ. Kiel
Arctic Climate Impact Assessment (ACIA) marine, 2000-2004
VISTA programme (Statoil/Academy of Science and Letters, Oslo), steering
board
Working group on overgrazing of kelp forests (Ministry of Fisheries), chairman
Marine Science Board, European Science Foundation
Conference on Marine Environment in North West of Russia (MAREN), St.
Petersburg, Russia, deputy chairman of Science Committee; NTNU delegate in
1999 and 2000:
Scientific Committee of Antarctic Research (SCAR) Working Group on
Biology
1st Nordic Marine Sciences Meeting, Gothenburg. Initiator and chairman of
planning committee Gothenburg, March 1995; 2nd Nordic Sciences Meeting,
Hirtshals, 1999; planning committee
Joint Global Ocean Flux Study, Scientific Steering Committee, executive member
1995-2000; Task Team on Photosynthetic Measurements 1993-1999, chairman
International Committee for Evaluation of Finnish Antarctic Research, chairman
Strategic Programme for Marine Science and Technology (MODTEQ, at NTNU,
funded by NFR); chairman
University Studies at Svalbard (UNIS), science committee; chairman of the
Biology Committee 1998-2001
International Committee for Evaluation of Swedish Polar Research (Forskningsrådsnämden, Stockholm)
Research Program for North-Norwegian Coastal Ecology (Mare Nor)
Research programmes Mare Nor, Causes of the Occurrence of Harmful
Phytoplankton Blooms, and Marine Pollution", member of planning committees
Albert Einstein prize (Consejo Cultural Mundial, Mexico), Kyoto Prize, Japan
Prize, Water Prize (Stockholm), Volvo Prize, nominator
The Museum of Natural History and Archaeology, NTNU, member of the
executive board
Scientific Committee for Antarctic Research (SCAR), Group of Specialists on the
Ecology of the Southern Ocean, member
APPRISE (field programme in Auke Bay, Alaska; Univ. Alaska, Oceanic Inst.
Hawaii), science advisory group
Editorial board, J. Plankton Research, European J. of Phycology (1993-)
Norwegian Antarctic Expedition of 1985-1986, member of steering committee
1983-1991
1982
1978-83
1976-1980
1974-85
1971-85
1971-81
1971-72
Pro Mare (Norwegian Research Programme for Marine Arctic Ecology,
chairman
Writing of the comprehensive plan for Pro Mare (Norwegian Research
Programme for Marine Arctic Ecology), with Bjørn Gulliksen,
Working group on biological decomposition and effects on micro-organisms,
FOH (Research Programme on Oil Pollution), chairman
Steering committee, Norwegian Coastal Current Programme (NFR)
Budget and Planning Committee, the Museum, member 1974-1985; chairman
1979-1985
Norwegian Oceanographic Committee
Central Budget and Planning Committee of the University of Trondheim,
chairman 1980-1981
Plankton committee, ICES
Consultant/committee member for promotions and appointments (tenure) at research institutions
and universities: Norway (16), USA (16), Germany (2), Sweden (1)
Opponent, Ph.D (15; Norway, Sweden, Germany, France)
Numerous evaluations of research proposals to research proposals: Norway, USA, Europe
Convener of 3 symposia (University of Alaska, Pro Mare, Trondheim, JGOFS Photosynthesis
Task Team, Svalbard)
Publications: (citations until 2010: H/I Index 26)
Publications are listed in three categories: refereed papers and review articles (series A), nonrefereed and popularized science (series B), and books (papers) I have edited (series C).
Science Citation Index (1968-2010): 2900; H/I Index 26
Short version, chronological (2005-2009:
Sakshaug, E. and Johnsen, G. 2005. Absorption, fluorescence excitation and
photoacclimation. Pp. 685-714 in Subba Rao, D.D. (ed) Algal cultures – Analogues of
blooms and applications. Science Publishers, Plymouth U.K., 971 p (2 vols)
Chauton, M. S, Johnsen, G., Sakshaug, E. 2005. Nuclear Magnetic Resonance spectroscopy of
marine microalgae: Metabolic profiling and species discrimination from Highresolution Magic Angle Spinning NMR analysis of whole-cell samples. Ph.D thesis,
113 p, NTNU Trondheim
Sakshaug, Egil; Johnsen, Geir. 2005. Algal cultures, Analogues of Blooms and
Applications. Multi-Science Publishing (ISBN 1-57808-393-1), 27 s.
Carmack, E., Barber, D., Christensen, J., Macdonald, R., Rudels, B., Sakshaug, E. 2006.
Climate variability and physical forcing of the food webs and the carbon budget on
panarctic shelves. Prog Oceanogr; 71: 145-181.
Olsen, L.M., Öztürk, M., Sakshaug, Egi., Johnsen G. 2006. Photosynthesis-induced phosphate
precipitation in seawater: ecological implications for phytoplankton. Marine Ecology
Progress Series 2006 ;Volum 319:103-110.
Hancke, K., Johnsen, G., Sakshaug, E. 2007. Bio-optical properties and spectral attenuation of
photosynthetically available radiation: relation to primary productivity and
phytoplankton composition. In Proc.Arctic Frontiers: balancing human use and
ecosystem protection. University of Tromsø
Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2007. Rate of O2 production from PAM
fluorescence: Testing three bio-optical approaches against measured O2 production
rate. AquaFluo Conference, Tromsø, 28 May-June 2007
Johnsen, G., Sakshaug, E 2007. Application of in vivo fluorescence excitation spectra from
different pigment-groups of phytoplankton. AquaFluo meeting; 2007-05-31,
NTNU
Johnsen, G., Sakshaug, E. 2007. Biooptical characteristics of PSII and PSI in 33 species (13
pigment groups) of marine phytoplankton, and the relevance for pulse-amplitudemodulated and fast-repetition-rate fluorometry. J. Phycol. 43: 1236-1251
Johnsen, G., Volent, Z., Sigernes, F., Berge, J., Hovland, E.K., Sakshaug, E. 2007
Local hyperspectral imaging of ocean colour from tethered balloons in the Arctic.
Proc. Arctic Frontiers Meeting: Balancing human use and ecosystem protection. :
University of Tromsø
Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2008. Rate of O2 production derived
from pulse-amplitude-modulated fluorescence: Testing three biooptical approaches
against measured O2-production rate. J. Phycol. 44: 803-813.
Fox, C., Harris, R., Sundby, S., Achterberg, E., Allen, J.I., Allen, J., Baker, A, Brussaard,
C.P.D; Buckley, P; Cook, E.J; Dye, S.R.; Edwards, M; Fernand, L; Kershaw, P;
Metcalfe, J; Osterhus, S; Potter, T; Sakshaug, Egil; Speirs, D.; Stenevik, E; John,
M.S.; Thingstad, F., Wilson, B. 2009. Transregional linkages in the north-eastern
Atlantic – an ‘end-to-end’ analysis of pelagic ecosystems. Oceanogr. Mar. Biol. Ann.
Rev. 47:1-75.
