. There are ~ 12,000 annelid species in 2 classes. Exam 3 Mar 22 Cycloneuralia Mollusca I Mollusca II Mollusca III There are ~ 12,000 annelid species in 2 classes. Annelids are triploblastic protostomes with true coeloms. Gastrula Clitellata ~4,000 spp. Polychaeta ~8,000 spp. Blastopore Mouth 1 . Annelids share 4 features 1. Setae Annelids share 4 features 2. Metamerism Annelids share 4 features 2. Metamerism Annelids share 4 features 3. Movement via circular & longitudinal muscles Circular muscles segments Longitudinal muscles septa Earthworm cross section 2 . Annelids share 4 features 3. Movement via circular & longitudinal muscles Annelida is divided into 2 classes. Polychaeta Annelids share 4 characteristics. 4. Closed Circulatory System Class Polychaeta have many setae on parapodia. Clitellata segments Many setae on parapodia Few setae; no parapodia Clitellum for reproduction Parapodia with setae Nereis Sense organs & mouth 3 . Parapodia are un-jointed appendages that bear setae. Polychaetes can be motile (can move around) or sedentary (anchor to substrate). setae Acicula A benthic motile polychaete with many parapodia & setae A polychaete parapodium under a microscope Tube worms are marine polychaetes that have adapted to live along deep-sea hydrothermal vents. Tube worms are peizophiles. Sessile infaunal christmas tree worms that live in coral Tube worms have chemosynthetic bacterial endosymbionts. Bacteria Tubeworms Bacteriocytes https://www.youtube.com/watch?v=dg6ZewBTRj8 4 . Class Clitellata have few setae, no parapodia and reproduce via the clitellum. This class includes earthworms & leeches. Clitellata mate via the clitellum. Clitellum Earthworm egg case Subclass Oligochaeta includes earthworms. Earthworms play a critical role in ecosystems. Few setae 5 . However, invasive earthworms have over populated some U.S. forests. Thick, healthy leaf litter layer (dark brown) prior to earthworm invasion. When leaf litter is broken down too quickly, native plant populations are reduced. Very thin leaf litter layer, most of which has been converted to soil (dark brown) after earthworm invasion. Some species of earthworms are used in vermicomposting. Subclass Hirudinea are the leeches. Most leeches are hematophagous ectoparasites of both vertebrates & invertebrates. 6 . Historically, leeches were used in a medical process called blood-letting. Recently leech therapy has been used in modern operating rooms. Hirudo medicinalis - medicinal leech 7