Coral Ecology Coral reefs • Coral reefs are hard, wave-resistant structures composed of individual coral animals (polyps) • Individual coral polyps: • • • • • • Are about the size of an ant Are related to jellyfish Feed with stinging tentacles Live attached to the sea floor in large colonies Construct hard calcium carbonate structures for protection Contain symbiotic photosynthetic zooxanthellae algae Coral reef distribution and diversity Coral reef zonation Coral Ecology • Ecology = the study of the interaction of organisms with their environments. • It involves understanding biotic and abiotic factors influencing the distribution and abundance of living things. • The word "ecology" coined from Greek word "oikos", which means "house" or "place to live”. The Scope of Ecology • population growth • competition between species • symbiotic relationships • trophic (=feeding) relationships • origin of biological diversity • interaction with the physical environment Population Growth Idealized models predict patterns of population growth ▪ The rate of population increase under ideal conditions is called exponential growth. It can be calculated using the exponential growth model equation, G = rN, in which • G is the growth rate of the population, • N is the population size, and • r is the per capita rate of increase (the average contribution of each individual to population growth). ▪ Eventually, one or more limiting factors will restrict population growth. Population size (N) 500 450 400 350 300 250 200 150 100 50 0 0 1 2 3 4 5 6 7 8 9 10 11 12 Time (months) Idealized models predict patterns of population growth • The logistic growth model is a description of idealized population growth that is slowed by limiting factors as the population size increases. • To model logistic growth, the formula for exponential growth, rN, is multiplied by an expression that describes the effect of limiting factors on an increasing population size. • K stands for carrying capacity, the maximum population size a particular environment can sustain. (K − N) G = rN K Breeding male fur seals (thousands) 10 8 6 4 2 0 1915 1925 1935 Year 1945 Multiple factors may limit population growth • The logistic growth model predicts that population growth will slow and eventually stop as population density increases. • At increasing population densities, densitydependent rates result in • declining births and • increases in deaths. Multiple factors may limit population growth • Intraspecific competition is • competition between individuals of the same species for limited resources and • is a density-dependent factor that limits growth in natural populations. Multiple factors may limit population growth • Limiting factors may include • • • • food, nutrients, retreats for safety, or nesting sites. 100 Survivors (%) 80 60 40 20 0 20 40 60 80 100 120 Density (beetles/0.5 g flour) Multiple factors may limit population growth • In many natural populations, abiotic factors such as weather may affect population size well before density-dependent factors become important. • Density-independent factors are unrelated to population density. These may include • • • • fires, storms, habitat destruction by human activity, or seasonal changes in weather (for example, in aphids). Number of aphids Exponential growth Apr May Jun Sudden decline Jul Aug Sep Oct Nov Dec Month EVOLUTION CONNECTION: Evolution shapes life histories • The traits that affect an organism’s schedule of reproduction and death make up its life history. • Key life history traits include • • • • age of first reproduction, frequency of reproduction, number of offspring, and amount of parental care. Evolution shapes life histories • Populations with so-called r-selected life history traits • produce more offspring and • grow rapidly in unpredictable environments. • Populations with K-selected traits • raise fewer offspring and • maintain relatively stable populations. • Most species fall between these two extremes. © 2012 Pearson Education, Inc. K-Selected Species • Poor colonizers • Slow maturity • Long-lived • Low fecundity • High investment in care for the young • Specialist • Good competitors r-Selected Species • Good colonizers • Reach sexual maturity rapidly • Short-lived • High fecundity • Low investment in care for the young • Generalists • Poor competitors Competition Among Species • Ecological Niche Concept • Ecological niche = the "role" a species "plays" in the ecosystem. • Contrast the ecological niche with the "habitat" which is the physical environment in which the organism lives. • The ecological niche of a species, therefore, includes not just the species’ habitat, but also the ways in which it interacts with other species and the physical environment. Competition Among Species • Competitive Exclusion Principle • No two similar species occupy the same niche at the same time. • Possible outcomes of competition • extinction of one species • resource partitioning: splitting the niche • character displacement: two similar species evolve in such a way as to become different from each other by accentuating their initial minor differences Keystone Species • A species whose presence in the community exerts a significant influence on the structure of that community. Keystone Species Keystone Species Human as Hyperkeystone Ecological Succession • The progressive change in the species composition of an ecosystem. Ecological Succession • Bare Substrate • Colonizing Stage Successionist Stage • Climax Stage Trophic Levels • 1st Trophic Level = Primary Producers • 2nd Trophic Level = Herbivores or Primary Consumers • 3rd Trophic Level and Higher = Carnivores or Secondary (or higher) Consumers • Highest Trophic Level = Top Carnivore • Decomposers Some Feeding Types • • • • • • • • • • Algal Grazers and Browsers Suspension Feeding Filter Feeding Deposit Feeding Benthic Animal Predators Plankton Pickers Corallivores Piscivores Omnivores Scavengers Overview: Chemical Cycles Biogeochemical cycles_________________ The various material circuits, which involve both the nutrient and physical components of an ecosystem. Carbon, Nitrogen, Phosphorous and Water are needed by every organism on Earth. C is needed to build organic molecules. N is needed for nucleic acids. P is needed for energy molecules. Water is needed to maintain life. How does every organism on Earth have access to these limited resources? Chemical Cycling Carbon cycle Nitrogen cycle Phosphorous cycle Water cycle