Selection of Breeding Program An S 426 Fall 2007 Selection of Breeding Program Setting of goals is of major importance and includes goals for: 1. 2. 3. 4. Reproduction Calf performance Income Etc. Breeding management will impact each of above in varying degrees: 1. Breeding management practice with greatest impact on reproduction is crossbreeding 2. Selection is best management practice for improving carcass quality Selection of Breeding Program Once goals established, herd assessment is needed – where is herd at 1. Determine breed makeup 2. Determine production level – need accurate records 3. Determine average weight and frame size of cow herd Larger frame cows have higher maintenance costs Table 1. Frame relationship to mature size and carcass weight Frame Score 3 4 5 6 7 8 9 Yearling Hip Height (in) Expected Weight (lbs) Bulls Mature Cows 1025 1100 1175 1245 1320 1395 1465 45 47 49 51 53 55 57 Heifers 43 45 47 49 51 53 55 Steer Harvest 950 1050 1150 1250 1350 1450 1550 Steer Carcass 600 660 725 785 850 915 975 Figure 1. Comparison of growth curves of small and large framed cattle. Selection of Breeding Program Larger frame feedlot cattle 1. Grow at faster rate 2. Require greater amounts of feed and in feedyard longer, thus greater expenses 3. Heavier finish weights generate more income 4. Real problem if variation in frame sizes in same feedyard, may need to sort for: a. b. Selection of Breeding Program Genetic change 1. Genetic change may not result in improved animals 2. Genetic improvement means superior animals with greater profitability Brought about in two ways: 1. 2. Selection of Breeding Program Selection – factors affecting: 1. 2. 3. 4. Variation – Heritability – Selection differential – Generation interval – Predicting genetic change = Genetic change per year = Progress through selection dependent upon number of traits selected where n = number of traits selected Selection of Breeding Program Single-vs. multiple-trait selection Single-trait selection produces rapid genetic change, but unfortunately likely results in undesirable changes in correlated traits Increasing frame size resulted in increased: 1. 2. 3. 4. Selection of Breeding Program Multiple-trait selection first step towards more effective selection, but challenges remain: 1. Rate of improvement in any one trait decreases 2. Unfavorable genetic correlations remain for economic relative traits (ERT) Selection of Breeding Program Multiple-trait selection first step towards more effective selection, but challenges remain: 3. Methods a. Tandem – emphasis on one trait until desired level attained and then move to next trait b. Independent culling - choosing minimum or maximum levels for each trait - more traits added, more relaxation occurs in order to find animals meeting criteria - once thresholds identified, application of this method is easy and popular Selection of Breeding Program 3. Methods c. Selection indexes – incorporate the cost and income resulting from production 1.) A description of every index should include: a.) Traits included in the index b.) Description of information used in the index, such as EPD or individual phenotypic performance c.) source of economic information and performance levels used to calculate economic weights d.) Relative economic emphasis of each trait to the overall index Selection of Breeding Program c. Selection indexes – incorporate the cost and income resulting from production 2.) Successful application of generalized indexes relies on logical implementation: a.) Identify production and marketing system 1.) When will animals be marketed (at what age) 2.) How will animals be marketed 3.) What is current performance and genetic level of herd Selection of Breeding Program c. Selection indexes – incorporate the cost and income resulting from production 2.) Successful application of generalized indexes relies on logical implementation: b.) Identify the index appropriate to the production system outlined above - questions to be addressed 1.) What traits are included in the index 2.) What are the relative economic values used to weigh the traits 3.) Decide on the appropriate index for evaluation based on the most similarity between above points 1 and 2 Selection of Breeding Program c. Selection indexes – incorporate the cost and income resulting from production 3.) Most breed selection indexes are generalized for overall breed improvement and use average costs and income from production rather than accounting for a specific producer’s marketing and production systems a.) Has led