EFFECTS OF CLIMATIC VARIABILITY AND CHANGE ON FOREST ECOSYSTEMS – EMPIRICAL EVIDENCE FROM THE WESTERN MOUNTAIN INITIATIVE NETWORK PETERSON, DAVE (1); FAGRE, DAN (2); MCKENZIE, DON (1); ALLEN, CRAIG (3); BARON, JILL (4); STEPHENSON, NATE (5) (1) USDA Forest Service, PNW Research Station, Seattle, WA, (2) USGS Northern Rocky Mountain Science Center, West Glacier, MT, (3) USGS Fort Collins Science Center, Los Alamos, NM, (4) USGS Fort Collins Science Center, Fort Collins, CO, (5) USGS California Science Center, Three Rivers, CA The effects of variability and directional trends in climate during the past century are measurable in forest ecosystems at several locations in the Western Mountain Initiative network. In the American Southwest, prolonged drought has caused dieback and reduced productivity in large areas, with the potential for severe fires due to fuel accumulations. In the Sierra Nevada, drought has also affected forests, with reduced tree vigor and increased mortality. Mixed conifer forests in this region are also stressed by elevated levels of oxidant air pollution. In the Pacific Northwest, subalpine forests have experienced rapid in-fill of subalpine meadows throughout the region. In the Northern Rocky Mountains, subalpine forests at altitudinal treeline have also filled in and transitioned from krummholz to a more upright growth form at some locations. In the central Rocky Mountains, no significant changes are apparent, although high-elevation forests are subject to elevated levels of nitrogen deposition. There appears to be a significant trend of increased tree growth throughout the West, focused primarily on high-elevation forest and secondarily on Pacific coastal forest – perhaps a signal of carbon dioxide fertilization. Potential signals of the effects of climatic change must be differentiated from effects of the Pacific Decadal Oscillation, which is an important multi-decadal influence on biophysical variability in Western mountain ecosystems.