Cell 2 12 The Cell Anatomy: Physiology: The two categories of cells are prokaryotic cells and eukaryotic cells. Prokaryotic cell lack membrane bound organelles and their Genetic material is not surrounded by a nuclear membrane, while Eukaryotic cells have their Genetic material surrounded by a nuclear membrane. Prokaryotic organisms are classified within the domains Archaea and Bacteria, whereas eukaryotic organisms are classified within the domain Eukarya. The genetic material, usually DNA, sends information via the messenger RNA [mRNA]. Ribosomes of the cell read the information on the mRNA to from protein frameworks which are further modified within the cytoplasm by other organelles such as, for the eukaryotic cell, the golgi apparatus. The final product remains within the cell or for eukaryotic cells may form other organelles [such as lysosomes or peroxisomes] or transported by secretory vesicles to exit the cell. The structures of cells are unique with specific functions. Eukaryotic cells, for the most part tend to be larger than prokaryotic cells. All cells have a plasma membrane, cytoplasm and genetic material. Prokaryotic organisms are classified within the domains Archaea and Bacteria, whereas eukaryotic organisms are classified within the domain Eukarya. The structures of cells are unique with specific functions. Eukaryotic cells, for the most part tend to be larger than prokaryotic cells. Cell Eukaryotic Prokaryotic Prkarya Archaea Prkarya Bacteria Animal Plant Fungi Protists © 2019 Damaris-Lois Y Lang, Content Description licensed under a Creative Commons Attribution-NonCommercial 4.0 International License. 13 © 2019 Damaris-Lois Y Lang, Content illustrations and diagrams are licensed under a Creative Commons Attribution-NonCommercial 4.0 International License. CELL STRUCTURE EUKARYOTES PROKARYOTES BACTERIA ARCHAEA Size ~0.5–1 μM ~0.5–1 μM ~5–20 μM Surface area-tovolume ratio High High Low Nucleus No No Yes Genome characteristics • Single chromosome • Single chromosome • Multiple chromosomes • Circular • Circular • Linear • Haploid • Haploid • Haploid or diploid • Lacks histones • Contains histones • Contains histones Cell division Binary fission Binary fission Mitosis, meiosis Membrane lipid composition • Ester-linked • Ether-linked • Ester-linked • Straight-chain fatty acids • Branched isoprenoids • Straight-chain fatty acids • Bilayer • Bilayer or monolayer • Sterols • BilayerW Cell wall composition • Peptidoglycan, or • Pseudopeptidoglycan, or • Cellulose (plants, some algae) • None • Glycopeptide, or • Chitin (molluscs, insects, • Polysaccharide, or • Protein (S-layer), or crustaceans, and fungi) • Silica (some algae) Motility structures Rigid spiral flagella composed of flagellin • None • Most others lack cell walls Rigid spiral flagella composed of Flexible flagella and cilia archaeal flagellins composed of microtubules Membranebound organelles No No Yes Endomembrane system No No Yes (ER, Golgi, lysosomes) Ribosomes 70S 70S • 80S in cytoplasm and rough ER • 70S in mitochondria, chloroplasts © 2013 OpenStax CNX, Table content from textbook produced by OpenStax is licensed under a Creative Commons Attribution License 14 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 Sporangiospores are produced in a sporangium (Figure 24.9). Eukaryotic Fungi 635 635 (a) Apicomplexans are parasitic protists. They have a characteristic apical complex that enab ells. (b) Plasmodium, the causative agent of malaria, has a complex life cycle typical of apic dification of work by CDC) which include Paramecium and Tetrahymena, are a group of protists 10 to 3,000 micrometers in n rows, tufts, or spirals of tiny cilia. By beating their cilia synchronously or in waves, ciliates ca ements and ingest food particles. Certain ciliates have fused cilia-based structures that function l ns. Ciliates Fungi also are surrounded by a pellicle, providing protection without compromising agility includes protists that have organized their cilia into a plate-like primitive mouth, called an oral gr Figure 24.9 This bright field light micrograph shows the release of spores from a sporangium at th pture anda digest bacteria (Figure 23.15). captured in the oraloften groove enters a food vacuo called sporangiophore. The organism is aFood Mucor sp. fungus, a mold found indoors. (credit: m h digestive enzymes. byRussell) an exocytic vesicle that fuses at a specific r by Dr. Lucille Georg,Waste CDC; particles scale-bar are dataexpelled from Matt e, called the anal pore. In addition to a vacuole-based digestive system, Paramecium also uses Reproduction ichSexual are osmoregulatory vesicles that fill with water as it enters the cell by osmosis and then contrac e cell. 4.5 Fungal hyphae may be (a) septated or (b) coenocytic = "common"; = "cell") with of many nuclei In fungi, sexual reproduct Sexual reproduction introduces genetic(coenovariation into -cytic a population fungi. n a single hypha. A bright field light micrograph of (c) Phialophora richardsiae shows septa that divide the response to adverse environmental conditions. During sexual reproduction, credit c: modification of work by Dr. Lucille Georg, CDC; scale-bar data from Matt Russell) two mating types are pro Heterothallic m ngal hyphae may be (a) septated or (b) coenocytic (coeno- = "common"; -cytic = "cell") with many nuclei mating types are present in the same mycelium, it is called homothallic, or self-fertile. ngle hypha. A bright field light micrograph of (c) Phialophora richardsiae shows septa that divide the ive in environments that are moist andmycelia slightly and can grow with without light. They vary in but compatible, to reproduce sexually. c: different, modification of work by Dr. Lucille Georg, acidic, CDC; scale-bar data fromorMatt Russell) gen requirement. Most fungi are obligate aerobes, requiring oxygen to survive. Other species, such as the mycota that reside in the rumen of cattle, are are obligate