Animals are multicellular and heterotrophic. Animal kingdom is most diverse than the plant kingdom. Based on the organization of the cells, there are 3 levels of organization Cellular level of organization: Cells are arranged as loose cell aggregates. E.g. sponges Tissue level of organization: Cells are arranged into tissues. E.g. coelenterates, ctenophores Organ level of organization: Tissues are arranged into organs and the organs are associated to form organ systems. Each system performs a specific physiological function. E.g. All higher animals (from platyhelminthes to chordates). Organ systems of different animals show complexities. For example, The digestive system of cnidarians and platyhelminthes is incomplete (only a single opening functioning as mouth and anus). Complete digestive system has 2 openings (mouth and anus). Circulatory system is 2 types (open and closed). It is the arrangement of similar body parts on 2 sides of main axis of body. Based on symmetry, animals are 2 types: Symmetry is 2 types: Radial symmetry Bilateral symmetry Radial symmetry: Body can be divided into 2 similar parts by any plane along oral- aboral axis of body. E.g. some Sponges, Coelenterates, Echinoderms. Bilateral symmetry: Body can be divided into 2 right and left halves by a section passing through the longitudinal axis. Such animal has An upper or vertebral dorsal side A lower ventral side Left and right lateral sides Anterior (cephalic) side and Posterior (anal) side. E.g. All vertebrates and many invertebrates. The layers of gastrula from which body organs are formed. Based on the number of germ layers, animals are 2 types- Diploblastic and Triploblastic. Diploblastic animals: 2 germ layers- outer ectoderm and inner endoderm. E.g. Sponges and Coelenterates. Triploblastic animals: 3 germ layers- Outer ectoderm, middle mesoderm and inner endoderm. E.g. Flat worms to mammals. It is the space between body wall and gut wall. Coelom separates the muscles of gut and body wall. On the basis of nature of coelom animals are 3 types. Acoelomate Pseudocoelomate Eucoelomate (True coelomate) Acoelomate: No coelom. The space between body wall and digestive cavity is filled with matrix (parenchyma). E.g. Porifers, Coelenterates and flat worms. Pseudocoelomates: Here, blastocoel becomes the coelom. It is not lined by peritoneal layer. E.g. roundworms. Eucoelomates (True coelomates): The coelom in animals except the roundworms arises from the mesoderm. Coelom is lined by peritoneal layer and filled with coelomic fluid. E.g. Annelids to mammals. Functions of coelom: It accommodates visceral organs Coelomic fluid provides moist environment to visceral organs Acts as shock absorber Friction is minimized. In some animals, the body is formed of similar parts (segments or metameres). This segmentation is called metamerism. E.g. Annelids, Arthropods. In vertebrates: only internal metamerism. It is a supporting rod formed on the dorsal side during embryonic development in some animals. Animals with notochord are called chordates and those without notochord are called non-chordates. Levels of organisation Germ layers Symmetry Coelom Phylum Cellular level Diploblastic Asymmetrical / radial Acoelomate Porifera Tissue Diploblastic Radial Acoelomate Cnidaria Ctenophora Acoelomate Platyhelminthes Pseudocoelomate Aschelminthes Coelomates Annelida Arthropoda Mollusca Echinodermata Hemichordata Chordata Organ/ Organ system Triploblastic Bilateral Habit: Aquatic. Sedentary. Solitary or colonial. Digestive system: Absent. Digestion is intracellular. Water transport (canal system) to gather food. Respiratory system: Absent. Respiration by canal system. Circulatory system: Absent. Circulation by canal system. Excretory system: Absent. Excretion by canal system. Nervous system: Absent Reproductive system: Hermaphrodite. ARP by fragmentation. SRP by the formation of sperms and ova. Fertilization internal. Development is indirect. Other features: Millions of minute pores (ostia). Body wall with outer layer of flat cells (pinacocytes) and inner layer of flagellated cells (choanocytes or collar cells). Body is supported by spicules & spongin fibres. Highest degree of regenerating capacity. Habit: Aquatic. Solitary or colonial. Sessile or free swimming. Digestive system: Incomplete. Intracellular and extracellular digestion. Respiratory system: Absent Circulatory system: Absent Excretory system: Body surface Nervous system: Only a network of neurons. Reproductive system: Polyp reproduces asexually (budding) and medusa sexually. Fertilization external. Development is indirect. Other features: Tentacles for food collection and defence. Tentacles possess cnidoblasts (stinging cells) with poison filled capsule (nematocyst). Cnidoblast is used to kill or paralyse the enemies & prey. CNIDOBLAST Polyp and Medusa are 2 kinds of body forms. Show polymorphism and alternation of