Sakshaug, E. 2009. Egil Sakshaugs Saga. In: Å forstå livet. Institutt for biologi NTNU - 50 år.
De biologiske fagenes historie i Trondheim og ved NTNU. Trondheim: Institutt for
biologi, NTNU 2009 ISBN 82-471-6062-5. p. 88-101.
Sakshaug, E., Kovacs, K., Johnsen, G. (eds) 2009. Ecosystem Barents Sea. Tapir Academic
Publishers, Trondheim. ISBN 978-82-519-2461-0, 587 p.
Sakshaug, E., Johnsen, G., Kristiansen, S., von Quillfeldt, C., Rey, F., Slagstad, D.,
Thingstad, F. 2009. Phytoplankton and primary production. In: Ecosystem Barents
Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 167-208
Sakshaug, E., Johnsen, G., Volent, Z. 2009. Light. In: Ecosystem Barents Sea. Tapir
Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 117-138
Johnsen, G., Volent, Z., Sakshaug, E., Sigernes, F., Pettersson, Lasse H. 2009. Remote
sensing in the Barents Sea. In: Ecosystem Barents Sea. Tapir Academic Publishers,
Trondheim. ISBN 978-82-519-2461-0. p. 139-166
Sakshaug, E., Kovacs, K.M. 2009. Introduction. I: Ecosystem Barents Sea. Tapir
Academic Publishers, Trondheim. ISBN 978-82-519-2461-0.p. 9-32
Sakshaug, E., Rey, F., Ardelan, M.V. 2009. Seawater, its constituents and chemistry. In:
Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-5192461-0. s. 83-116
von Quillfeldt, C., Hegseth, E.N., Johnsen, G. Sakshaug, E. Syvertsen, E.E. 2009. Ice algae.
In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82519-2461-0. s. 285-302
FULL LIST of LITERATURE
A. Refereed Papers and Review Articles
1A. Sakshaug, E. 1970. Quantitative phytoplankton investigations in near-shore water
masses. K. Norske Vidensk. Selsk. Skr. 1970 (3), 8pp.
2A. Jensen, A. and Sakshaug, E. 1970. Producer-consumer relationships in the sea. I. Preliminary studies on phytoplankton density and Mytilus pigmentation. J. Exp. Mar. Biol. Ecol.
5, 180-186.
3A. Jensen, A. Sakshaug, E. 1970. Producer-consumer relationships in the sea. I. Correlation between Mytilus pigmentation and the density and composition of phytoplankton
populations in inshore waters. J. Exp. Mar. Biol. Ecol. 5, 246-253.
4A. Sakshaug, E. and Jensen, A. 1971. Gonyaulax tamarensis and paralytic mussel toxicity in
Trondheimsfjorden, 1963-1969. Kgl. Norske Vidensk. Selsk. Skr. 1972 (15), 15 pp.
5A. Sakshaug, E., 1972. Phytoplankton investigations in Trondheimsfjord, 1963-1966. K. Norske
Vidensk. Selsk. Skr. 1972 (1), 56 pp.
6A. Haug, A., Myklestad, S. and Sakshaug, E. 1973. Studies on the phytoplankton ecology of the
Trondheimsfjord. I. The chemical composition of phytoplankton populations. J. Exp.
Mar. Biol. Ecol. 11, 15-26.
7A. Jensen, A. and Sakshaug, E. 1973. Studies on the phytoplankton ecology of the Trondheimsfjord. II. Chloroplast pigments in relation to abundance and physiological state of
the phytoplankton. J. Exp. Mar. Biol. Ecol. 11, 137-155.
8A. Sakshaug, E. and Myklestad, S. 1973: Studies on the phytoplankton ecology of the Trondheimsfjord. III. Dynamics of phytoplankton blooms in relation to environmental factors,
bioassay experiments and parameters for the physiological state of the populations. J.
Exp. Mar. Biol. Ecol. 11, 157-188.
9A. Sakshaug, E., Myklestad, S., Krogh, T. and Westin, G. 1973: Production of protein and
carbohydrate in the dinoflagellate Amphidinium carteri. Some preliminary results. Norw.
J. Bot. 20, 211-218.
10A. Sakshaug, E. 1976. Dynamics of phytoplankton blooms in Norwegian fjords and coastal
waters. Pp. 139-143 in Skreslet, S., Leinebø, R. and Sakshaug, E. (eds), Freshwater on
the sea. Norwegian Association of Oceanographers, Oslo.
11A. Sakshaug, E. 1977. Limiting nutrients and maximum growth rates for diatoms in Narragansett Bay. J. Exp. Mar. Biol. Ecol. 28, 109-123.
12A. Sakshaug, E. and Holm-Hansen, O. 1977. Chemical composition of Skeletonema costatum
(Grev.) Cleve and Pavlova lutheri (Droop)Green as a function of nitrate-, phosphate-,
and iron-limited growth. J. Exp. Mar. Biol. Ecol. 29, 1-34.
13A. Sakshaug, E. 1978. The influence of environmental factors on the chemical composition of
cultivated and natural populations of marine phytoplankton. Dr. philos. thesis,
University of Trondheim, 83 pp.
14A. Sakshaug, E. and Jensen, A. 1978: The use of cage cultures in studies of the biochemistry
and ecology of marine phytoplankton. Oceanogr. Mar. Biol. Ann. Rev. 16, 81-106.
15A. Sakshaug, E. 1980: Problems in the methodology of studying phytoplankton. Pp. 57-91 in
Morris, I. (ed.): The physiological ecology of phytoplankton. Blackwell Scientific
Publications, 625 pp.
16A. Bell, W.H. and Sakshaug, E. 1980: Bacterial utilization of algal extracellular products. II. A
kinetics study on natural populations. Limnol. Oceanogr. 25, 1021-1033.
17A. Sakshaug, E., Myklestad, S., Andresen, K., Hegseth, E.N. and Jørgensen, L. 1981
Phytoplankton off the Møre Coast in 1975-1976: distribution, species composition,
chemical composition and conditions for growth. Pp. 681-711 (vol. 2) in Sætre, R. and
Mork, M. (eds), The Norwegian Coastal Current, University of Bergen, 795 pp (vols 1 &
2).
18A. Sakshaug, E., Andresen, K., Myklestad, S. and Olsen, Y. 1983: Nutrient status of
phytoplankton communities in Norwegian waters (marine, brackish, and fresh) as
revealed by their chemical composition. J. Plankton Res. 5, 175-196.
19A. Hegseth, E. and Sakshaug, E. 1983. Seasonal variation in light- and temperature-dependent
growth of marine planktonic diatoms in in situ dialysis cultures in the Trondheimsfjord,
Norway (63N). J. Exp. Mar. Biol. Ecol. 67, 199-220.
20A. Myklestad, S. and Sakshaug, E. 1983: Alkaline phosphatase activity of Skeletonema
costatum populations in the Trondheimsfjord. J. Plankton Res. 5, 557-564.
