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Chapter 50
An Introduction to Ecology
and the Biosphere
PowerPoint Lectures for
Biology, Seventh Edition
Neil Campbell and Jane Reece
Lectures by Chris Romero
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
• Overview: The Scope of Ecology
• Ecology
– Is the scientific study of the interactions
between organisms and the environment
• These interactions
– Determine both the distribution of organisms
and their abundance
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• Ecology
– Is an enormously complex and exciting area of
biology
– Reveals the richness of the biosphere
Figure 50.1
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• Concept 50.1: Ecology is the study of
interactions between organisms and the
environment
• Ecology
– Has a long history as a descriptive science
– Is also a rigorous experimental science
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Ecology and Evolutionary Biology
• Events that occur in ecological time
– Affect life on the scale of evolutionary time
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Organisms and the Environment
• The environment of any organism includes
– Abiotic, or nonliving components
– Biotic, or living components
– All the organisms living in the environment, the
biota
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• Environmental components
– Affect the distribution and abundance of
organisms
Kangaroos/km2
> 20
10–20
5–10
1–5
0.1–1
< 0.1
Limits of
distribution
Figure 50.2
Climate in northern Australia
is hot and wet, with seasonal
drought.
Southern Australia has
cool, moist winters and
warm, dry summers.
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Red kangaroos
occur in most
semiarid and arid
regions of the
interior, where
precipitation is
relatively low and
variable from
year to year.
Southeastern Australia
has a wet, cool climate.
Tasmania
• Ecologists
– Use observations and experiments to test
explanations for the distribution and
abundance of species
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Subfields of Ecology
• Organismal ecology
– Studies how an organism’s structure,
physiology, and (for animals) behavior meet
the challenges posed by the environment
Figure 50.3a
(a) Organismal ecology. How do humpback whales
select their calving areas?
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• Population ecology
– Concentrates mainly on factors that affect how
many individuals of a particular species live in
an area
(b) Population ecology.
What environmental
factors affect the
reproductive rate of
deer mice?
Figure 50.3b
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• Community ecology
– Deals with the whole array of interacting
species in a community
(c) Community ecology.
What factors influence
the diversity of species
that make up a
particular forest?
Figure 50.3c
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• Ecosystem ecology
– Emphasizes energy flow and chemical cycling
among the various biotic and abiotic
components
(d) Ecosystem ecology. What
factors control photosynthetic
productivity in a temperate
grassland ecosystem?
Figure 50.3d
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• Landscape ecology
– Deals with arrays of ecosystems and how they
are arranged in a geographic region
Figure 50.3e
(e) Landscape ecology. To what extent do the trees lining the
drainage channels in this landscape serve as corridors of
dispersal for forest animals?
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• The biosphere
– Is the global ecosystem, the sum of all the
planet’s ecosystems
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Ecology and Environmental Issues
• Ecology
– Provides the scientific understanding
underlying environmental issues
• Rachel Carson
– Is credited
with starting
the modern
environmental
movement
Figure 50.4
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• Most ecologists follow the precautionary
principle regarding environmental issues
• The precautionary principle
– Basically states that humans need to be
concerned with how their actions affect the
environment
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• Concept 50.2: Interactions between organisms
and the environment limit the distribution of
species
• Ecologists
– Have long recognized global and regional
patterns of distribution of organisms within the
biosphere
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• Many naturalists
– Began to identify broad patterns of distribution
by naming biogeographic realms
Palearctic
Nearctic
Tropic
of Cancer
(23.5N)
Oriental
Ethiopian
Equator
Neotropical
Figure 50.5
(23.5S)
Tropic of
Capricorn
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Australian
• Biogeography
– Provides a good starting point for
understanding what limits the geographic
distribution of species
Species absent
because
Yes
Dispersal
limits
distribution?
No
Area inaccessible
or insufficient time
Behavior
limits
distribution?
Yes
Habitat selection
Yes
No
Biotic factors
(other species)
limit
distribution?
No
Predation, parasitism, Chemical
competition, disease factors
Water
Abiotic factors
limit
distribution?
Oxygen
Salinity
pH
Soil nutrients, etc.
Temperature
Physical Light
factors Soil structure
Fire
Moisture, etc.
