THE GENUS HOMO Early homo 2.5-1.6 mya Louis Leakey (1960) Olduvai Gorge, Northern Tanzania 1.8 mya Homo habilis Significance of the name Early tool makers Ancestors to modern humans Other Plio-Pleistocene sites EAST AFRICA – Omo, Southern Ethiopia Koobi Fora, KMN-ER 1470 – Richard Leakey Olduvai Gorge SOUTH AFRICA Sterkfontein Swartkrans General trends of early homo in comparison to Australopithecines CRANIAL ANATOMY Brain expansion 600-800 cc Thin walled – no crests Rounded vault Weakly projecting ridges Variable facial proportions between individuals No sagittal crest Reduced prognathism Rounded cranial vault KNM ER 1470 (homo rudolfensis?) Koobi Fora, Richard Leakey (1972) 1.9 mya 752 cc Rounded cranial vault No heavy crests Robust face Back teeth still quite large (but < australopithecines) Associated crude stone tools More human like brain organization Evolutionary Role played by early homo TRENDS: Increased encephalization Dental reduction Standardized tool making ? Opportunistic tools - Manuports 5 – 2.5 mya Unaltered objects carried some distance before use LOWER PALEOLITHIC (old stone age) Approx. 2.5 mya Important in brain evolution Emphasizes manual dexterity Requires a complex, abstract thought process OLDOWAN INDUSTRIAL COMPLEX Earliest identifiable stone tools 2.5 mya – 1.4 mya Sites: lake Turkana Olduvai Gorge Hadar Omo Characterized by Oldowan Chopping Tool All purpose chopping tool Made from smooth water worn cobbles Multi-purpose Butcher meat, split bones, defense Hard hammer/PERCUSSION METHOD Hammerstone – Bipolar Percussion - Rock used to strike flakes off cobble Anvil – Direct Percussion - Large rock that you strike the cobble against or place the cobble on Concoidal fracture Produced by striking one stone against another Flake - Chip of rock off the cobble - Sharp, useful edges - scrapers Cultural Significance of Oldowan tool tradition New additions to diet: Meat group sharing scavenging Saved time & labor for food acquisition Probably resulted from an adaptation to a new environment Bipedalism freed hands Large predatory carnivores Significance of Oldowan tool tradition Evolution of brain Manual dexterity Fine motor manipulation Set stage for language acquisition Brain lateralization handedness Homo erectus 1st hominin species to exist outside of Africa an African origin an impressive longevity Extensive geographical spread t/o Old World HOMO ERECTUS First Find: Eugene Dubois, 1891 Trinil, Java Femur Pithecanthropus erectus JAVA SITES (1.8 mya – 500,000 ya) ZHOUKOUDIAN, China Gunnar Andersson Pei Wenshong Davidson Black Franz Weinreich Best collection of 600,000 – 300,000 yr old hominids ever found Minimum 40 individuals – adult male, female & subadults 1000cc aver > 100,000 artifacts East African H. erectus (h.ergaster) Nariokotome, West Turkana 1984 Kamoya Kimeu WT 15000 (Nariokotome boy) 1.6 mya Boy approx. 12 y.o. Stature 5’3” Postcranial similar to modern humans 880 cc Issues at Turkana – h. ergaster or h. erectus? Large amount of variation Too large for one species? Groves & Mazak (1975) 2. Temporal relationships inconsistencies in the chronological transition of morphological changes from h. habilis to h. ergaster to h. erectus African specimens Bones thinner Smaller brow ridges Asian specimens Shorter & stockier ridges Characteristics of homo erectus Cranial anatomy 780 – 1225 cc (1000 cc average) Wide at base from behind Low cranial vault Sagittal keel Nuchal torus Supraorbital torus Robust jaw Teeth larger than modern human, but smaller than h. habilis Homo erectus postcranial anatomy Postcranial skeleton similar to modern humans Robust limb bones Conical rib cage (different from humans who have barrel shaped rib cage) Narrow hips Stature: human range < sex dimorphism than h. habilis H. erectus in Europe or Northern Africa? Dmanisi, Georgia North of the Levantine Corridor Approx 1.8 mya 5 individuals Average 650 cc Homo georgicus or erectus? (Gabounia et. al., 2002) Skull vault elongated, narrow, low From behind skull widest at base Saggital keel Postorbital constriction Supraorbital torus Differences from Homo erectus small face More prognathic Reminiscent of early homo Homo naledi Lee Berger et al., 2015 Dinaledi chamber, South Africa At least 15 individuals Cranial: 513 cc Sagittal keel, supraorbital torus Postcranial: small human Human hands & feet Australo: trunk, shoulder, pelvis Currently undated Homo erectus culture Transition from Oldowan to Acheulean @ Olduvai 3 stage transition (Mary Leakey, 1975) Increased sophistication in Oldowan protobiface Coexistence of Oldowan & Acheulean Disappearance of Oldowan & further development of Acheulean Diversified function Symbolism? Punctuated transition (not gradual) (Issac 1969) ACHEULEAN TOOL TRADITION 1ST appeared 1.7 mya in East Africa Spreads to Middle East & Europe 750,000 ya HANDAXE (biface) pear shaped/pointed First fully conceived implements Significant step in conceptualization More standardized form TOOL DIVERSIFICATION CLEAVERS Handaxes with a straight cutting edge Butcher meat, chopping, breaking bones FLAKE TOOLS Strike a flint core with hammerstone Retouched to make A. side scraper B. point C. end scraper D. burin LATE ACHEULEAN - NEW TECHNIQUE SOFT HAMMER PERCUSSION (baton) Involved using a bone or antler punch to hit the edge of the flint core producing shallow flake scars Longevity of Acheulean tool tradition originated in Africa - approx 1.7 mya Disappeared about 250,000 ya Middle East (Ubeidiya, Israel) - 1 mya Europe - 500,000 – 600,000 ya NE Pakistan - 400,000 – 730,000 ya The CHOPPING TOOL TRADITION developed by H. erectus in ASIA 460,000 – 230,000 ya More reminiscent of Oldowan Probably grew out of Oldowan Large choppers with jagged edges Zhoukoudian NO acheulean sites in Far east (China) and SE Asia – WHY? Different resources available for tool making (i.e. bamboo) (Pope, 1993) Movius line: divides the acheulean and non acheulean regions (Hallam Movius, 1948) o Bose site, Southern China (Yamei et al., 2000) - bifaces - Challenges Movius hypothesis Different functional requirements (Schick & Toth, 1993) The new dates from Java (1.8 mya) place homo erectus in SE Asia @ least 100,000 years BEFORE the advent of the Acheulean in Africa Regional variation within China (Shen & Qi, 2004) Lack of suitable raw stone Other Accomplishments of homo erectus FIRE? Evidence: 1.6-1.5 mya Koobi Fora & Chesowanja, Kenya? Swartkrans, S Africa? Hearths? Zhoukoudian 670,000 – 410,000 Gesher Benot Ya’aqov, Israel 790,000 ya Homo erectus diet Scavengers Zhoukoudian Boaz & Ciochon (2004) Binford & Stone (1986) Increased meat consumption: Higher energy foods (over plants) Easily digestible Need less food and smaller digestive organs Requires less digestive effort than earlier hominins Provided plenty of energy for their expanding brains Communication speech and language? Protolanguage with rudimentary words but no grammatical rules Terrence Deacon (1997) communicated symbolically short phrases