21A. Sakshaug, E., Granéli, E., Elbrächter, M. and Kayser, H. 1984. Chemical composition and
alkaline phosphatase activity of nutrient-saturated and P-deficient cells of four marine
dinoflagellates. J. Exp. Mar. Biol. Ecol. 84, 241-254.
22A. Sakshaug, E. and Holm-Hansen, O. 1984. Factors governing pelagic production in polar
oceans. Pp. 1-18 in Holm-Hansen, O., Bolis, L. and Gilles, R. (eds), Marine
phytoplankton and productivity. Lecture notes on coastal and estuarine studies, vol. 8.
Springer Verlag.
23A. Hewes, C.D., Holm-Hansen, O. and Sakshaug, E. 1985. Alternate carbon pathways at
lower trophic levels in the Antarctic food web. Pp. 277-283 in Siegfried, W.R., Condy,
P.R. and Laws, R.M. (eds), Antarctic nutrient cycles and food webs, Springer Verlag,
700 pp.
24A. Sakshaug, E. and Olsen, Y. 1986: Nutrient status of phytoplankton blooms in Norwegian
waters and algal strategies for nutrient competition. Can. J. Fish. Aq. Sci. 43, 389-396.
25A. Sakshaug, E. and Holm-Hansen, O. 1986. Photoadaptation in Antarctic phytoplankton:
variations in growth rate, chemical composition and P versus I curves. J. Plankton Res. 8,
459-473.
26A. Sakshaug, E. and Andresen, K. 1986. Effect of light regime upon growth rate and chemical
composition of a clone of Skeletonema costatum from the Trondheimsfjord, Norway. J.
Plankton Res. 8, 619-638.
27A. Sakshaug, E., Demers, S. and Yentsch, C.M. 1987. Thalassiosira oceanica and T.
seudonana: two different photoadaptational responses. Mar. Ecol. Prog. Ser. 41, 275282.
28A. Sakshaug, E., Andresen, K. and Kiefer, D.A. 1989. A steady-state model of growth and
light absorption in the marine plankton diatom Skeletonema costatum. Limnol. Oceanogr.
34, 198-205.
29A. Sakshaug, E. and Skjoldal, H.R. 1989. Life at the ice edge. Ambio 18, 60-67.
30A. Sakshaug, E. 1989. The physiological ecology of polar phytoplankton. Pp. 61-89 in Rey, L.
and Alexander, V. (eds): Marine living systems of the far North (6th Conf. Comité
Arctique Int., Fairbanks, Alaska), E.J. Brill, Leiden, The Netherlands. 637 pp.
31A. Hewes, C.D., Sakshaug, E.H., Reid, F.M.H. and Holm-Hansen, O. 1990. Microbial
autotrophic and heterotrophic eucaryotes in Antarctic waters: Relationships between
biomass and chlorophyll, adenosine triphosphate and particulate organic carbon. Mar.
Ecol. Prog. Ser. 63, 27-35.
32A. Thingstad, T.F. and Sakshaug, E. 1990. Control of phytoplankton growth in nutrient
recycling ecosystems. Theory and terminology. Mar. Ecol. Prog. Ser. 63, 261-272.
33A. Gilstad, M. and Sakshaug, E. 1990. Growth rates of ten diatom species from the Barents
Sea at different irradiances and day lengths. Mar. Ecol. Prog. Ser. 64, 169-173.
34A. Sakshaug, E. 1990. Arctic planktonic ecosystems. Pp. 47-51 in Priddle, J. and Medlin, L.
(eds) Polar diatoms - a manual, British Antarctic Survey, Cambridge.
35A. Smith, W.O. and Sakshaug, E. 1990. Primary production in Polar Oceans. Pp. 477-525 in
Smith, W.O. (ed.) Polar Oceanography, part B. Academic Press, 760 pp (2 vols).
36A. Sakshaug, E., Johnsen, G., Andresen, K. and Vernet, M. 1991. Modeling of lightdependent algal photosynthesis and growth: Experiments with the Barents Sea diatoms
Thalassiosira nordenskioeldii and Chaetoceros furcellatus. Deep-Sea Res. 38, 415-430.
37A. Sakshaug, E. 1991. Food webs and primary production in the Barents Sea. Proc. 12th NIPR
Symp. Polar Biol. (Tokyo) 4, 1-8.
38A. Sakshaug, E. and Slagstad, D. 1991. Factors controlling the development of phytoplankton blooms in the Antarctic Ocean - A mathematical model. Mar. Chem. 35, 259272.
39A. Sakshaug, E. and Slagstad, D. 1991. Light and productivity of phytoplankton in polar
marine ecosystems. Polar Res. 10, 69-85.
40A. Verity, P.G., Smayda, T.J. and Sakshaug, E. 1991. Photosynthesis, excretion, and growth
rates of Phaeocystis colonies and solitary cells. Polar Res. 10, 117-128.
41A. Johnsen, G., Sakshaug, E. and Vernet, M. 1992. Pigment composition, spectral
characterization and photosynthetic parameters in Chrysochromulina polylepis. Mar.
Ecol. Prog. Ser. 83, 241-249.
42A. Sakshaug, E. Slagstad, D. 1992. Sea-ice and wind: Effects on primary production in the
Barents Sea. Atmosphere-Ocean 30, 579-571.
43A. Gilstad, M., Johnsen, G. and Sakshaug, E. 1993. Diurnal- and daylength-dependent
variations in photosynthetic parameters, pigment composition and the respiration rate of
Skeletonema costatum. J. Plank. Res. 15, 939-951.
44A. Sakshaug, E. 1993. The relationship between phytoplankton growth rate and production
with emphasis on respiration and excretion. ICES Mar. Symp. Ser. 197, 63-68.
45A. Nielsen, M. and Sakshaug, E. 1993. Adaptation to spectrally different light regimes:
Studies of the growth rate and Carbon to Chlorophyll a ratio of the marine diatom
Skeletonema costatum. Limnol. Oceanogr. 38, 1576-1581.
46A. Johnsen, G. and Sakshaug, E. 1993. Bio-optical characteristics and photoadaptive
responses in the toxic and bloom-forming dinoflagellates Gyrodinium aureolum,
Gymnodinium galatheanum, and two strains of Prorocentrum minimum. J. Phycol. 29,
627-642.
47A. Cullen, J.J., Geider, R.J., Ishizaka, J., Kiefer, D.A., Marra, J., Sakshaug, E. and Raven, J.A.
1993. Toward a general model of phytoplankton growth for biogeochemical models. Pp
153-176 in Evans, G.T. and Fasham, M.J.R. (eds): Towards a model of ocean
biogeochemical processes. NATO ASI Series I, vol. 10, Springer-Verlag, Berlin,
Heidelberg, 350 pp.
48A. Sakshaug, E. 1993. Variability in photosynthetic parameters. Pp. 2-10 in Eilertsen, H.C.
(ed.) Proc. Europto Series, SPIE 2048, 190 pp.