generation. Habit: Marine, solitary and pelagic. Digestive system: Incomplete. Intracellular & extracellular digestion. Respiratory system: Absent Circulatory system: Absent Excretory system: Body surface Nervous system: Only a network of neurons. Reproductive system: Only SRP. Hermaphrodite. Fertilization external. Development is indirect. Other features: Locomotion is by 8 vertical external rows of ciliated comb plates. Tentacles present. Shows Bioluminescence. Habit: Parasites. Some are free-living. Mainly aquatic. Digestive system: Branched alimentary canal with mouth. Anus absent. In parasites alimentary canal absent. Respiratory system: Absent Circulatory system: Absent Excretory system: Protonephridia with flame cells (solenocytes). Ammonotelic Nervous system: Ladder-like. A brain and nerve cords, connected by transverse nerves. Reproductive system: ARP by fragmentation and regeneration. Most are hermaphrodite. In SRP fertilization is internal. Development is indirect Other features: Unsegmented flat leaf like body (except tape worms). The first group showing cephalization. Examples: Taenia solium, Planaria, Liver fluke Habit: Free living or parasitic Digestive system: Tubular alimentary canal with mouth, pharynx, intestine and anus. Mouth may possess teeth Respiratory system: Absent Circulatory system: Absent Excretory system: H-shaped Renett cells. Ammonotelic Nervous system: Consists of circumpharyngeal ring with dorsal and ventral nerves. Reproductive system: SRP. Internal fertilization. Development is direct. Other features: Syncitial epidermis. Thick cuticle. Sexual dimorphism. Examples: Ascaris, Ancylostoma, Wuchereria, Enterobius Habit: Terrestrial, fresh water or marine or parasitic. Digestive system: Pharynx, oesophagus, stomach, intestine. Respiratory system: Cutaneus respiration. Some have branchial (gill) respiration Circulatory system: Closed type Excretory system: Nephridia. Aq. forms are ammonotelic and terr. forms ureotelic. Nervous system: A pair of cerebral ganglia (brain) and a double ventral nerve cord. Reproductive system: Earth worms and leeches are hermaphrodites. Free swimming trochophore larva. Other features: True segmentation. Covered by thick cuticle. Locomotory organs are setae or parapodia. Examples: Earthworm, Leech, Neries, Aphrodite, Chaetopterus Habit: Cosmopolitan Digestive system: Well developed. Mouth parts are different types. Respiratory system: Gills, trachea, book gills or book lungs. Circulatory system: Open type Excretory system: Either Antennary glands or Malpighian tubules or coxal glands. Aq. forms are ammonotelic. Ter. forms are uricotelic. Nervous system: A nerve ring followed by a double ventral nerve cord, which is ganglionated. Reproductive system: Sexes are separate. External fertilization in aq.forms. Internal fertilization in land forms. Oviparous. Some are viviparous. Other features: Jointed appendages. 3 regions: head, thorax and abdomen. Body is covered by cuticle. Ecdysis (moulting) present. Metamorphosis present. Examples: Spider, Scorpion, Crab, Prawn, Lepisma, insects etc. Habit: Generally aquatic. Few are terrestrial. Digestive system: Well developed. Salivary glands and liver present. Respiratory system: Gills in aq.forms and pulmonary sac in terrestrial forms. Circulatory system: Open type Excretory system: Organs of Bojanus. Ammonotelic or ureotelic. Nervous system: Paired ganglia and their connectives. Reproductive system: Sexes are separate. Development direct or indirect. Larval forms are glochidium, trochophore and veliger. Other features: Body is divisible into head, visceral mass (visceral hump) & foot. Univalve or bivalve shell. Mantle and radula present. Examples: Pila, Achatina, Pinctada, Sepia, Loligo, Octopus, Habit: Exclusively marine. Digestive system: Simple and complete. Respiratory system: Dermal branchiae (gills) or papulae and tube feet. Circulatory system: Reduced Excretory system: Diffusion through gills. Ammonotelic Nervous system: Circum oral nerve ring and some radiating nerves. No brain. Reproductive system: Fertilization external. Development indirect. Ciliated free swimming larva. , Other Features: Body is covered with spines for protection. Head absent. Endoskeleton (ossicles) present. Water vascular system for locomotion and food capture. Great power of autotomy and regeneration. Examples: Star fish, Echinus, Echinocardium, Antedon, Cucumaria Habit: Marine Digestive system: Complete Respiratory system: Gills Circulatory system: Open type Excretory system: Proboscis gland Nervous system: Reproductive system: infra-epidermal type Sexes are separate. Fertilization external. Development is indirect. Other features: Cylindrical body composed of an anterior proboscis, a collar and a long trunk. Examples: Balanoglossus, Saccoglossus These are animals with notochord, dorsal tubular nerve cord and pharyngeal gill slits. Notochord