Figure 50.6
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Dispersal and Distribution
• Dispersal
– Is the movement of individuals away from
centers of high population density or from their
area of origin
– Contributes to the global distribution of
organisms
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Natural Range Expansions
• Natural range expansions
– Show the influence of dispersal on distribution
New areas
occupied
Figure 50.7
Year
1996
1989
1974
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Species Transplants
• Species transplants
– Include organisms that are intentionally or
accidentally relocated from their original
distribution
– Can often disrupt the communities or
ecosystems to which they have been
introduced
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Behavior and Habitat Selection
• Some organisms
– Do not occupy all of their potential range
• Species distribution
– May be limited by habitat selection behavior
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Biotic Factors
• Biotic factors that affect the distribution of
organisms may include
– Interactions with other species
– Predation
– Competition
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• A specific case of an herbivore limiting
distribution of a food species
EXPERIMENT
W. J. Fletcher tested the effects of two algae-eating animals, sea urchins and limpets, on seaweed
abundance near Sydney, Australia. In areas adjacent to a control site, either the urchins, the limpets, or both were removed.
RESULTS
Fletcher observed a large difference in seaweed growth between areas with and without sea urchins.
100
Sea
urchin
Seaweed cover (%)
80
Both limpets
and urchins
removed
Only
urchins
removed
60
Limpet
40
Only limpets removed
Control (both
urchins and
limpets present)
20
Removing both
limpets and
urchins or
removing only
urchins increased
seaweed cover
dramatically.
Almost no
seaweed grew
in areas where
both urchins and
limpets were
present, or where
only limpets were
removed.
0
August
1982
Figure 50.8
February
1983
August
1983
February
1984
CONCLUSION
Removing both limpets and urchins resulted in the greatest increase of seaweed cover, indicating that both
species have some influence on seaweed distribution. But since removing only urchins greatly increased seaweed growth while
removing only limpets had little effect, Fletcher concluded that sea urchins have a much greater effect than limpets in limiting
seaweed distribution.
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Abiotic Factors
• Abiotic factors that affect the distribution of
organisms may include
– Temperature
– Water
– Sunlight
– Wind
– Rocks and soil
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Temperature
• Environmental temperature
– Is an important factor in the distribution of
organisms because of its effects on biological
processes
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Water
• Water availability among habitats
– Is another important factor in species
distribution
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Sunlight
• Light intensity and quality
– Can affect photosynthesis in ecosystems
• Light
– Is also important to the development and
behavior of organisms sensitive to the
photoperiod
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Wind
• Wind
– Amplifies the effects of temperature on
organisms by increasing heat loss due to
evaporation and convection
– Can change the morphology of plants
Figure 50.9
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Rocks and Soil
• Many characteristics of soil limit the distribution
of plants and thus the animals that feed upon
them
– Physical structure
– pH
– Mineral composition
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Climate
• Four major abiotic components make up
climate
– Temperature, water, sunlight, and wind
• Climate
– Is the prevailing weather conditions in a
particular area
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• Climate patterns can be described on two
scales
– Macroclimate, patterns on the global, regional,
and local level
– Microclimate, very fine patterns, such as those
encountered by the community of organisms
underneath a fallen log
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Global Climate Patterns
• Earth’s global climate patterns
– Are determined largely by the input of solar
energy and the planet’s movement in space
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• Sunlight intensity
– Plays a major part in determining the Earth’s
climate patterns
LALITUDINAL VARIATION IN SUNLIGHT INTENSITY
North Pole
60N
Low angle of incoming sunlight
30N
Tropic of
Cancer
Sunlight directly overhead
0 (equator)
Tropic of
Capricorn
30S
Low angle of incoming sunlight
60S
South pole
Figure 50.10
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Atmosphere
SEASONAL VARIATION IN SUNLIGHT INTENSITY
60N
June solstice: Northern
Hemisphere tilts toward
sun; summer begins in
Northern Hemisphere;
winter begins in
Southern Hemisphere.
30N
0 (equator)
March equinox: Equator faces sun directly;
neither pole tilts toward sun; all regions on Earth
experience 12 hours of daylight and 12 hours of
darkness.
30S
Constant tilt
of 23.5
September equinox: Equator faces sun
directly; neither pole tilts toward sun; all
regions on Earth experience 12 hours of
daylight and 12 hours of darkness.
Figure 50.10
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December solstice: Northern
Hemisphere tilts away from sun;
winter begins in Northern
Hemisphere; summer begins
in Southern Hemisphere.