49A. Johnsen, G., Samseth, O., Granskog, L. and Sakshaug, E. 1993. In vivo absorption
characteristics in 10 classes of phytoplankton: Taxonomic characteristics and responses
to photoadaptation by means of discriminant and HPLC analysis. Mar. Ecol. Prog. Ser.
105, 149-157.
50A. Baumann, M.E.M., Lancelot, C., Brandini, F.P., Sakshaug, E. and John, D.M. 1994. The
taxonomic identity of the cosmopolitan prymnesiophyte Phaeocystis: a morphological
and ecophysiological approach. J. Mar. Systems 5, 5-22.
51A. Sakshaug, E, Bjørge, A., Gulliksen, B., Loeng, H. and Mehlum, F. 1994. Structure,
biomass distribution, and energetics of the pelagic ecosystem in the Barents Sea: A
synopsis. Polar Biol. 14, 405-411.
52A. Dalløkken, R., Sandvik, R. and Sakshaug, E. 1994. Variations in bio-optical properties in
the Greenland/Iceland/Norwegian Seas. Ocean Optics 12. SPIE 2258, 266-276.
53A. Dalløkken, R., Sandvik, R. and Sakshaug, E. 1995. Seasonal variations in the vertical light
attenuation coefficient in the Greenland Sea: Effect of phytoplankton light absorption.
Pp. 33-43 in Skjoldal, H.R., Hopkins, C.C.E., Erikstad, K.E. and Leinaas, H.P. (eds).
Ecology of fjords and coastal waters. Elsevier Science, Amsterdam.
54A. Sakshaug, E., Rey, F. and Slagstad, D. 1995. Wind forcing of marine primary productivity
in the atmospheric low-pressure belts. Pp. 15-25 in Skjoldal, H.R., Hopkins, C.C.E.,
Erikstad, K.E. and Leinaas, H.P. (eds). Ecology of fjords and coastal waters. Elsevier
Science, Amsterdam.
55A. Johnsen, G. and Sakshaug, E. 1995. Light harvesting in bloom-forming marine
phytoplankton: species-specificity and photoadaptation. Scientia Marina 60 (Suppl.), 4756.
56A. Vernet, M., Mitchell, B.G., Sakshaug, E., Johnsen, G., Iturriaga, R. and Wassmann, P.
1996. Evidence for a novel pigment with in vivo absorption maximum at 708 nm
associated with Phaeocystis cf. pouchetii blooms. Mar. Ecol. Prog. Ser. 133, 253-262.
57A. Sakshaug, E. 1996. Biomass, productivity distributions and their variability in the Barents
Sea. ICES mar. Sci. Symp. 54, 341-350.
58A. Grzymski, J., Johnsen, G. and Sakshaug, E. 1997. The significance of intracellular selfshading on the bio-optical properties of brown, red and green macroalgal species. J.
Phycol. 33, 408-414.
59A. Sakshaug, E., Bricaud, A., Dandonneau, Y., Falkowski, P.G., Kiefer, D.A., Legendre, L.,
Morel, A., Parslow, J. and Takahashi, M. 1997. Parameters of photosynthesis:
definitions, theory and interpretation of results. J. Plank. Res. 19, 1637-1670.
60A. Johnsen, G., Volent, Z., Tangen, K. and Sakshaug, E. 1997. Time series of harmful and
benign phytoplankton blooms in northwest European waters using the Seawatch buoy
system. Pp. 115-143 in Kahru, M. and Brown, C.W. (eds). Monitoring algal blooms:
New techniques for detecting large-scale environmental change. Landes Bioscience,
New York.
61A. Johnsen, G., Samset, O. and Sakshaug, E., 1997. Bio-optical classification of potential
harmful chlorophyll c3-containing prymnesiophyte blooms in Scandinavian waters.
62A. Sakshaug, E. 2000. Wind forcing of marine primary production in the northern belt of
westerlies. Pp. 19-36 in Hanson, R., Ducklow, H. and Field. J.G. (eds). The changing
ocean carbon cycle. IGBP Series 5, Cambridge Univeristy Press, Cambridge, U.K., 514
p.
63A. Sakshaug, E. and Walsh, J. 2000. Marine biology: Biomass, productivity distributions and
their variability in the Barents and Bering Seas. Pp. 161-195 in Nuttall, M. and
Callaghan, T.V. The Arctic: a guide to research in the natural and social sciences.
Harwood, London.
64A. Johnsen, G. and Sakshaug, E. 2000. Monitoring of harmful algal blooms along the
Norwegian coast. S. Afr. J. Mar. Sci. 22, 309-321.
65A. Aas E, Højerslev NK, Sandvik R, Sakshaug E 2002. Spectral properties and UVattenuation in Arctic marine waters. Pp. 23-56 in Hessen, D (ed) Radiation and Arctic
ecosystems. Springer, Heidelberg-Berlin.
66A. Özturk, M., Steinnes, E. And Sakshaug, E. 2002. Iron speciation in the Trondheim Fjord
from the perspective of iron limitation for phytoplankton. Est. Coastal Shelf Sci 55, 197212.
67A. Sakshaug, Egil (2003). Primary production in Arctic and subarctic waters. Pp 57-81 in
Stein R, Macdonald R (eds) The organic arctic arbon cycle. Springer, BerlinHeidelberg, 363 p.
68A. Macdonald, RW, Sakshaug, E and Stein, R. 2003: The Arctic Ocean: Modern Status
and climate change. Pp. 6-21 in Stein, R. and Macdonald RW (eds): The organic
carbon cycle in the Arctic Ocean. Springer, 363 p. 70A.
69A. Öztürk, M., Vadstein, O. and Sakshaug, E. 2003. The effects of enhanced phytoplankton
production on iron speciation and removal in mesocosm experiments in a landlocked
basin of Hopavågen, Norway. Mar. Chem. 84, 3-17.
70A. Öztürk, M., Croot, P., Bertilsson, S., Abrahamsson, K., Karlson, B., Davis, R., Fransson,
A. and Sakshaug, E. 2004. Iron enrichment and photoreduction of iron under UV and
PAR in the presence of hydrocarboxylic acid: Implication for4 phytoplankton growth in
the Southern Ocean. Deep-Sea Res. II 51, 2841-2856
71A. Sakshaug, E. and Johnsen, G. 2005. Absorption, fluorescence excitation and
photoacclimation. Pp. 685-714 in Subba Rao, D.D. (ed) Algal cultures – Analogues of
blooms and applications. Science Publishers, Plymouth U.K., 971 p (2 vols)
72A. Chauton, M. S, Johnsen, G., Sakshaug, E. 2005. Nuclear Magnetic Resonance
spectroscopy of marine microalgae: Metabolic profiling and species discrimination
from High-resolution Magic Angle Spinning NMR analysis of whole-cell samples.
Ph.D thesis, 113 p, NTNU Trondheim
73A. Sakshaug, Egil; Johnsen, Geir. 2005. Algal cultures, Analogues of Blooms and
Applications. Multi-Science Publishing (ISBN 1-57808-393-1), 27 s.
