Biology, Seventh Edition Solomon • Berg • Martin Chapter 40 Neural Regulation Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Nerve nets • Radially symmetrical invertebrates • Nerve cells scattered throughout the body Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Hydra’s nerve net Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Radial nervous systems • Radially symmetrical invertebrates • Nerve ring and nerves that extend to various parts of the body Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Bilateral nervous systems • Bilaterally symmetrical animals • Nerve cells form nerves, nerve cords, ganglia, and brain • Sense organs concentrated in the head region Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Bilateral nervous systems • Wide range of responses –Increased number of association neurons –More complex synaptic contacts permit a planarian flatworms Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Planarian flatworms • Cerebral ganglia • Two solid ventral nerve cords connected by transverse nerves Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Ladder-type nervous system in flatworms Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Annelids and arthropods • Ventral nerve cord • Numerous ganglia • Cerebral ganglia of arthropods have specialized regions • Cephalopod mollusks • Complex nervous systems • Neurons in a central region Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Annelid and arthropod nervous systems Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Evolutionary trends • Increased numbers and concentration of nerve cells • Specialization of function • Increased number of association neurons • More complex synaptic contacts • Formation of a head Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Vertebrate nervous system • Central nervous system (CNS) –Brain –Dorsal, tubular spinal cord • Peripheral nervous system (PNS) –Sensory receptors –Nerves Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Vertebrate brain development • Brain and spinal cord arise from the neural tube • Anterior end of the tube differentiates into –Forebrain –Midbrain –Hindbrain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Early development of the vertebrate nervous system Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Section through the human brain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Hindbrain subdivides into • Metencephalon • Myelencephalon Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Myelencephalon develops into the medulla • Vital centers • Other reflex centers • Fourth ventricle –Communicates with the central canal of the spinal cord Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Metencephalon gives rise to the cerebellum • Muscle tone • Posture • Equilibrium • Pons –Connects various parts of the brain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Midbrain • Largest part of the brain in fishes and amphibians • Main association area, linking sensory input and motor output • Center for visual and auditory reflexes in reptiles, birds, and mammals Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Brain stem • Medulla • Pons • Midbrain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Forebrain differentiates to form the diencephalon and telencephalon • Diencephalon develops into the thalamus and hypothalamus • Thalamus • Relay center for motor and sensory information Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Hypothalamus • Controls autonomic functions • Links nervous and endocrine systems • Controls temperature, appetite, and fluid balance • Emotional and sexual responses Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Telencephalon develops into the cerebrum and olfactory bulbs • Cerebrum • Divided into right and left hemispheres • In fishes and amphibians, integrates sensory information Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • In birds, the corpus striatum controls complex behavior patterns, such as flying and singing • In mammals, the neocortex accounts for a large part of the cerebral cortex Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Human brain and spinal cord • Protected by bone and three meninges –Dura mater –Arachnoid –Pia mater • Cushioned by cerebrospinal fluid (CSF) Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Spinal cord • Transmits impulses to and from the brain • Controls many reflex actions • Gray matter contains nuclei that serve as reflex centers Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Structure of the spinal cord Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Ascending tracts • Transmit information to the brain • Descending tracts • Transmit information from the brain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Withdrawal reflex • Sensory receptors • Sensory neurons • Interneurons • Motor neurons • Effectors Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Reflex action Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Human cerebral cortex • Gray matter –Forms folds or convolutions • Fissures –Deep furrows between the folds Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Cerebrum • Sensory areas –Receive incoming sensory information • Motor areas –Control voluntary movement Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Association areas –Link sensory and motor areas –Learning –Language –Thought –Judgment Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • The cerebrum consists of lobes • Frontal lobes • Parietal lobes • Temporal lobes • Occipital lobes Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Human brain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • White matter of the cerebrum • Lies beneath the cerebral cortex • Corpus callosum –Large band of white matter –Connects right and left hemispheres Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Basal ganglia –Cluster of nuclei within the white matter –Important centers for motor function Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Sleep–wake pattern • Regulated by the hypothalamus and brain stem Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Alpha wave patterns • Relaxed states • Beta wave patterns • Accompany heightened mental activity Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Theta and delta waves • Slower frequency • Higher-amplitude • Non-REM sleep Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation EEGs showing electrical activity in the brain Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Reticular activating system (RAS) • Arousal system • Neurons filter sensory input • Selects which information is transmitted to the cerebrum Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Non-REM sleep • Electrical activity of the cerebral cortex slows • Metabolic rate slows • REM sleep • Dreaming Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Suprachiasmatic nucleus • Main biological clock • Receives information about light and dark • Transmits it to other nuclei that regulate sleep Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Limbic system • Emotional aspects of behavior • Motivation • Sexual behavior • Autonomic responses • Biological rhythms Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Hippocampus • Categorizes information • Consolidated memories • Amygdala • Evaluates incoming information • Signals danger Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Learning • Acquiring information as a result of experience • Memory • Information is encoded, stored, and retrieved Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Implicit memory • Unconscious memory for perceptual and motor skills • Explicit memory • Factual memory of people, places, or objects Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Short-term memory • Recall information for a few minutes • Information can be transferred to long-term memory Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Synaptic plasticity • Ability of the nervous system to modify synapses during learning and remembering • Long-term memory storage • Gene activation • Long-term functional changes at synapses Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Long-term potentiation (LTP) • Increased sensitivity to an action potential by a postsynaptic neuron • Long-term depression (LTD) • Long-lasting decrease in the strength of synaptic connections Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Peripheral nervous system • Somatic division • Responds to changes in the external environment • Autonomic division • Regulates the internal activities of the body Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Sympathetic system • Permits the body to respond to stressful situations • Parasympathetic system • Influences organs to conserve and restore energy Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Sympathetic and parasympathetic nerves • Innervate many organs • Function in an opposite way –Sympathetic system increases heart rate –Parasympathetic system decreases heart rate Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation Dual innervation of the heart and stomach Copyright © 2005 Brooks/Cole — Thomson Learning Biology, Seventh Edition CHAPTER 40 Neural Regulation • Drug actions and effects • Alter mood • Increase or decrease the concentrations of specific neurotransmitters within the brain Copyright © 2005 Brooks/Cole — Thomson Learning