to the development of Interactive Decision Support (IDS) systems which allow breeders to input parameters specific to their production systems – two classes: Selection of Breeding Program 1.) Herd – level systems a.) Designed to evaluate overall genetic changes in herd level – not individuals b.) Angus Assoc. Optimal Milk Model – designed for producers to decide the appropriate range of milk EPD given the mature weight of their cows, annual cow costs and variability in feed resources Selection of Breeding Program 2.) Individual animal systems a.) Designed to evaluate individual animal selection decisions and the impact of those decisions on profitability b.) Charolais Assoc. Terminal Sire Profitability Index (TS) – designed to evaluate the selection of sires based on their relative impact on profitability in a terminal sire mating system; allows input of current herd production characteristics and sources of income by the producer Selection of Breeding Program Mating systems 1. Inbreeding – a. Main genetic effect is to increase the number of pairs of homozygous genes b. Usually results in a decline in traits related to physical fitness 1.) 2.) Selection of Breeding Program Mating systems 2. Outbreeding – a. Main genetic effect is to increase the number of pairs of heterozygous genes b. By increasing heterozygosity there is a tendency to cover up detrimental recessive genes and improve traits related to physical fitness 1.) 2.) Selection of Breeding Program Mating systems Crossbreeding – two reasons for its use: 1. For certain traits may result in heterosis 2. Breed complementarity Selection of Breeding Program Table 1. Summary of heritability and level of heterosis by trait type. A Trait Carcass / end product Skeletal measurements Mature weight Growth rate Birth weight Weaning weight Yearling weight Milk production Maternal ability Reproduction Health Cow longevity Overall cow productivity a Adapted from Kress and MacNeil. 1999 Heritability Level of Heterosis Medium Medium (5 to 10 %) Low High (10 to 30 %) High Low (0 to 5 %) Selection of Breeding Program 1. For certain traits may result in heterosis average of average of a. % heterosis =crossbred - purebreds x 100 average of purebreds 1.) b. Traits not expressing heterosis will equal the average of the parent breeds for that trait in the crossbred offspring Selection of Breeding Program 1. For certain traits may result in heterosis c. Why important in beef cattle business 1.) Largest benefit to commercial production is crossbred cows – improves: a.) b.) c.) d.) d. Keys for successful crossbreeding program: 1.) 2.) e. Crossbreeding systems Selection of Breeding Program Breed complementarity – implies using breeds in a crossbreeding program where their strengths and weaknesses complement one another 2. a. Example 1.) Selection of Breeding Program Composite breeds – 1. Maintain heterosis 2. Utilize breed complementarity 3. Once formed, can be managed like a purebred herd Formed from 2-breed 3-breed 4-breed Retained % maximum heterosis 50 67 75 Improved produc. of cow herd(%) 12 15 17 Table 1. Breed performance levels for seven traits in beef cattle. a,b. Table 2. 2004 Adjustment factors to add to EPD of 15 different breeds to estimate across-breed EPD. a,b. Breed Angus Beefmaster Brahman Brangus Braunvieh Charolais Gelbvieh Hereford Limousin Maine Anjou Pinzgauer Red Angus South Devon Salers Shorthorn Simmental Tarentaise Birth Weight 0.0 9.7 13.0 5.7 6.5 10.5 5.4 3.4 4.5 6.7 7.7 3.6 6.7 4.9 7.8 6.4 3.6 Weaning Weight 0.0 39.0 34.8 20.0 30.0 38.4 7.1 -2.0 1.8 17.6 28.3 -1.4 21.7 30.7 31.4 22.4 30.1 Yearling Weight 0.0 37.9 -4.4 20.4 13.9 53.4 -21.1 -13.7 -19.9 5.5 25.5 0.7 40.8 46.1 44.5 21.9 13.4 Milk . 0.0 --24.6 --22.2 2.6 1.7 -17.8 -15.9 7.6 6.1 -7.8 3.5 9.0 12.1 10.0 17.8 Van Vleck and Cundiff, 2004 b Updates to across-breed EPD adjustments can be found at www.beefimpovement.org a Table 3. Average across-breed EPD for nonparent animals born in 2002-2003 by breed. a. Breed Angus Beefmaster Brahman Brangus Braunvieh Charolais Gelbvieh Hereford Limousin Maine Anjou Pinzgauer Red Angus Salers Shorthorn Simmental South Devon a Birth Weight 2.6 10.1 15.1 7.7 7.6 12.0 6.4 7.2 6.9 9.2 7.6 4.1 7.8 6.7 9.7 6.4 Weaning Weight 35 45 51 41 37 57 44 33 36 34 29 27 34 44 65 40 Yearling Weight 65 49 22 54 21 85 48 46 44 37 26 49 60 66 102 45 Adjustment using the 2004 across-breed EPD adjustments listed in Table 2. Milk 17 --32 --22 8 19 -5 2 11 5 6 12 11 18 16 . . Selection of Breeding Program Successful breeding program depends on: 1. 2. a. b.