74A Carmack, E., Barber, D., Christensen, J., Macdonald, R., Rudels, B., Sakshaug, E. 2006.
Climate variability and physical forcing of the food webs and the carbon budget on
panarctic shelves. Prog Oceanogr; 71: 145-181.
75A. Olsen, L.M., Öztürk, M., Sakshaug, Egi., Johnsen G. 2006. Photosynthesis-induced
phosphate precipitation in seawater: ecological implications for phytoplankton. Marine
Ecology Progress Series 2006 ;Volum 319:103-110.
76A. Hancke, K., Johnsen, G., Sakshaug, E. 2007. Bio-optical properties and spectral
attenuation of photosynthetically available radiation: relation to primary productivity
and phytoplankton composition. In Proc.Arctic Frontiers: balancing human use and
ecosystem protection. University of Tromsø
77A. Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2007. Rate of O2 production from
PAM fluorescence: Testing three bio-optical approaches against measured O2
production rate. AquaFluo Conference, Tromsø, 28 May-June 2007
78A. Johnsen, G., Sakshaug, E 2007. Application of in vivo fluorescence excitation spectra
from different pigment-groups of phytoplankton. AquaFluo meeting; 2007-05-31,
NTNU
79A. Johnsen, G., Sakshaug, E. 2007. Biooptical characteristics of PSII and PSI in 33 species
(13 pigment groups) of marine phytoplankton, and the relevance for pulse-amplitudemodulated and fast-repetition-rate fluorometry. J. Phycol. 43: 1236-1251
80A. Johnsen, G., Volent, Z., Sigernes, F., Berge, J., Hovland, E.K., Sakshaug, E. 2007
Local hyperspectral imaging of ocean colour from tethered balloons in the Arctic.
Proc. Arctic Frontiers Meeting: Balancing human use and ecosystem protection. :
University of Tromsø
81A. Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2008. Rate of O2 production
derived from pulse-amplitude-modulated fluorescence: Testing three biooptical
approaches against measured O2-production rate. J. Phycol. 44: 803-813.
82A. Fox, C., Harris, R., Sundby, S., Achterberg, E., Allen, J.I., Allen, J., Baker, A,
Brussaard, C.P.D; Buckley, P; Cook, E.J; Dye, S.R.; Edwards, M; Fernand, L;
Kershaw, P; Metcalfe, J; Osterhus, S; Potter, T; Sakshaug, Egil; Speirs, D.; Stenevik,
E; John, M.S.; Thingstad, F., Wilson, B. 2009. Transregional linkages in the northeastern Atlantic – an ‘end-to-end’ analysis of pelagic ecosystems. Oceanogr. Mar.
Biol. Ann. Rev. 47:1-75.
83A. Sakshaug, E. 2009. Egil Sakshaugs Saga. In: Å forstå livet. Institutt for biologi NTNU 50 år. De biologiske fagenes historie i Trondheim og ved NTNU. Trondheim: Institutt
for biologi, NTNU 2009 ISBN 82-471-6062-5. p. 88-101.
84A. Sakshaug, E., Kovacs, K., Johnsen, G. (eds) 2009. Ecosystem Barents Sea. Tapir
Academic Publishers, Trondheim. ISBN 978-82-519-2461-0, 587 p.
85A. Sakshaug, E., Johnsen, G., Kristiansen, S., von Quillfeldt, C., Rey, F., Slagstad, D.,
Thingstad, F. 2009. Phytoplankton and primary production. In: Ecosystem Barents
Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 167-208
86A. Sakshaug, E., Johnsen, G., Volent, Z. 2009. Light. In: Ecosystem Barents Sea. Tapir
Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 117-138
87A. Johnsen, G., Volent, Z., Sakshaug, E., Sigernes, F., Pettersson, Lasse H. 2009. Remote
sensing in the Barents Sea. In: Ecosystem Barents Sea. Tapir Academic Publishers,
Trondheim. ISBN 978-82-519-2461-0. p. 139-166
88A. Sakshaug, E., Kovacs, K.M. 2009. Introduction. I: Ecosystem Barents Sea. Tapir
Academic Publishers, Trondheim. ISBN 978-82-519-2461-0.p. 9-32
89A. Sakshaug, E., Rey, F., Ardelan, M.V. 2009. Seawater, its constituents and chemistry. In:
Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-5192461-0. s. 83-116
90A. von Quillfeldt, C., Hegseth, E.N., Johnsen, G. Sakshaug, E. Syvertsen, E.E. 2009. Ice
algae. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 97882-519-2461-0. s. 285-302
B. Popular and non-refereed publications
1B. Sakshaug, E. 1969. Planteplankton (Phytoplankton). Pp. 22-39 in Strømgren, T. (ed.): The
Borgenfjord Investigations, Preliminary report 1968. The Royal Norwegian Society of
Sciences and Letters, The Museum, Trondheim (in Norwegian).
2B. Sakshaug, E. and Jensen, A. 1971. Gonyaulax tamarensis og paralytisk muslinggift i
Trondheimsfjorden, 1970-1971 (Gonyaulax tamarensis and paralytic mussel toxicity in
the Trondheimsfjord, 1970-1971). Pp. 14-20 in Strømgren, T. (ed.): The Borgenfjord
Investigations, Preliminary report 1970. The Royal Norwegian Society of Sciences and
Letters, The Museum, Trondheim (in Norwegian).
3B. Sakshaug, E., Jensen, A. and Prakash, A. 1971. Gonyaulax tamarensis - the causative
organism of mussel toxicity in Trondheimsfjord. ICES C.M. 1971, Helsinki. Report no.
L:14. 13 pp.
4B. Sakshaug, E. and Haug, A. 1972. Forurensinger i marint miljø (Pollution in the marine
environment). Pp. 125-142 in Carlsen, K., Celius, H., Halmø, T. and Nestaas, I. (eds)
Miljøkunnskap (Environmental Science). Tapir, Trondheim. 308 pp (in Norwegian).
5B. Sakshaug, E., Haug, A., Jensen, A. and Myklestad, S. 1972. Phytoplankton ecology of the
Trondheimsfjord. ICES, C.M. 1972, Copenhagen. Report no. L:13, 20 pp.
6B. Sakshaug, E. and Brun, P. 1974. Plantenæringsstoffer og planteplankton i Trondheimsfjorden (Plant nutrients and phytoplankton in the Trondheimsfjord). Vann 1974 (3)
1-12.
7B. Sakshaug, E. 1975. Noen tanker om modeller for akvatisk produktivitet (Some thoughts on
models for aquatic productivity). Pp. 195-211 in Rept 10th Nordic Symp. Water Res.,
Værløse, Denmark, May 1974. Nordforsk Publication 1975 (1). 559 pp.
8B. Sakshaug, E. 1977. Plantenæringsstoffer og klorofyll a (Plant nutrients and chlorophyll a).
