Plant Anatomy AP Biology 2006-2007 Basic plant anatomy root root tip root hairs shoot (stem) nodes internodes buds terminal or apical buds axillary buds flower buds & flowers leaves AP Biology mesophyll tissue veins (vascular bundles) Meristem Regions of growth stem cells: perpetually embryonic tissue regenerate new cells apical shoot meristem growth in length primary growth apical root meristem growth in length primary growth lateral meristem growth in girth secondary growth AP Biology Apical meristems AP Biology shoot root Leaves Function of leaves photosynthesis energy production CHO production gas exchange transpiration AP Biology Modified leaves tendrils (peas) AP Biology succulent leaves spines (cacti) colored leaves (poinsetta) AP Biology Interdependent systems Both systems depend on the other roots depend on sugars produced by photosynthetic leaves shoots depend on water & minerals absorbed from the soil by roots AP Biology sugars water & minerals Plant TISSUES Dermal epidermis (“skin” of plant) single layer of tightly packed cells that covers & protects plant Ground bulk of plant tissue photosynthetic mesophyll, storage Vascular AP Biology transport system in shoots & roots xylem & phloem vessel elements Xylem vessel element Vascular tissue move water & minerals up from roots dead cells at functional maturity only cell walls remain need empty pipes to efficiently move H2O transpirational pull dead cells Aaaah… Structure–Function again! tracheids AP Biology Phloem: food-conducting cells carry sugars & nutrients throughout plant sieve tube companion cell sieve plate plasmodesmata AP Biology living cells Putting it all together Obtaining raw materials sunlight leaves = solar collectors CO2 stomates = gas exchange H2O uptake from roots nutrients uptake from roots AP Biology Transport in Plants AP Biology 2006-2007 Transport in plants H2O & minerals transport in xylem transpiration evaporation, adhesion & cohesion negative pressure Sugars transport in phloem bulk flow Calvin cycle in leaves loads sucrose into phloem positive pressure Gas exchange photosynthesis CO2 in; O2 out stomates AP Biology respiration O2 in; CO2 out roots exchange gases within air spaces in soil Why does over-watering kill a plant? Ascent of xylem fluid Transpiration pull generated by leaf AP Biology Water & mineral absorption Water absorption from soil osmosis aquaporins Mineral absorption active transport proton pumps active transport of H+ aquaporin root hair AP Biology proton pumps H2O Mineral absorption Proton pumps active transport of H+ ions out of cell chemiosmosis H+ gradient creates membrane potential difference in charge drives cation uptake creates gradient cotransport of other solutes against their gradient AP Biology Transport of sugars in phloem Loading of sucrose into phloem flow through cells via plasmodesmata proton pumps cotransport of sucrose into cells down proton gradient AP Biology Pressure flow in phloem Mass flow hypothesis “source to sink” flow direction of transport in phloem is dependent on plant’s needs phloem loading active transport of sucrose into phloem increased sucrose concentration decreases H2O potential water flows in from xylem cells increase in pressure due to increase in H2O causes flow AP Biology On a plant… What’s a source…What’s a sink? can flow 1m/hr Control of Stomates Guard cell Epidermal cell Uptake of K+ ions by guard cells proton pumps water enters by osmosis guard cells become turgid H2O K+ H2O K+ Loss of K+ ions Nucleus Chloroplasts H2O K+ H2O K+ K+ H2O K+ H2O K+ H2O K+ H2O Thickened inner cell wall (rigid) by guard cells AP Biology water leaves by osmosis H2O K+ guard cells become flaccid H2O K+ H2O K+ H2O K+ Stoma open Stoma closed water moves into guard cells water moves out of guard cells Plant Reproduction In a nutshell AP Biology 2007-2008 Plant Diversity mosses ferns conifers flowering plants Bryophytes Pteridophytes Gymnosperm Angiosperm non-vascular land plants seedless vascular plants pollen & “naked” seeds flowers & fruit flowers & fruits pollen & seeds vascular