Chapter 5 Hormonal Responses to Exercise EXERCISE PHYSIOLOGY Theory and Application to Fitness and Performance, 6th edition Scott K. Powers & Edward T. Howley © 2007 McGraw-Hill Higher Education. All rights reserved. Objectives • Describe the hormone-receptor interaction • Identify four factors that influence the contraction of a hormone in the blood • Describe how steroid hormones act on cells • Describe “second messenger” hormone action • Describe the role of hypothalamus-releasing factors in the control of hormone secretion from the anterior and posterior pituitary © 2007 McGraw-Hill Higher Education. All rights reserved. Objectives • Identify the site of release, stimulus for release, and the predominate action of the following hormones: epinephrine, norepinephrine, glucagon, insulin, cortisol, aldosterone, thyroxine, growth hormone, estrogen, and testosterone • Discuss the use of anabolic steroid and growth hormone on muscle growth and their potential side effects © 2007 McGraw-Hill Higher Education. All rights reserved. Objectives • Contrast the role of plasma catecholamines with intracellular factors in the mobilization of muscle glycogen during exercise • Graphically describe the changes in the following hormones during graded and prolonged exercise and discuss how those changes influence the four mechanisms used to maintain the blood glucose concentration: insulin, glucagon, cortisol, growth hormone, epinephrine, and norepinephrine © 2007 McGraw-Hill Higher Education. All rights reserved. Objectives • Describe the effect of changing hormone and substrate levels in the blood on the mobilization of free fatty acids from adipose tissue © 2007 McGraw-Hill Higher Education. All rights reserved. Neuroendocrinology • Endocrine glands release hormones directly into the blood • Hormones alter the activity of tissues that possess receptors to which the hormone can bind • The plasma hormone concentration determines the magnitude of the effect at the tissue level © 2007 McGraw-Hill Higher Education. All rights reserved. Blood Hormone Concentration Determined by: • Rate of secretion of hormone from endocrine gland • Rate of metabolism or excretion of hormone • Quantity of transport protein • Changes in plasma volume © 2007 McGraw-Hill Higher Education. All rights reserved. Control of Hormone Secretion • Rate of insulin secretion from the pancreas is dependent on: – Magnitude of input – Stimulatory vs. inhibitory © 2007 McGraw-Hill Higher Education. All rights reserved. Factors That Influence the Secretion of Hormones Fig 5.1 © 2007 McGraw-Hill Higher Education. All rights reserved. Hormone-Receptor Interactions • Trigger events at the cell • Magnitude of effect dependent on: – Concentration of the hormone – Number of receptors on the cell – Affinity of the receptor for the hormone © 2007 McGraw-Hill Higher Education. All rights reserved. Hormone-Receptor Interactions • Hormones bring about effects by: – Altering membrane transport – Stimulating DNA to increase protein synthesis – Activating second messengers • Cyclic AMP • Ca++ • Inositol triphosphate • Diacylglycerol © 2007 McGraw-Hill Higher Education. All rights reserved. Mechanism of Steroid Hormones Fig 5.2 © 2007 McGraw-Hill Higher Education. All rights reserved. Cyclic AMP “Second Messenger” Mechanism Fig 5.3 © 2007 McGraw-Hill Higher Education. All rights reserved. Other “Second Messenger” Systems Fig 5.4 © 2007 McGraw-Hill Higher Education. All rights reserved. Hormones: Regulation and Action • Hormones are secreted from endocrine glands – Hypothalamus and pituitary glands – Thyroid and parathyroid glands – Adrenal glands – Pancreas – Testes and ovaries © 2007 McGraw-Hill Higher Education. All rights reserved. Hypothalamus • Controls activity of the anterior and posterior pituitary glands • Influenced by positive and negative input © 2007 McGraw-Hill Higher Education. All rights reserved. Positive and Negative Input to the Hypothalamus Fig 5.6 © 2007 McGraw-Hill Higher Education. All rights reserved. Anterior Pituitary Gland Fig 5.5 © 2007 McGraw-Hill Higher Education. All rights reserved. Growth Hormone • Secreted from the anterior pituitary gland • Essential for normal