Pp. 134-151 in Lande, E. (ed.): Resipientundersøkelsen av Trondheimsfjorden, 1972-75
(The Trondheimsfjord Investigations in 1972-75). The Royal Norwegian Society of
Sciences and Letters, The Museum, Rept Zool. Series 1977 (11), in Norwegian.
9B. Sakshaug, E. 1978: Trondhjem Biologiske Stasjon - en god historie (Trondhjem Biological
Station - a good story). Akvarie-Tanken, no 3. The Royal Norwegian Society of
Sciences and Letters, The Museum, 16 pp (in Norwegian).
10B. Sakshaug, E. and Andersen, B. 1980: Norsk oseanografi - fortid og fremtid (Norwegian
Oceanography - past and future), Forskningsnytt 1980 (1), 26-28 (in Norwegian).
11B. Sakshaug, E. 1980. Organiske partikler i sjøvann (Organic particles in seawater). Naturen
(2), 29-40 (in Norwegian).
12B. Paden, C.A., Hewes, C.D., Neori, A., Holm-Hansen, O., Weaver, E., Kiefer, D.A. and
Sakshaug, E. 1981. Phytoplankton studies in the Scotia Sea. Ant. J. U.S. 16(5), 163.
13B. Sakshaug, E. 1982. Vassdragsregulering og liv i hav (Regulation of rivers and the effect on
life in the sea). Trøndersk Natur 1982 (1), 8-12 (in Norwegian).
14B. Sakshaug, E., 1982. Krill - mat i mengder i Sørishavet (Krill - food in abundance in the
Southern Ocean). Syn og Segn, 88, 482-489 (in Norwegian).
15B. Hewes, C.D., Holm-Hansen, O. and Sakshaug, E. 1983. Nanoplankton and microplankton
studies during the circumnavigation cruise. Ant. J. U.S. 1983, 169-171.
16B. Legendre, L., Collos, Y., Elbrächter, M., Fasham, M.J.R., Gieskes, W.W.C., Herbland, A.,
Holligan, P.M., Margalef, R., Perry, M.J., Platt, T., Sakshaug, E. and Smith, D.F. 1984.
Studies on marine autotrophs: Recommendations for the 1980s. Pp. 629-639 in Fasham,
M.J.R. (ed.), Flows of energy and materials in marine ecosystems. Plenum Press.
17B. Sakshaug, E. 1984. Going north - the need for ecological studies in the Barents and
Spitsbergen areas. Annual meeting, Norw. Petroleum Society, Bergen, May 1984.
18B. Sakshaug, E. 1986. Pro Mare - The Norwegian research program for marine arctic
ecology. Scandinavian Review 74, 44-47.
19B. Sakshaug, E. 1987. Fosfor og nitrogen som begrensende faktorer for algevekst Phosphorus and nitrogen as limiting factors for algal growth). Pp. 247-255 in: Rept 22nd Nordic
Symp. Water Res., Laugarvatn, Iceland, May 1986. Nordforsk Publication, Nature
Conservation Series, No. 1987(1), Helsinki.
20B. Wassmann, P. and Sakshaug, E. 1987. Barentshavets økologiske grunntrekk (Short review
of the ecology of the Barents Sea), Naturen 1987(4), 151-158 (in Norwegian).
21B. Sakshaug, E. 1988. Pro Mare 1984-1989. UnitNytt 18 (2), 8-10 (in Norwegian).
22B. Sakshaug, E. and Tangen, K. 1988. Tilførsel av plantenæringsstoffer: potensielle effekter
på algevekst (Plant nutrient supply: Potential effects on algal growth). Resipientundersøkelse for Trondheim, Delrapport 9 (Investigations of the Trondheim basin as a
recipient for sewage, Report no. 9). Oceanor Trondheim, (in Norwegian) 40 pp.
23B. Sakshaug, E. 1989. Spleiselag i Arktis. UnitNytt 19, 8-13.
24B. Sakshaug, E. 1990. Mot en ny istid? (Towards a new glacial age?). UnitNytt 20 (4), 3-4 and
18 (in Norwegian).
25B. Sakshaug, E. 1990. Marine ecology and climate. Compendium for Norsk Hydro A/S, 24 pp
(Norwegian and English edition).
26B. Sakshaug, E. and Tangen, K. 1990. The Chrysochromulina bloom in Skagerrak and
Kattegat summer 1988. OECD Environmental Report, 1990.
27B. Sakshaug, E., Johnsen, G., Samseth, O and Volent, Z. 1991. Identification of phytoplankton
blooms by means of remote sensing. Environment Northern Seas Meeting, Stavanger 28
August 1991, 91-100.
28B. Sakshaug, E. 1991. Den biologiske karbonpumpen: Primærproduksjon i havet (The
biological carbon pump: Primary production in the sea). Kjemi 52, 27-32 (in Norwegian;
English summary).
29B. Sakshaug, E. 1992. The Barents Sea: The physical-biological connection. ("Anton Bruun
Lecture" of IOC/UNESCO). Pp 23-26 In Bruun Memorial Lectures, 1991: Modelling
and prediction in marine science. IOC/UNESCO, Paris, 33 p.
30B. Sakshaug, E. 1992. Brukes "føre var"-prinsippet på en uforsvarlig måte? Biolog, 1992 (1),
18-26.
31B. Sakshaug, E., Bjørge, A., Gulliksen, B., Loeng, H. and Mehlum, F. 1992. Innledning
(Introduction). Pp. 11-19 in Økosystem Barentshavet (7C; in Norwegian).
32B. Sakshaug, E., Bjørge, A., Gulliksen, B., Loeng, H. and Mehlum, F. 1992. Epilog
(Epilogue). Pp. 275-285 in Økosystem Barentshavet (7C; in Norwegian).
33B. Sakshaug, E., Volent, Z., Kögeler, J. and Slagstad, D. 1992. Lys i polarstrøk (Light in polar
areas). Pp. 65-78 in Økosystem Barentshavet (7C; in Norwegian).
34B. Sakshaug, E., Kristiansen, S., Syvertsen, E., Eilertsen, H.-C., Gilstad, M., Hegseth, E.N.,
Hewes, C.D., Holm-Hansen, O., Johnsen, G., Mitchell, G.B., Rey, F., Slagstad, D., StøleHansen, K., Thingstad, T.F. and Vernet, M. 1992. Planteplankton (Phytoplankton). Pp.
79-108 in Økosystem Barentshavet (7C; in Norwegian).
35B. Rey, F., Sakshaug, E. and Myklestad, S. 1992. Sjøvannskjemi (Seawater chemistry). Pp.
54-63 in Økosystem Barentshavet (7C; in Norwegian).
36B. Slagstad, D., Sakshaug, E., Loeng, H., Tande, K and Ådlandsvik, B. 1992. Matematisk
modellering (Mathematical modelling). Pp. 250-262 in Økosystem Barentshavet (7C; in
Norwegian).
37B. Vinje, T., Gulliksen, B., Loeng, H., Hegseth, E.N., Sakshaug, E. and Syvertsen, E.S. 1992.
Havis (Sea ice). Pp. 125-126 in Økosystem Barentshavet (7C; in Norwegian).
