Ichthyology “jawed fishes” • Class Chondrichthyes- cartilaginous • Subclass Elasmobranchii- sharks, rays, and skates • Subclass Holocephali- chimeras & ratfishes • Class Osteichthyes- bony fishes • Subclass Dipnoi- lung fishes • Subclass Crossopterygii- coelocanth • Subclass Actinopterygii- modern bony fishes Class Chondrichthyes: (Cartilaginous Fishes) • • • • • • general diagnostic characters: reduced dermal armor cartilaginous endoskeleton & girdles tricuspid teeth gill slits fins: two dorsal, no anal, stiff pectorals, flexible pelvic • heterocercal tail Sharks Sting Ray Manta Ray Chimera or Ratfish 1st Elasmobranch Radiation diagnostic characters: 1. shark-like appearance 2. terminal mouth; amphistylic jaw suspension 3. unique tooth replacement 4. claspers Claspers nd 2 Elasmobranch Radiation diagnostic characters: 1. heterodont dentition 2. unique pectoral & pelvic fin arrangements 3. heterocercal tail Extant Forms • Elasmobranchii [sharks (360 species) , skates & rays (456 species); appear in the Jurassic diagnostic characters of sharks 1. hyostylic jaw suspension & rostrum 2. cranial kinesis- allows consumption of large food items 3. pectoral girdle fused ventrally 4. placoid scales 5. lack gas bladder; liver provides buoyancy (~ 25% of body mass). Hyostylic Jaw Suspension sensory systems 1. lateral line system (mechanoreceptors) 2. ampullae of Lorenzini (detects electrical impulses) 3. acute chemosensory 4. tapetum lucidum 5. internal fertilization and large female investment; oviparity, ovoviviparity, & viviparity Ampullae of Lorenzini Placoid Scales Placoid Scales Shark Egg Case diagnostic characters of skates & rays 1. hard teeth with flat crowned plates; mostly adapted for feeding on benthic invertebrates 2. swimming via undulations of pectoral fins 3. stingrays (family Dasyatidae) modified placoid scales form a sting at the base of the tail 4. manta rays (family Mobulidae) plankton feeders Cartilaginous Skeleton Shark Anatomy Link Click Here Class Osteichthyes: “bony fishes” • most abundant and diverse vertebrates >21,000 species • bony scales (enlarged in the head region) • operculum covering the gills • stream-lined; laterally compressed • ossified endoskeleton; secondarily lost in some species Operculum Dipnoans 1. subclass Dipnoi- "double breathers"; lungfishes 2. upper jaw fused to the brain case; large, crushing tooth plates 3. two dorsal fins 4. paired fins Crossopterygians • dominant fresh water predators during the Devonian period • rhipidistians (ancestor to amphibians) & coelacanth; Latimeria (extant) important structural characteristics: • lobed fins with elements connecting to pectoral & pelvic girdles • autostylic jaw suspension- jointed braincase (jaw connects directly to the brain case) • labyrinth tooth- complex folding of the enamel Coelocanth (Latimeria) Actinopterygians (ray-finned fishes) • most modern bony fishes; subclass Actinopterygii • important structural characteristics: • ganoid scales • distinctive pectoral & pelvic girdles • fins attach to the body via flexible fin rays instead of a fleshy lobe • branchiostegal rays • no internal nares Branchiostegal Rays Ganoid Scales Ctenoid Scales Cycloid Scales “bony fish” Paddlefish (Polydon) Sturgeon Fish Diversity Smallmouth Bass Green Sunfish Catfish Oklahoma Non-game Fish