Biology, Seventh Edition Solomon • Berg • Martin Chapter 31 Plant Structure, Growth, and Differentiation Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Plants • Herbaceous –In temperate climates, aerial parts die back –Annuals, biennials, perennials • Woody –In temperate climates, aerial parts persist –Perennials Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Annuals • Grow, reproduce, and die in one year or less –Corn –Geranium –Marigold Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Duckweeds, with Wolffia on frog’s back Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Biennials • Take two years to complete their life cycles before dying –Carrot –Queen Anne’s lace Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Perennials • Have the potential to live for more than two years –Asparagus –Oak trees Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Eucalyptus regnans Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Life history strategies • Long-lived trees compete successfully against smaller plants in favorable environments (e.g., tropical rain forests) • Small, short-lived plants thrive in unfavorable environments (e.g., deserts after a rainy spell) Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular plant body • Root system –Generally underground –Obtains water and dissolved nutrient minerals –Anchor plant in place Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation The plant body: Arabidopsis thaliana Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular plant body, cont. • Shoot system consists of –A vertical stem bearing leaves (organs of photosynthesis) –Reproductive structures (flowers and fruits in flowering plants) –Buds (undeveloped embryonic shoots that develop on stems) Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular plant body, shoot system, cont. • Generally aerial • Obtains sunlight and exchanges gases, such as CO2, oxygen, and water vapor Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular plant body, cont. • Plant body is composed of –Ground tissue system –Vascular tissue system –Dermal tissue system • Although separate organs exist, tissue systems are integrated throughout plant body Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation The three tissue systems in the plant body Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Ground tissue system • Parenchyma tissue • Collenchyma tissue • Sclerenchyma tissue Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Parenchyma cells Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Collenchyma cell Schlerenchyma cells Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Parenchyma tissue • Composed of living parenchyma cells with thin primary cell walls • Functions –Photosynthesis –Storage –Secretion Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Collenchyma tissue • Consists of collenchyma cells with unevenly thickened primary cell walls • Function is to provide flexible structural support Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Sclerenchyma tissue • Composed of sclerenchyma cells (sclereids or fibers) with –Primary cell walls –Secondary cell walls • Function is to provide structural support, although Sclerenchyma cells are often dead at maturity Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Parenchyma cell type Sclerenchyma cell type Collenchyma cell type Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular tissue system • Conducts materials throughout plant body • Provides strength and support • Consists of –Xylem –Phloem Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition Tracheid (xylem cell CHAPTER 31 Plant Structure, Growth, and Differentiation Sieve tube element Vessel element (xylem cell) Copyright © 2005 Brooks/Cole — Thomson Learning Phloem tissue Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular tissue system, cont. • Xylem –Complex tissue conducting water and dissolved nutrient minerals –Conducting cells are tracheids and vessel elements Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Vascular tissue system, cont. • Phloem –Complex tissue conducting sugar in solution –Conducting cells are sieve tube elements; they are assisted by companion cells Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Tracheid Vessel element Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Sieve tube element Companion cell Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Dermal tissue system • Outer protective covering of the plant body • Consists of –Epidermis –Periderm Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Dermal tissue system, cont. • Epidermis –Complex tissue covering herbaceous plant body –Epidermis of aerial parts secretes a waxy cuticle to reduce water loss –Stomata permit gas exchange –Trichomes (outgrowths or hairs) Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Dermal tissue system, cont. • Periderm –Complex tissue covering woody parts of plant body in woody plants Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Epidermal cell Guard cell Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Trichome Cork cell Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Growth in plants versus animals • Localized in meristems • Involves –Cell division –Cell elongation –Cell differentiation Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Primary growth • Increase in stem or root length • Occurs in all plants • Secondary growth • Increase in stem or root girth • Localized • Typically occurs in long cylinders of meristematic cells throughout length of older stems and roots Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Apical meristems • Localized –At tips of roots –At tips of shoots –Within buds of stems • Activity herein results in primary growth Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation A root tip Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation • Lateral meristems • Vascular cambium • Cork cambium • Responsible for secondary growth Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Longitudinal section through shoot apex Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 31 Plant Structure, Growth, and Differentiation Secondary growth Copyright © 2005 Brooks/Cole — Thomson Learning