38B. Sakshaug, E. 1993. Discussant's report: primary production in the Antarctic pelagial - a
view from the North. Proc. BIOMASS symposium, Bremerhaven, September 1990. p.
125-126.
39B. Sakshaug, E. 1995. A synopsis of the biomass distribution and energetics of the pelagic
ecosystem of the Barents Sea. ICES.CM 1995/Mini:3, 11 p.
40B. Sakshaug, E., Olsen, Y. and Hoell, E. 1995. Maricult research programme: Fertilising
marine ecosystems to harvest more. ICES.CM 1995/T:1, 16 p.
41B. Sakshaug, E. 1995. Weather and climate: effects on life in the sea. Kgl. Norske Vidensk.
Selsk. Forh. 1995, 103-123.
42B. Sakshaug, E. 1995. Biomass and biological production in Arctic seas. Proc. EuroMab
Conf., Kangerlussuaq, Greenland, September 1995 (submitted).
43B. Sakshaug, E. 1997. The Nordic region: cycles in climate and marine life. Pp. 40-51 in
Andresen, K.H., Arlov, T.B., Hagen, D., Løset, S., Reinertsen, R.E. and Thomassen, J.
(eds). Sustainable development in the Arctic. Consequences of industrial development in
the European Arctic region. Centre for environment and development, Norwegian
University of Science and Technology, Trondheim.
44B. Sakshaug, E. 1997. The nordic region: Cycles in climate and marine life. Forh. norske
vidensk.Akad.
45B. Sakshaug, E. 1997. Istid, ikke nå igjen (Ice age, not now again). Universitsavisa, 21 August
1997.
46B. Sakshaug, E. 1998. El Nino er her igjen (El Nino is here again). Universitetsavisa,
November 1998.
47B. Sakshaug, E. and Slagstad, D. 2000. Fjordens posisjon og solens gang )The location of the
fjord and the solar track). P. 18 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden.
Tapir, Trondheim, 336 p (in Norwegian).
48B. Sakshaug, E. 2000. Hvorfor istider? (Why ice ages?). Pp. 28-29 in Sakshaug, E. and J.-A.
Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).
49B. Sakshaug, E. 2000. Trondheimsfjorden og havet utenfor (The Trondheimsfjord and the sea
outside). Pp. 38-41 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir,
Trondheim, 336 p (in Norwegian).
50B. Sakshaug, E. 2000. Om salt, CO2 og organisk stoff i sjøen- Pp. 59-64 in Sakshaug, E. and
J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).
51B. Sakshaug, E. and Killingtveit, Å. 2000. Elvene: et stort bidrag (The rivers: a large
contribution. Pp. 65-75.
52B. Sakshaug, E. and Olsen, Y. 2000. Litt om økologi (A little about ecology). Pp. 76-82 in
Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in
Norwegian).
53B. Sakshaug, E. and Tangen, K. 2000. Lyset, næringssaltene og planteplanktonet (The light,
the nutrients and the phytoplankton). Pp. 83-95 in Sakshaug, E. and J.-A. Sneli (eds)
Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).
54B. Tangen, E. and Sakshaug, E. 2000. Skadelige alger (Harmful algae). Pp. 96-102 in
Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in
Norwegian).
55B. Tokle, N. and Sakshaug, E. 2000. Dyreplanktonet (The zooplankton). Pp. 103-109 in
Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in
Norwegian).
56B. Sakshaug, E. 2000. Matalger (Edible seaweeds). P. 164 in Sakshaug, E. and J.-A. Sneli
(eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).
57B. Sakshaug, E. 2000. Søl - sagaens alge (Palmaria palmata - the seaweed of the sagas). P.
167 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p
(in Norwegian).
58B. Sakshaug, E. and Sneli, J.-A. 2000. Trondhjem biologiske stasjon: en sprek hundreåring
(Trondhjem Biological Station: a vigorous centenarian). Pp. 316-318 in Sakshaug, E. and
J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).
59B. Sakshaug, E. 2000. Units. Pp. 319-322 in Sakshaug, E. and J.-A. Sneli (eds)
Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).
60B. Sakshaug, E., Christie, H., Dale, T., Fosså, J.H., Fredriksen, S., Hedlund, N. og Sivertsen,
K. 2002. Nedbeiting av tareskog i Norge. Fiskeridepartementet, Oslo, 47 p.
61B: Laws, E, Sakshaug, E., Babin, M., Dandonneau, Y., Falkowski, P., Geider, R., Legendre,
L., Morel, A., Sondergaard, M., Takahashi, M. And Williams, P. leB. 2002.
Photosynthesis and primary productivity in marine ecosystems. Practical aspects and
application of techniques. Joint Global Ocean Flux Study, Rapport 36; Univ. Bergen, 88
p.
62B: Sakshaug, E. 2002. Primary Production and marine climate in the Nordic and Arctic Seas.
IRISEN II (Integrated regional impact studies in the European North), Advanced Study
Courses, Abisko, Sverige, 26. juli 2002, 10 p (manuscript).
63B: Sakshaug, E. 2002. Phytoplankton. Advanced Nordic courses: Sensitivity of the
Westnordic Marine Ecosystem to past, present and future climate. Laugarvatn, Island, 19
august, 2 p (manuscript).
64B: Sakshaug, E. Phytoplankton. DKNVS Arctic Symposium, Kongsvoll, 23. august, 55 p
(manuscript)
65B: Sakshaug, E. 2002. Primary production and marine climate in Nordic and Arctic Seas.
Workshop, Havforskningsinstituttet, Thórshavn, Færøyane, 25. september, 9 p.
(manuscript).
66B: Sakshaug E. and Walsh, J.J. 2002. Productivity, distributions and their variability in the
Barents Sea and the Bering shelf: a summary. Univ. South Florida, St. Petersburg,
Florida, 18. oktober, 9 p (manuscript).
67B: Sakshaug, E. 2002. Variability in primary production in Arctic Seas. Climate workshop,
Nordic Arctic Research Programme, Sigulda, Latvia, 4. november, 35 p.
68B: Sakshaug E. 2002. Arctic primary production vs. climate variation. Dept Biology,
University of California Santa Barbara, 12. november (set of overhead copies)
69B. Holm-Hansen, O.and Sakshaug, E. 2002. Polar Marine Phytoplankton. Pp. 2554-2565
in Bitton, G. (ed.). Encyclopedia of Environmental Microbiology, vol. 5. John Wiley
& Sons, New York.
70B Sakshaug, E. 2004. Primary production and marine climate in the arctic seas. Pp. 283297 in Lange, M.A. and Pöszig, D. (eds) IRISEN II. Proc. Adv. Stud. Course, Abisko,
Sweden. Centre Environ. Res. Univ. Münster, Münster, 484 p.