system AP Biology common ancestor xylem cells tracheids Animal vs. Plant life cycle Animal Plant diploid multicellular individual 2n mitosis meiosis fertilization haploid unicellular gametes 1n AP Biology diploid multicellular sporophyte 2n mitosis fertilization meiosis gametes 1n spores 1n mitosis mitosis haploid multicellular gametophyte 1n alternation of generations Bryophytes Mosses & liverworts non-vascular diploid swimming sperm multicellular dominant haploid gametophyte sporophyte dependent sporophyte 2n mitosis spores for reproduction meiosis fertilization gametes 1n spores 1n mitosis mitosis haploid multicellular gametophyte 1n AP Biology alternation of generations Pteridophytes Ferns vascular swimming sperm dominant sporophyte independent gametophyte fragile diploid multicellular sporophyte 2n mitosis fertilization meiosis gametes 1n spores 1n mitosis mitosis spores for reproduction haploid archegonia antheridia AP Biology alternation of generations haploid multicellular gametophyte 1n Gymnosperm Conifers vascular diploid pollen (sperm) multicellular dominant sporophyte sporophyte dependent reduced gametophyte 2n mitosis cones, seeds for reproduction fertilization meiosis female male AP Biology gametes 1n spores 1n mitosis mitosis haploid multicellular gametophyte 1n Angiosperm Flowering plants vascular diploid pollen (sperm) multicellular dominant sporophyte sporophyte dependent reduced gametophyte 2n mitosis flowers, fruits, seeds for meiosis fertilization reproduction male gametophyte in pollen (haploid) female gametophyte in ovary (haploid) AP Biology gametes 1n spores 1n mitosis mitosis haploid multicellular gametophyte 1n Angiosperm: flowering plants AP Biology polar nuclei Angiosperm life cycle male gametophyte in pollen (haploid) pollen grains egg cell fertilization female gametophyte in ovary (haploid) AP Biology sporophyte in seed (diploid) Seed dispersal Plants produce enormous numbers of seeds to compensate for low survival rate vast amount of genetic variation for natural selection to screen Seed Aviation r-strategy AP Biology K-strategy Plant Responses to Stimuli AP Biology Plant hormones auxin gibberellins abscisic acid ethylene and more… AP Biology Auxin (IAA) Effects controls cell division & differentiation phototropism growth towards light asymmetrical distribution of auxin cells on darker side elongate faster than cells on brighter side AP Biology apical dominance Gibberellins Family of hormones over 100 different gibberellins identified Effects stem elongation fruit growth seed germination AP Biology plump grapes in grocery stores have been treated with gibberellin hormones while on the vine Abscisic acid (ABA) Effects slows growth seed dormancy high concentrations of abscisic acid germination only after ABA is inactivated or leeched out survival value: seed will germinate only under optimal conditions light, temperature, moisture AP Biology Ethylene Hormone gas released by plant cells Effects fruit ripening leaf drop like in Autumn apoptosis One bad apple spoils the whole bunch… AP Biology Fruit ripening Adaptation hard, tart fruit protects developing seed from herbivores ripe, sweet, soft fruit attracts animals to disperse seed Mechanism triggers ripening process breakdown of cell wall softening conversion of starch to sugar sweetening positive feedback system ethylene triggers ripening ripening stimulates more ethylene production AP Biology Other Responses Gravitropism AP Biology Occurs as soon as seed germinates, ensures root grows into soil and shoot grows toward sunlight. (Auxin) Other Responses Thigmotropism AP Biology Response to touch, allows the plant to use mechanical supports as it grows, also can result in rapid leaf movements (similar to action potentials in nerves) Other Responses Photoperiodism A physical response, such as flowering to the photoperiod Actually controlled by night length not day length Long-day plants flower in late spring or early summer Short-day plants flower in late summer fall or winter AP Biology Other Responses Photoperiodism AP Biology