is a flexible rod located in the mid dorsal line between the alimentary canal and the nerve cord in the embryo. Chordata Non-chordata Notochord is found in the embryonic stage Absent Central nervous system is dorsal, hollow and single Pharyngeal gill slits present Ventral heart A post-anal tail is present Endoskeleton and exoskeleton present Hepatic portal system present Ventral, solid and double Absent Dorsal heart (if present) Absent Only exoskeleton Absent Marine Heterocercal caudal fin Cartilaginous Gill slits without operculum Skin with placoid scales Teeth are modified placoid scales which are backwardly directed No air bladder so they have to swim constantly to avoid sinking Sexes are separate. In males pelvic fins bear claspers Internal fertilization Many of them viviparous E.g. Scoliodon (Dog fish), Pristis (Saw fish), Carcharodon (Great white shark), Trygon (Sting ray), Torpedo (have electric organ). Marine and fresh water Homocercal caudal fin Bony endoskeleton Terminal mouth Gills (4 pairs) covered by an operculum on each side Cycloid, ctenoid or ganoid scales Air bladder for buoyancy Sexes are separate Fertilisation external Mostly oviparous Development direct Marine Fresh water Aquarium Exocoetus (flying fish) Labeo (Rohu) Betta (Fighting Hippocampus (sea horse) Catla (Katla) fish) Sardine Clarias (Magur) Pterophyllum Mackeral Tilapia (Angel fish) etc. Tuna Anabas (Climbing Remora (sucker fish) perch) Pomfret Cybium Aquatic larval life and terrestrial adult life For breeding they require a watery environment Pentadactyl limbs Moist skin without scales A tympanum represents the ear Alimentary canal, urinary and reproductive tracts open into a common chamber (Cloaca) which opens to exterior Respiration is by gills (in larva), lungs and through skin. 3-chambered heart (2 auricles + 1 ventricle) Poikilotherms Sexes are separate. Fertilisation external. Oviparous. Development is indirect E.g. Bufo (Toad), Rana (Frog), Hyla (Tree frog), Salamandra (Salamander), Ichthyophis (Limbless amphibia), Ambystoma (Tiger salamander), Rhacophorus (Flying frog) Dry & cornified skin, epidermal scales or scutes Snakes and lizards shed their scales as skin cast. Tympanum represents ear. Limbs- 2 pairs (if present) 3-chambered heart (but a septum partially separates ventricle). Heart is 4-chambered in crocodiles. Poikilotherms Fertilisation internal Oviparous Development is direct Examples: Chelone (Turtle), Testudo (Tortoise) Chameleon (Tree lizard), Calotes (Garden lizard) Crocodilus (Crocodile), Alligator, Gavialis (gharial) Hemidactylus (Wall lizard), Sphenodon (Tautara), Draco (flying dragon or flying lizard), Phrynosoma (horned lizard), Varanus (monitor), Poisonous snakes: Naja (Cobra), Bangarus (Krait), Vipera (Viper), Crotalus (rattle snake), Enhydrina (sea snake) etc. Non-poisonous snakes: Python, Typhlos (blind snake), Dryophis (tree snake) Presence of feathers Possess beak Forelimbs are modified into wings Hind limbs have scales and are modified for walking, swimming, or clasping tree branches Skin is dry without glands except the oil gland at the base of tail Long bones are hollow and pneumatic Digestive tract with additional chambers, the crop and gizzard Heart is 4-chambered Homoiotherms (warm-blooded). Double respiration. Air sacs connected to lungs. No urinary bladder. Fertilisation is internal. Oviparous. Development is direct. E.g. Corvus (Crow), Columba (Pigeon), Psittacula (Parrot), Struthio (Ostrich), Pavo (Peacock), Gullus (Fowl), Bubo, Aptenodytes (Penguin), Neophron (Vulture) etc. Presence of mammary glands (milk producing glands). 2 pairs of limbs for walking, running, climbing, burrowing, swimming or flying. Skin with hair. External ear (Pinnae) Heterodont, thecodont, diphyodont. 4-chambered heart. Homoiotherms. Respiration by lungs. Sexes are separate. Fertilisation internal. Viviparous (except Echidna and Platypus). Development is direct. E.g. Ornithorhynchus (Platypus), Macropus (Kangaroo), Pteropus (flying fox), Camelus (Camel), Macaca (Monkey), Rattus (Rat), Canis (dog), Felis (Cat), Elephas (Elephant), Equus (Horse), Delphinus (Common Dolphin), Balaenoptera (blue whale), Panthera tigris (Tiger), Panthera leo (lion) Protochordata (Acraniata) Urochordata (Tunicata) Vertebrata (Craniata) Cephalochordata Notochord present Notochord extends from Possess notochord during the only in larval tail head to tail region and is embryonic period Body is covered by persistent throughout Notochord is replaced by a test made up of the life cartilaginous or bony tunicin Exclusively marine vertebral column in the adult Exclusively marine Fish-like Ventral muscular heart Retrogressive No definite coelom Kidneys for excretion and metamorphosis Sexes are separate osmoregulation Hermaphrodite E.g. Branchiostoma Paired appendages which E.g. Ascidia, Salpa, (Amphioxus or Lancelet) may be fins or limbs Doliolum. Ascidia