Official reports:
1976: Norwegian Oceanography ─ Status and Perspectives (2C in the list of publications), the
Norwegian Oceanographic Committee (NAVF)
1979: Norwegian Oceanography ─ Recent Development and Perspectives, the Norwegian
Oceanographic Committee (NAVF)
1982 Comprehensive Program for the Norwegian Research Program for Marine Arctic
Ecology (Pro Mare, published by NFFR and NAVF
C. Books/Editorial work
1C. Skreslet, S., Leinebø, R., Matthews, J.B.L. and Sakshaug, E. 1976. Freshwater on the sea.
Proc. Symp. on the influence of fresh-water outflow on biological processes in fjords and
coastal waters, Geilo, Norway, April 1974. Norwegian Association of Oceanographers,
Oslo. 246 p.
2C. Sakshaug, E., Wedege, N.P. and Dahl, F.E. 1976. Norsk oseanografi - status og perspektiver (Norwegian Oceanography - status and perspectives). Norwegian Oceanographic Committee, Norwegian Research Council for Science and the Humanities, Oslo.
248 p (with English summary).
3C. Sakshaug, E. 1979: Norsk oseanografi - nyere utvikling og perspektiver (Norwegian
Oceanography - recent development and perspectives). Norwegian Oceanographic
Committee, Norwegian Research Council for Science and the Humanities, Oslo. 100 p.
4C. Sakshaug, E. and Gulliksen, B. 1983. Marinøkologisk forskningsprogram i Arktis:
rammeprogram (A comprehensive plan for the Norwegian Research Program for Marine
Arctic Ecology, Pro Mare). The Norwegian Fisheries Research Council, the Norwegian
Research Council for Science and the Humanities, Trondheim/Oslo. 55 p.
5C. Sakshaug, E. and Hegseth, E.N. 1984. Sluttrapport fra faggruppe IV: Biologisk nedbrytning
og effekter på mikroorganismer. II. Studier med planktonalger (Final report from
scientific committee IV: Biological degradation and effects on microorganisms. II.
Studies on planktonic algae. Pp. 88-111 in Lange, R. (ed.): Faggruppenes sluttrapporter
(Final reports from the scientific committees). Forskningsprogram om Havforurensninger
(The Norwegian marine pollution research and monitoring programme (FOH).
Miljøverndepartementet (The Ministry of the Environment), Oslo (in Norwegian).
6C. Sakshaug, E., Hopkins, C.C.E. and Øritsland, N.A. (eds.), 1991: Proceedings of the Pro
Mare Symposium on Polar Marine Ecology. Polar Res. 10, 662 p.
7C. Sakshaug, E., Bjørge, A., Gulliksen, B., Loeng, H. Mehlum, F. (eds) 1992. Økosystem
Barentshavet (Ecosystem Barents Sea). NAVF, NFFR, Ministry of the Environment, 320
p. 2nd edition: Universitetsforlaget, Oslo, 1994.
8C. Sakshaug, E. and J.-A. Sneli (eds) 2000. Trondheimsfjorden. Tapir, Norway, 336 p (in
Norwegian).
Comments re 2C-8C:
2C: Pp. 5-31, 185-188, and 209-232 have been written by myself. This includes the chapters:
"Resources of the sea", "What is oceanography?", "Costs and personnel in Norwegian
oceanographical research, 1963-1972", and "A review of the history of Norwegian oceanographical research up to the second world war"; the latter was written in cooperation with Professor
H. Mosby, University of Bergen. The final version was approved by the Norwegian
Oceanographical Committee
3C: The first 63 pages were written by myself. The final version was approved by The
Norwegian Oceanographical Committee
4C: Introductory parts and parts covering pelagic research were written by myself. The
document was extended with a chapter covering questions related to oil pollution and approved
by the two research councils in question and the Ministry of the Environment in May 1983
7C: This book in Norwegian has been edited and in part written by the editorial committee on
basis of input from participants in Pro Mare. For resons mainly of style the final versions of the
chapters were in the main written by me. This book is the final report from Pro Mare to
Norwegian fisheries and environmental managements as well as stafthe research councils. It also
is used by ocean- and fisheries-relevant organizations and as a textbook for students (universities,
district colleges etc). 2500 copies were printed; 2000 more were printed in spring 1994.
8C: This book has been written/edited on basis of inputs from NTNU researchers and colleagues
at other research institutions in Trondheim. It was presented as a millenium gift from the
Museum to the people of the Trondheimsfjord rregion.
Series D. Posters/Abstracts:
1D. Johnsen, G., Volent, Z., Tangen, K. and E. Sakshaug 1996. Time-series of potential toxic
phytoplankton blooms in North european waters in 1993 using the SEAWATCHEurope buoy system. Eos, suppl. 5, 35.
2D. Sakshaug, E. and Walsh, J. 1999. Barents Sea versus Bering Shelf ecology. Gordon
Research Conference on Polar Marine Carbon Cycle, Ventura, California, March.
3D. Zielinski, O., Barth, H., Breves, W., Chauton, M., Godoy, J., Figueroa, F., Johnsen, G.,
Perez-Rodriguez, E., Reuter, R., Sakshaug, E. and M. Villgarcia. 1999. Underwater
light field in relation to particulate and dissolved matter north of the Canary Islands.
Final CANIGO meeting in Las Palmas. 12-16 September.
4D. Chauton, M., Zielinsky, O., Perez, E., Lorenzo, L. M., Johnsen, G., Egeland, E.,
Sakshaug, E., Reuter, R., Figueroa, F. and F. Figueiras. 1999. Pigment composition
and in vivo fluorescence excitation spectra from the Canary Islands and Azores region.
Final CANIGO meeting in Las Palmas. 12-16 September.
5D. Legrand, C. Johnsen, G., Granéli, E., Sakshaug, E. 2000. Effect of polyamines on growth
and toxicity of Chrysochromulina leadbeateri and Prymnesium patelliferum
(Haptophytae). 9th International conference on Harmful Algal Blooms. Hobart,
Tasmania, 7-11 February 2000.P. 32.
6D. Johnsen, G., Legrand, C.C., Granéli, E, Sakshaug, E. 2000. The use of a DIVING-PAM
for detection of irradiance, biomass and photosystem II fluorescence yields in harmful
algal bloom experiments: Discussion of methodological aspects. 9th International
conference on Harmful Algal Blooms. Hobart, Tasmania, 7-11 February 2000. P 144.
7D. Chauton, M., Johnsen, G., Legrand, C., Granéli, E., Sakshaug, E. 2000. Development of a
potential harmful algal bloom in mesocosm experiment monitored through pigment
analysis and bio-optical measurements. 9th International conference on Harmful Algal
Blooms. Hobart, Tasmania, 7-11 February.
8D. Granéli, E., Legrand, C., Carlsson, P., Maestrini, S. Y., Hummert, C., Reichelt, M.,
Johnsen, G., Pallon, J., Luckas, B. and E. Sakshaug 2000. Nitrogen And phosphorus
deficiency favours the dominance and toxin content of harmful species in summer
Baltic Sea phytoplankton. 9th International conference on Harmful Algal Blooms.
Hobart, Tasmania, 7-11 February 2000.