growth – Stimulates protein synthesis and long bone growth • Increases during exercise – Mobilizes fatty acids from adipose tissue – Aids in the maintenance of blood glucose © 2007 McGraw-Hill Higher Education. All rights reserved. Growth Hormone Fig 5.6 © 2007 McGraw-Hill Higher Education. All rights reserved. Posterior Pituitary Gland • Secretes antidiuretic hormone (ADH) or vasopressin • Reduces water loss from the body to maintain plasma volume • Stimulated by: – High plasma osmolality and low plasma volume due to sweating – Exercise © 2007 McGraw-Hill Higher Education. All rights reserved. Change in the Plasma ADH Concentration During Exercise Fig 5.7 © 2007 McGraw-Hill Higher Education. All rights reserved. Thyroid Gland • Triiodothyronine (T3) and thyroxine (T4) – Important in maintaining metabolic rate and allowing full effect of other hormones • Calcitonin – Regulation of plasma Ca++ • Parathyroid Hormone – Also involved in plasma Ca++ regulation © 2007 McGraw-Hill Higher Education. All rights reserved. Adrenal Medulla • Secretes Epinephrine and Norepinephrine • Increases –HR, glycogenolysis, lypolysis, © 2007 McGraw-Hill Higher Education. All rights reserved. Adrenal Cortex • Mineralcorticoids (aldosterone) – Maintain plasma Na+ and K+ – Regulation of blood pressure © 2007 McGraw-Hill Higher Education. All rights reserved. Change in Mineralcorticoids During Exercise Fig 5.8 © 2007 McGraw-Hill Higher Education. All rights reserved. Adrenal Cortex • Glucocorticoids (Cortisol) – Stimulated by exercise and long-term fasting – Promotes the use of free fatty acids as fuel – Stimulates glucose synthesis – Promotes protein breakdown for gluconeogenesis and tissue repair © 2007 McGraw-Hill Higher Education. All rights reserved. Control of Cortisol Secretion Fig 5.9 © 2007 McGraw-Hill Higher Education. All rights reserved. Pancreas • Secretes digestive enzymes and bicarbonate into small intestine • Releases – Insulin - Promotes the storage of glucose, amino acids, and fats – Glucagon - Promotes the mobilization of fatty acids and glucose – Somatostatin - Controls rate of entry of nutrients into the circulation © 2007 McGraw-Hill Higher Education. All rights reserved. Testes • Release testosterone – Anabolic steroid • Promotes tissue (muscle) building • Performance enhancement – Androgenic steroid • Promotes masculine characteristics © 2007 McGraw-Hill Higher Education. All rights reserved. Control of Testosterone Secretion Fig 5.10 © 2007 McGraw-Hill Higher Education. All rights reserved. Estrogen • Establish and maintain reproductive function • Levels vary throughout the menstrual cycle © 2007 McGraw-Hill Higher Education. All rights reserved. Control of Estrogen Secretion Fig 5.11 © 2007 McGraw-Hill Higher Education. All rights reserved. Muscle Glycogen Utilization • Breakdown of muscle glycogen is under dual control – Epinephrine-cyclic AMP – Ca2+-calmodulin • Delivery of glucose parallels activation of muscle contraction • Glycogenolysis – breakdown of glycogen Fig 5.16 © 2007 McGraw-Hill Higher Education. All rights reserved. Control of Glycogenolysis Glycogenolysis Fig 5.16 © 2007 McGraw-Hill Higher Education. All rights reserved. Muscle Glycogen Utilization • Glycogenolysis is related to exercise intensity – High-intensity of exercise results in greater and more rapid glycogen depletion Fig 5.13 • Plasma epinephrine is a powerful simulator of glycogenolysis – High-intensity of exercise results in greater increases in plasma epinephrine Fig 5.14 © 2007 McGraw-Hill Higher Education. All rights reserved. Glycogen Depletion During Exercise Fig 5.13 © 2007 McGraw-Hill Higher Education. All rights reserved. Plasma Epinephrine Concentration During Exercise Fig 5.14 © 2007 McGraw-Hill Higher Education. All rights reserved. Maintenance of Plasma Glucose During Exercise • Mobilization of glucose from liver glycogen stores • Mobilization of FFA from adipose tissue – Spares blood glucose • Gluconeogenesis from amino acids, lactic acid, and glycerol • Blocking the entry of glucose into cells – Forces use of FFA as a fuel © 2007 McGraw-Hill Higher Education. All rights reserved. Blood Glucose Homeostasis During Exercise • Permissive