anaerobes, in that they only use anaerobic respiration nxygen environments moist and acidic, and can grow with or without light.This They vary in will disrupt that their are metabolism or killslightly them. Yeasts are intermediate, being faculative anaerobes. means equirement. Most fungiofare obligate requiringbut oxygen to survive. Other species, such as the grow best in the presence oxygen using aerobes, aerobic respiration, can survive using anaerobic respiration when anotthat reside The in the rumen of cattle, areyeast are fermentation obligate anaerobes, in that use anaerobic respiration available. alcohol produced from is used in wine andthey beeronly production. n will disrupt their metabolism or kill them. Yeasts are intermediate, being faculative anaerobes. This means best in the presence of oxygen using aerobic respiration, but can survive using anaerobic respiration when mals, fungi are heterotrophs; they use complex organic compounds as a source of carbon, rather than fix carbon vailable. The alcohol produced from yeast fermentation is used in wine and beer production. rom the atmosphere as do some bacteria and most plants. In addition, fungi do not fix nitrogen from the re. Like animals, they must obtain it from their diet. However, unlike most animals, which ingest food and then nternally in specialized organs, fungi perform these steps in the reverse order; digestion precedes ingestion. First, ungi are heterotrophs; they use complex organic compounds as a source of carbon, rather than fix carbon mes are transported out of the hyphae, where they process nutrients in the environment. Then, the smaller molecules he atmosphere as do some bacteria and most plants. Inarea addition, fungi do As notwith fixanimal nitrogen by this external digestion are absorbed through the large surface of the mycelium. cells,from the the ke animals, they must obtain it from their diet. However, unlike most animals, which ingest food and then aride of storage is glycogen, rather than starch, as found in plants. lly in specialized organs, fungi perform these steps in the reverse order; digestion precedes ingestion. First, mostly saprobes (saprophyte is an equivalent term): organisms that derive nutrients from decaying organic transported out of the hyphae, where they process nutrients in the environment. Then,Fungal the smaller molecules 5ehey Paramecium primitive mouth an plant oralmaterial. groove) to ingest food, and an anal pore obtainProtist their nutrientshas from a dead or decomposing organic(called matter: mainly exoenzymes s external digestion are absorbed through the large surface area of the mycelium. As with animal cells, the down insoluble polysaccharides, such as the to cellulose and ligninexcess of dead wood, into readily absorbable eoofbreak vacuoles allow the organism excrete water. Cilia enable the organism to m storageThe is glycogen, rather than as foundarein thus plants. molecules. carbon, nitrogen, and starch, other elements released into the environment. Because of their micrograph”: modification of ecological work by dataas from Russell) etabolic pathways, fungi fulfill an important roleNIH; and arescale-bar being investigated potentialMatt tools in ly saprobes (saprophyte is an equivalent term): organisms that derive nutrients from decaying organic ation. For example, some species of fungi can be used to break down diesel oil and polycyclic aromatic btain their nutrients from dead or decomposing organic matter: mainly plant material. Fungal exoenzymes ons (PAHs). Other species take up heavy metals, such as cadmium and lead. k down insoluble polysaccharides, such as the cellulose and lignin of dead wood, into readily absorbable gi areThe parasitic, infecting eitherand plants or animals. Smut and elm disease affect plants, whereas athlete’s their ules. other elements are from thusDutch released into the environment. Because © 2013nitrogen, OpenStax CNX, Images and illustrations textbook produced by OpenStax is licensed under aof Creative Commons Attribution 15carbon, andidiasis (thrush) are medically important fungal infections in humans. In environments poor in nitrogen, ic pathways,License fungi 3.0 fulfill anThis important role and are being investigated as potentialsome tools in license. is availableecological for free at http://cnx.org/content/col11496/1.8 ort to predation of nematodes (small non-segmented roundworms). Species of Arthrobotrys fungi have a number Prokaryotic Chapter 4 | Prokaryotic Diversity Bacteria Archaea Figure 4.26 Sulfolobus, an archaeon of the class Crenarchaeota, oxidizes sulfur and sto granules. Another genus, Thermoproteus, is represented by strictly anaerobic organisms with an © 2013 OpenStax of CNX,85 Images illustrations textbookand, produced by OpenStaxare is licensed underThermoproteus a Creative Commons Attribution 16 °C.and They havefrom flagella therefore, motile. has a cellular me License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 Plant Cell 17 © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 Animal Cell © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution 18 License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 Fill out the table Structures Cell Organelles Molecules Atoms Work Overflow 19 Sketch and Label / Illustrations Functions/ Roles © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution 20 License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 (condensed) 21 © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution 22 License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 Extracellular Intracellular 23 © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 © 2013 OpenStax CNX, Images and illustrations from textbook produced by OpenStax is licensed under a Creative Commons Attribution 24 License 3.0 license. This is available for free at http://cnx.org/content/col11496/1.8 Personal Notes: -----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------25