• Air circulation and wind patterns
– Play major parts in determining the Earth’s
climate patterns
GLOBAL AIR CIRCULATION AND PRECIPITATION PATTERNS
60N
30N
Descending
dry air
absorbs
moisture
0 (equator)
30S
0
60S
Figure 50.10
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Ascending
moist air
releases
moisture
Descending
dry air
absorbs
moisture
Arid
zone
Tropics
Arid
zone
GLOBAL WIND PATTERNS
Arctic
Circle
60N
Westerlies
30N
Northeast trades
Doldrums
0
(equator)
Southeast trades
30S
Westerlies
60S
Antarctic
Circle
Figure 50.10
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Regional, Local, and Seasonal Effects on Climate
• Various features of the landscape
– Contribute to local variations in climate
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Bodies of Water
• Oceans and their currents, and large lakes
– Moderate the climate of nearby terrestrial
environments
2 Air cools at
high elevation.
3 Cooler
air sinks
over water.
4 Cool air over water
moves inland, replacing
rising warm air over land.
Figure 50.11
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1 Warm air
over land rises.
Mountains
• Mountains have a significant effect on
– The amount of sunlight reaching an area
– Local temperature
– Rainfall
1 As moist air moves in
off the Pacific Ocean and
encounters the westernmost
mountains, it flows upward,
cools at higher altitudes,
and drops a large amount
of water. The world’s tallest
trees, the coastal redwoods,
thrive here.
2 Farther inland, precipitation
increases again as the air
moves up and over higher
mountains. Some of the world’s
deepest snow packs occur here.
3 On the eastern side of the
Sierra Nevada, there is little
precipitation. As a result of
this rain shadow, much of
central Nevada is desert.
Wind
direction
East
Pacific
Ocean
Sierra
Nevada
Coast
Range
Figure 50.12
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Seasonality
• The angle of the sun
– Leads to many seasonal changes in local
environments
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• Lakes
– Are sensitive to seasonal temperature change
– Experience seasonal turnover
Lake depth (m)
In winter, the coldest water in the lake (0°C) lies just
below the surface ice; water is progressively warmer at
deeper levels of the lake, typically 4–5°C at the bottom.
O2 (mg/L)
0
4
Spring
Winter
8
12
8
16
2
4
4
4
4C
24
O2 concentration
0
Lake depth (m)
1
2 In spring, as the sun melts the ice, the surface water warms to 4°C
and sinks below the cooler layers immediately below, eliminating the
thermal stratification. Spring winds mix the water to great depth,
bringing oxygen (O2) to the bottom waters (see graphs) and
nutrients to the surface.
O2 (mg/L)
0
4 8
12
8
16
4
4
4
4
4
4C
24
High
Medium
O2 (mg/L)
0
4
8
12
8
16
24
Figure 50.13
4
Autumn
4
4
4
4C
4
In autumn, as surface water cools rapidly, it sinks below the
underlying layers, remixing the water until the surface begins
to freeze and the winter temperature profile is reestablished.
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4
Thermocline
3
22
20
18
8
6
5
4C
Summer
Lake depth (m)
Lake depth (m)
Low
O2 (mg/L)
0
4
8
12
8
16
24
In summer, the lake regains a distinctive thermal profile, with
warm surface water separated from cold bottom water by a narrow
vertical zone of rapid temperature change, called a thermocline.
