ANIMAL EXCRETORY SYSTEM GRADE 10 OSMOREGULATION • OSMOREGULATION IS AN IMPORTANT PART OF HOMEOSTASIS. • OSMOREGULATION IS THE BALANCING OF WATER AND SALTS IN THE BODY. • THE OSMOREGULATORY SYSTEM IN ANIMALS ALSO REMOVES METABOLIC WASTES IN A PROCESS CALLED EXCRETION. The breakdown of proteins, amino acids and nucleic acids releases nitrogen and excess nitrogen must be excreted. Amino acids are broken down to generate energy or forms fats or carbohydrates. To do this the NH2 must be removed. NITROGENOUS WASTE PRODUCTS • THE REMOVAL OF NITROGEN IS DONE IN THE FORM OF AMMONIA, UREA OR URIC ACID. • THE PRODUCTION OF AMMONIA, UREA AND URIC ACID REQUIRE DIFFERENT AMOUNTS OF WATER AND ENERGY. AMMONIA • NH2 (FROM AMINO ACID) + H+ = AMMONIA • THIS REQUIRES VERY LITTLE ENERGY BUT IS VERY TOXIC SO NEEDS A LOT OF WATER TO DILUTE IT AS IT LEAVES THE BODY. • AMMONIA IS EXCRETED BY FISH AND AQUATIC ANIMALS WHO ARE SURROUNDED BY WATER. UREA • UREA IS LESS TOXIC THAN AMMONIA SO CAN BE EXCRETED IN MORE CONCENTRATED SOLUTIONS. • THE PRODUCTION OF UREA REQUIRES ENERGY BUT ALLOWS THE BODY TO SAVE WATER – THIS IS IMPORTANT FOR TERRESTRIAL (LAND) MAMMALS. • 2(NH2) + CO2 = UREA • SHARKS, AMPHIBIANS, MAMMALS URIC ACID • URIC ACID IS FORMED BY A COMPLEX SERIES OF ENZYMATIC REACTIONS. • IT REQUIRES A LOT OF ENERGY TO MAKE URIC ACID. • IT IS NOT VERY TOXIC SO CAN BE VERY CONCENTRATED. • IT IS NOT VERY WATER SOLUBLE – IT IS GOOD WHEN WATER IS IN SHORT SUPPLY. • INSECTS, REPTILES, BIRDS. • IN REPTILES AND BIRDS, URIC ACID GOES FROM THE KIDNEYS TO THE CLOACA – HERE WASTE FROM KIDNEYS, DIGESTIVE AND REPRODUCTIVE SYSTEMS COMBINE. • THEY ARE REFLUXED INTO THE INTESTINE WHERE WATER IS RE-ABSORBED. • THE WHITE PART OF BIRD FAECES IS URIC ACID. • URIC ACID IS EASY TO STORE INSIDE BIRD AND REPTILE EGGS AS IT IS NON-TOXIC AND CONCENTRATED. • HUMANS CAN PRODUCE URIC ACID FROM PURINE AND PYRIMIDINE NUCLEIC ACID BUT IN SOME PEOPLE IT CAUSES GOUT (JOINT PAIN). EXCRETORY ORGANS IN INVERTEBRATES • INVERTEBRATES HAVE TUBULAR EXCRETORY ORGANS FOR OSMOREGULATION. • PLANARIANS (FLATWORMS) HAVE 2 STRANDS OF BRANCHING TUBULES THAT RELEASE WASTE THROUGH EXCRETORY PORES. • EACH STRAND HAS A NUMBER OF FLAME CELLS THAT HAVE BEATING CILIA THAT PUSH/PROPEL EXCESS WATER AND WASTE TO THE PORES. • EARTHWORMS ARE DIVIDED INTO SEGMENTS – EACH SEGMENT HAS A PAIR OF NEPHRIDIA (EXCRETORY STRUCTURES). • EACH NEPHRIDIUM IS A TUBULE WITH A CILIATED OPENING AND AN EXCRETORY PORE. • AS FLUID FROM THE COELOM MOVES THROUGH THE TUBULE NUTRIENTS ARE ABSORBED BY CAPILLARIES (SMALL VEINS). • URINE (WATER, WASTES, SALTS) IS PASSED OUT THROUGH THE EXCRETORY PORES. • EARTHWORMS EXCRETE A VERY DILUTE URINE – THEY PRODUCE A VOLUME OF URINE EQUAL TO 60% OF THEIR BODY WEIGHT. INSECTS • INSECTS HAVE A LONG MALPIGHIAN TUBULE ATTACHED TO THE GUT. • URIC ACID IS ACTIVELY TRANSPORTED FROM HEMOLYMPH. • WATER FOLLOWS A SALT GRADIENT ESTABLISHED BY THE ACTIVE TRANSPORT OF K+. • WATER AND NUTRIENTS ARE REABSORBED BY THE RECTUM. • URIC ACID IS EXPELLED VIA THE ANUS. ARTHROPODS (EXOSKELETON) • IN AQUATIC ARTHROPODS, NITROGENOUS WASTE IS REMOVED BY DIFFUSION ACROSS THE GILLS. • SOME CRUSTACEANS (SHELL FISH) HAVE AN ORGAN CALLED THE GREEN GLANDS WHERE FLUID COLLECTS IN THE VENTRAL PORTION OF THE HEAD REGION. • IN SHRIMPS AND PILLBUGS THE EXCRETORY ORGANS ARE IN THE MAXILLARY SEGMENT AND ARE CALLED MAXILLARY GLANDS. • SPIDERS AND OTHER ARACHNIDS HAVE A COXAL GLAND WHICH ARE USED FOR EXCRETION.