Flower Morphology Basic Floral Structure • Essential/ reproductive structures - concerned with sporogenesis and gametogenesis - pistil/ carpel/ gynoecium and stamen/androecium • Accessory/ sterile structures - does not participate in spore/ sex cell formation - perianth, receptacle Presence/ absence of floral parts (corolla, calyx, stamen, pistil) Complete flower VS Incomplete flower Presence/ absence of essential reproductive structures • Perfect/ Bisexual flower VS Imperfect flower Symmetry • Actinomorphic/ Regular flower VS Zygomorphic flower Petal Attachment • Apopetalous/ Polypetalous flower VS Gamopetalous flower VS Apetalous flower Corolla tube Ovary position • Superior/ hypogynous flower VS Inferior/ epigynous flower VS Semi- inferior/ perigynous flower FLOWERS: Placentation. • 1. 2. 3. 4. 5. Pattern of ovule attachment and distribution in an ovary. Axile - ovules are borne on placentae on the central axis of a compound ovary. Basal - has a single ovule that is almost at the base of the ovary. Marginal - ovules are borne along the fused margins of placenta of a single ovary. Parietal - ovules are borne in placentae located on the inner surface of the outer wall of the ovary. Free central - ovules are borne on placentae on a free, central column within an ovary that has only one locule. FLOWERS: Composites. • A single flower with many tiny separate flowers densely clustered together on a common receptacle. • Example: Asteraceae family. 1. Sunflowers 2. Daisies 3. Chrysanthemums FLOWERS: Composites. • Inflorescence: head or capitate. • 2 kinds of flowers in the head: 1. Ray flowers: corolla looks like one petal. • found on the outer edge of the head 2. Disk flowers: corolla forms a tube of united or fused petals. • Constitutes the inner circle. • In some composites, the disk flowers are missing and the head consists only of ray flowers. • In others, it is the ray flowers that are missing and the head consists only of disk flowers. FLOWERS: Grasses. • Grasses: flowers, highly modified. • The individual grass flower, the floret is enveloped by a pair of bracts or modified leaves. • The outer and larger bract is termed the lemma and the inner and smaller one is called the palea. • The palea encloses the stamens and the pistil. • When the flower is mature, the stamens are large and usually extend well above the stigma. • The stigma is feathery and large Cross Pollination VS Self Pollination (Advantage and Disadvantage?) Microsporogenesis & Microgametogenesis Anther Cross Section Anther Cross Section Epidermis Vascular strand Endothecium Stomium Pollen grain/s Anther Cross Section Pollen sac Tapetum layer Anther Cross Section • Tapetum layer parenchyma cells that surrounds the Pollen sac • Epidermis Outer layer of anther • Endothecium Parenchyma cells that constitutes the rest of the anther Pollen grain Intine – inner layer of the pollen grain Exine – outer layer Tube Nucleus – give rise to the Pollen tube Generative Nucleus – for fertilization Pollen Grain with Pollen TUBE X-S ovule Ovule (3) (2) (2) (to be FERTILIZED) Fertilization Ovules becomes the SEED Seed (Dicot) Plumule Epicotyl Seed coat/testa Hypocotyl Radicle Cotlyedon Seed (Monocot) Monocot VS Dicot growth pattern HYPOGEAL EPIGEAL Seed germination 1.Absorption of water in the micropyle. 2. Increased metabolic rates 3. Cellular respiration proceeds 4. Cell division and growth proceeds. 5. Rupture of the seed coat 6. Embryo sprouting disperse their seed without any help from an external vector a vector or secondary agent is used to disperse seeds Ovary to Fruit (Matured, Ripened Ovaries) Infloresence Spadix infloresence Raceme Spike Corymb Head Panicle Umbel Cyme Fruits According to Flower Origin Multiple Simple Aggregate Nature of Pericarp Fleshy Dry Nature of Dispersal Berry Hesperidium Pepo Drupe Pome Dehiscent Indehiscent Legume Follicle Capsule Silique Achene Grain Samara Nut Multiple Fruit Pineapple (Ananas comosus) formed from an inflorescence. Each flower in the inflorescence produces a fruit, but these mature into a single mass in which each flower has produced a true fruit Jackfruit (Artocarpus heterophyllus) fruit that develops from the merger of several ovaries that were separate in a single flower. Aggregate Fruit Raspberry (Rubus strigosus) sugar-apple, or sweetsop (Annona squamosa) Blackberry (Rubus fruticosus) Fruits According to Flower Origin Multiple Simple Aggregate Nature of Pericarp Fleshy Dry Nature of Dispersal Berry Hesperidium Pepo Drupe Pome Dehiscent Indehiscent Legume Follicle Capsule Silique Achene Grain Samara Nut Fruit- Berry A fleshy fruit without a stone produced from a single flower containing one ovary Banana (Musa acuminata) Pepper (Capsicum annuum) Tomato (Solanum lycopersicum) Hesperidium A fleshy fruit with tough, leathery, aromatic rind. Orange (Citrus sinensis) Calamondin (Citrus microcarpa) Pepo hard, thick rind; typical fruit of the gourd family (Cucurbitaceae) Watermelon (Citrullus lanatus_ cucumber (Cucumis sativus) Drupe one-seeded fruit with the pericarp distinctly divided into thin skin-like exocarp, thick fleshy mesocarp and hard, stony endocarp coconut (Cocos nucifera) Coffee (Coffea arabica) Mango (Mangifera indica) Pome fruit with papery pericarp Pear (Pyrus amydaliformes) apple (Malus domestica) Fruits According to Flower Origin Multiple Simple Aggregate Nature of Pericarp Fleshy Dry Nature of Dispersal Berry Hesperidium Pepo Drupe Pome Dehiscent Indehiscent Legume Follicle Capsule Silique Achene Grain Samara Nut Legume/ Pod splits along two seams peanut(Arachis hypogaea) Tamarind (Tamarindus indica) splits in a variety of ways Capsule Lady finger’s (Abelmoschus esculentus) Cotton (Gossypium hirsutum) Follicle splits along one seam Milkweed (Asclepias syriaca) Silique two fused carpels that separate, leaving a persistent wall between them Mustard (Brassica campestris) Fruits According to Flower Origin Multiple Simple Aggregate Nature of Pericarp Fleshy Dry Nature of Dispersal Berry Hesperidium Pepo Drupe Pome Dehiscent Indehiscent Legume Follicle Capsule Silique Achene Grain Samara Nut Grain one seeded the coat of which is completely fused to the inner surface of the pericarp Rice (Oryza sativa) Achene one seed can be separated from the ovary wall except at point of attachment to the inside of the pericarp Nut one seeded fruit similar to achene but with a very hard and thick pericarp Walnut (Juglans regia) Chestnut (Castanea dentata) Oak (Quercus alba) Samara an achene-like fruit with wing-like outgrowth Narra (Pterocarpus inducus) Free Central - ovules are attached to a central axis of a uniloculate ovary Axile - ovules are attached to a central axis in a multiloculate ovary Parietal - ovules are found along the walls of a uniloculate ovary Basal Marginal