On the Function of Flowers 1. Review of reproductive biology -life cycle -variation in flowers 2. Pollination Ecology -precise placement -outcrossing -pollination syndromes On the Function of Flowers 1. Review of reproductive biology -life cycle -variation in flowers 2. Pollination Ecology -precise placement -outcrossing -pollination syndromes Three paths for pollen flow: selfing: 1. Within flower 2. Between flowers 3. outcrossing (between plants) Red Maple Flowers staminate pistillate Foxgloves and outcrossing Three components to outcrossing in foxglove 1. Young flowers are male, old are female 2. Young flowers are nearer the top 3. Bees fly up. Male zone Then start over at the bottom Bees fly up Female zone Primroses: Outcrossing with Heterostyly syndromes On the Function of Flowers 1. Review of reproductive biology -life cycle -variation in flowers 2. Pollination Ecology -precise placement -outcrossing -pollination syndromes -pollination syndromes Hummingbird --long tube -red color -abundant nectar - no odor - no landing -pollination syndromes Sphinx Moth --long tube -white color -abundant nectar - exquisite odor (periodic) - landing evening primrose (Oenothera, Onagraceae) Angraecum (Orchidaceae) Angraecum sesquipedale Xanthopan © Esko Puupponen -pollination syndromes Beetle -- two variants, nuptial chamber and painted bowl See Bernhardt 2000 to fix this slide and the next two. The beetle pollination syndrome nuptial-chamber variant • Many stamens often with appendages • Chamber to accommodate several animals • Night-flowering • Without markings • Scent attractant for beetles -pollination syndromes Carrion insect --contrasting light and dark pattern -brown-purple color -no nectar - foul odor wakerobin Carrioninsect flowers Jack in the pulpit Wild ginger Stapelia - African carrion-insect dispersed flower -pollination syndromes Wind--no perianth -no nectar -imperfect flowers -abundant pollen -big stigmas Grasses – family Poaceae (Gramineae) Sparganium - windpollinated aquatic -pollination syndromes Flowers that mimic insects -mate mimics -aggressive mimics Ophrys pollination On the Function of Flowers 1. Review of reproductive biology -life cycle -variation in flowers 2. Pollination Ecology -precise placement -outcrossing -pollination syndromes