and slow-acting hormones – Thyroxine – Cortisol – Growth hormone • Act in a permissive manner to support actions of other hormones © 2007 McGraw-Hill Higher Education. All rights reserved. Cortisol • Stimulates FFA mobilization from adipose tissue • Mobilizes amino acids for gluconeogenesis • Blocks entry of glucose into cells Fig 5.17 © 2007 McGraw-Hill Higher Education. All rights reserved. Role of Cortisol in the Maintenance of Blood Glucose Fig 5.17 © 2007 McGraw-Hill Higher Education. All rights reserved. Plasma Cortisol During Exercise • At low intensity – plasma cortisol decreases • At high intensity – plasma cortisol increases Fig 5.18 © 2007 McGraw-Hill Higher Education. All rights reserved. Changes in Plasma Cortisol During Exercise Fig 5.18 © 2007 McGraw-Hill Higher Education. All rights reserved. Growth Hormone • Important in the maintenance of plasma glucose – Decreases glucose uptake – Increases FFA mobilization – Enhances gluconeogenesis Fig 5.19 © 2007 McGraw-Hill Higher Education. All rights reserved. Growth Hormone in the Maintenance of Plasma Glucose Fig 5.19 © 2007 McGraw-Hill Higher Education. All rights reserved. Growth Hormone During Exercise: Effect of Intensity Fig 5.20 © 2007 McGraw-Hill Higher Education. All rights reserved. Growth Hormone During Exercise: Trained vs. Untrained Fig 5.20 © 2007 McGraw-Hill Higher Education. All rights reserved. Blood Glucose Homeostasis During Exercise • Fast-acting hormones – Norepinephrine and epinephrine – Insulin and glucagon • Maintain plasma glucose – Increasing liver glucose mobilization – Increased levels of plasma FFA – Decreasing glucose uptake – Increasing gluconeogenesis Fig 5.21 © 2007 McGraw-Hill Higher Education. All rights reserved. Role of Catecholamines in Substrate Mobilization Fig 5.21 © 2007 McGraw-Hill Higher Education. All rights reserved. Epinephrine & Norepinephrine During Exercise • Increase linearly during exercise • Favor the mobilization of FFA and maintenance of plasma glucose © 2007 McGraw-Hill Higher Education. All rights reserved. Change in Plasma Catecholamines During Exercise Fig 5.22 © 2007 McGraw-Hill Higher Education. All rights reserved. Epinephrine & Norepinephrine Following Training • Decreased plasma levels in response to exercise bout • Parallels reduction in glucose mobilization © 2007 McGraw-Hill Higher Education. All rights reserved. Plasma Catecholamines During Exercise Following Training Fig 5.23 © 2007 McGraw-Hill Higher Education. All rights reserved. Effects of Insulin & Glucagon Fig 5.24 © 2007 McGraw-Hill Higher Education. All rights reserved. Insulin During Exercise • Plasma insulin decreases during exercise – Prevents rapid uptake of plasma glucose – Favors mobilization of liver glucose and lipid FFA • Trained subjects during exercise – More rapid decrease in plasma insulin – Increase in plasma glucagon © 2007 McGraw-Hill Higher Education. All rights reserved. Changes in Plasma Insulin During Exercise Fig 5.25 © 2007 McGraw-Hill Higher Education. All rights reserved. Effect of Training on Plasma Insulin During Exercise Fig 5.25 © 2007 McGraw-Hill Higher Education. All rights reserved. Effect of Training on Plasma Glucagon During Exercise Fig 5.26 © 2007 McGraw-Hill Higher Education. All rights reserved. Effect of SNS on Substrate Mobilization Fig 5.28 © 2007 McGraw-Hill Higher Education. All rights reserved. Hormonal Responses to Exercise Fig 5.29a © 2007 McGraw-Hill Higher Education. All rights reserved. Hormonal Responses to Exercise Fig 5.29b © 2007 McGraw-Hill Higher Education. All rights reserved. Free Fatty Acid Mobilization During Heavy Exercise • FFA mobilization decreases during heavy exercise – This occurs in spite of persisting hormonal stimulation for FFA mobilization • May be due to high levels of lactic acid – Promotes resynthesis of triglycerides – Inadequate blood flow to adipose tissue – Insufficient transporter for FFA in plasma © 2007 McGraw-Hill Higher Education. All rights reserved. Effect of Lactic Acid on FFA Mobilization Fig 5.30 © 2007 McGraw-Hill Higher Education. All rights reserved. Chapter 5 Hormonal Responses to Exercise © 2007 McGraw-Hill Higher Education. All rights reserved.