Microclimate
• Microclimate
– Is determined by fine-scale differences in
abiotic factors
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Long-Term Climate Change
• One way to predict future global climate
change
– Is to look back at the changes that occurred
previously
Current
range
Predicted
range
Overlap
Figure 50.14
(a) 4.5C warming over
next century
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(b) 6.5C warming over
next century
• Concept 50.3: Abiotic and biotic factors
influence the structure and dynamics of aquatic
biomes
• Varying combinations of both biotic and abiotic
factors
– Determine the nature of Earth’s many biomes
• Biomes
– Are the major types of ecological associations
that occupy broad geographic regions of land
or water
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• The examination of biomes will begin with
Earth’s aquatic biomes
30N
Tropic of
Cancer
Equator
Tropic of
Capricorn
Continental
shelf
30S
Key
Figure 50.15
Lakes
Rivers
Estuaries
Coral reefs
Oceanic pelagic
zone
Intertidal zone
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Abyssal zone
(below oceanic
pelagic zone)
• Aquatic biomes
– Account for the largest part of the biosphere in
terms of area
– Can contain fresh or salt water
• Oceans
– Cover about 75% of Earth’s surface
– Have an enormous impact on the biosphere
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• Many aquatic biomes
– Are stratified into zones or layers defined by
light penetration, temperature, and depth
Intertidal zone
Neritic zone
Littoral
zone
Limnetic
zone
0
Oceanic zone
Photic zone
200 m
Continental
shelf
Pelagic
zone
Benthic
zone
Photic
zone
Aphotic
zone
Pelagic
zone
Benthic
zone
Aphotic
zone
2,500–6,000 m
Abyssal zone
(deepest regions of ocean floor)
(a) Zonation in a lake. The lake environment is generally classified on the basis
of three physical criteria: light penetration (photic and aphotic zones),
distance from shore and water depth (littoral and limnetic zones), and
whether it is open water (pelagic zone) or bottom (benthic zone).
Figure 50.16a, b
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(b) Marine zonation. Like lakes, the marine environment is generally
classified on the basis of light penetration (photic and aphotic zones),
distance from shore and water depth (intertidal, neritic, and oceanic
zones), and whether it is open water (pelagic zone) or bottom (benthic
and abyssal zones).
• Lakes
LAKES
Figure 50.17
An oligotrophic lake in
Grand Teton, Wyoming
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A eutrophic lake in Okavango
delta, Botswana
• Wetlands
WETLANDS
Figure 50.17
Okefenokee National Wetland Reserve in Georgia
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• Streams and rivers
STREAMS AND RIVERS
Figure 50.17
A headwater stream in the
Great Smoky Mountains
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The Mississippi River far
form its headwaters
• Estuaries
ESTUARIES
Figure 50.17 An estuary in a low coastal plain of Georgia
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• Intertidal zones
INTERTIDAL ZONES
Figure 50.17
Rocky intertidal zone on the Oregon coast
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• Oceanic pelagic biome
OCEANIC PELAGIC BIOME
Figure 50.17 Open ocean off the island of Hawaii
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• Coral reefs
CORAL REEFS
Figure 50.17
A coral reef in the Red Sea
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• Marine benthic zone
MARINE BENTHIC ZONE
Figure 50.17 A deep-sea hydrothermal vent community
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• Concept 50.4: Climate largely determines the
distribution and structure of terrestrial biomes
• Climate
– Is particularly important in determining why
particular terrestrial biomes are found in
certain areas
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Climate and Terrestrial Biomes
• Climate has a great impact on the distribution
of organisms, as seen on a climograph
Temperate grassland
Desert
Tropical forest
Annual mean temperature (ºC)
30
Temperate
broadleaf
forest
15
Coniferous
forest
0
Arctic and
alpine
tundra
15
100
Figure 50.18
200
300
Annual mean precipitation (cm)
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400
• The distribution of major terrestrial biomes
30N
Tropic of
Cancer
Equator
Tropic of
Capricorn
30S
Key
Tropical forest
Figure 50.19
Savanna
Desert
Chaparral
Temperate grassland
Temperate broadleaf forest
Coniferous forest
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Tundra
High mountains
Polar ice
General Features of Terrestrial Biomes
• Terrestrial biomes
– Are often named for major physical or climatic
factors and for their predominant vegetation
• Stratification
– Is an important feature of terrestrial biomes
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• Tropical forest
TROPICAL FOREST
Figure 50.20
A tropical rain forest in Borneo
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• Desert
DESERT
Figure 50.20 The Sonoran Desert in southern Arizona
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• Savanna
SAVANNA
Figure 50.20
A typical savanna in Kenya
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• Chaparral
CHAPARRAL
Figure 50.20
An area of chaparral in California
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• Temperate grassland
TEMPERATE GRASSLAND
Figure 50.20
Sheyenne National Grassland in North Dakota
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• Coniferous forest
CONIFEROUS FOREST
Rocky Mountain National Park in Colorado
Figure 50.20
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• Temperate broadleaf forest
TEMPERATE BROADLEAF FOREST
Figure 50.20
Great Smoky Mountains National Park in North Carolina
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• Tundra
TUNDRA
Figure 50.20
Denali National Park, Alaska, in autumn
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