The distribution of the cutthroat trout (Salmo clarki) in Montana by Delano A Hanzel A THESIS Submitted to the Graduate Faculty in partial fulfillment of the requirements for the degree of Master of Science in Fish and Wildlife Management Montana State University © Copyright by Delano A Hanzel (1959) Abstract: The distribution of cutthroat trout (Salmo clarki) and the factors affecting it were investigated during the summers of 1957 and 1958. Distribution records were obtained from the following sources: 100 streams surveyed, east of the Continental Divide; 219 records from fisheries biologists and 769 from creel census returns (Montana Fish and Game Department); 35 records from the Montana State College collection. Fifty-five (75 percent) of the streams surveyed had only cutthroat trout above barriers. The important barriers were natural falls, high gradient areas, and beaver dams. "Wherever rainbow and/or-eastern brook trout were present in association with cutthroat trout they were predominant. Cutthroat trout are presently restricted to the headwaters of streams which originally were entirely inhabited by them. Taxonomic determinations, were based upon the examination of 345 cutthroat trout (126 from streams that had never been stocked with rainbow trout), 54 rainbow; trout and 88 rainbow X cutthroat trout. Satisfactory separation for fish over 4.0 inches , in total length was achieved. Individual distribution records of cutthroat trout from 699 streams and 244 lakes were listed. They were predominant (only game fish present or ranked first in relation to other game fish) in 253 (38 percent) streams and l42 (58 percent) lakes. HHE DISTRIBUTION OF THE CUTTHROAT TROUT (SAEiMO CLARKI) IN MONTANA DELANO A. HANZEL Submitted to the Graduate Faculty partial fulfillment of the requirements for the degree of Master of Science in Fish and Wildlife Management Montana State College Approved8 Head, Maj oW Department’ Chai^An, Examining Committee Dean, Graduate/DiVision Bozeman,.Montana 1» THE AUTHOR , Delano A, Hanzel was born on March 2 0 1935 in Belt, Montana and graduated from Belt Valley High School in 1953» He entered Montana State College in 1953 and received a Bachelor of Science degree in Fish and Wildlife Management in June, 1957» During the summer months of 1952 - 1956, he was employed by the Montana Fish and Game Department as a student assistant» He was married to the former Betty L» Hill in 1958„ He began graduate studies at Montana State College in September 1957« This thesis fulfills part of the requirements for his Master of Science degree in Fish and Wildlife Management at Montana State College, Bozeman, Montana. 13S433 "*2*" TABLE OF CONTENTS Page ABSTRACT o o p p p e n i i p p p 9 9 P 3 9 h INTRODUCTION ................. DESCRIPTION OF THE STEJDI AREA 9 FIELD SURVEY METHODS . . . . . . FIELD SURVEY RESULTS . II TAIONOMIC CONSIDERATIONS . . . Ih CUTTHROAT TROUT DISTRIBUTION . AreapN o 8 I (Northwst) records . 6 9 6 ' B, 27 Area No« 2 (West central) records 37 Area No* 3 (Central) r e c o r d s ........ ' . Area No, It (Southwest) records Area No, ^ (South) records SUMMARY p Literaojhe o e o cited f l o . o o o o Ii8 ............... . . . . . O B l|,li O e o . . . . . ............................................... .. . . . . . « • . . ^3 . 57 ABSTRACT The distribution'of cutthroat:trout (Salmo clarkl) and the factors affecting it were investigated during the summers of 1957 and 1958, Dis­ tribution records were obtained f rom.t h e .following sources;, .100 .streams surveyed, east, of',the Continental Divide $ ,219 records from fisheries .biologists and 769 from creel census returns (Montana Fish, and Game Department); 3.5 records from, the Montana State College collection. Fiftyfive (?5 percent) of the streams surveyed had only cutthroat trout above barriers. The important barriers were natural falls, high gradient areas, and beaver dams. "Wherever rainbow and/or-eastern brook trout w ere■present in association with cutthroat trout they were predominant. Cutthroat trout are presently restricted to the headwaters of streams which origin­ ally were entirely inhabited by them. Taxonomic determinations, were based upon the examination of 3^5 cutthroat trout (126 from streams that had never been stocked with rainbow trout), rainbow; trout and 88 rain­ bow X cutthroat trout. Satisfactory separation for fish over I4.O inches , in total length was achieved. Individual distribution records of cut­ throat trout from 699 streams and 2hh lakes were listed'. They were pre­ dominant (only game fish present or ranked first in relation to other game fish) in 253 (38 percent) streams and llt.2 (58 percent) lakes. —-I].— INTRODUCTION The cutthroat trout (Salmo clarki) originally inhabited all the waters of Montana in and adjacent to the mountains except for a consider­ able number of small isolated virgin lakes. Other native game species which shared this range were the grayling (Thymallus arcticus) and the mountain whitefish (Prosopium williamsonj) on the eastern slopes of the Continental Divide with the dolly varden (Salvelinus alpInus) and the mountain whitefish on the western slopes. present (Table l ) . Other native:fishes were also ' Jordan (1889) reported that cutthroat trout were abundant in the upper Yellowstone River drainage. Evermann and Cox (I89I4) stated that the cutthroat trout attracted a large number of anglers in the upper waters of the Missouri River Basin and although the supply was large it had begun to diminish. This decline was attributed to fishing pressure and increased water and land use. Evermann (1893)" reported an abundance of cutthroat trout on the western slopes of the Rocky Mountains. There is evidence to show that cutthroat trout were abundant in the mountainous areas of Mon­ tana, however, this species was probably no more abundant than grayling and mountain whitefish in. many streams. While exotic trout have been introduced Into all the major drainages originally occupied b y cutthroat trout, a few small tributaries still re­ main unmolested. Rainbow trout (Salmo gairdneri) was first introduced in 1891 and has been most extensively stocked since' that time. Brown trout • (Salmo trutta) was originally introduced in 1891 and has become the pre- Table 1» list of the fishes associated with cutthroat trout (Salmo clarki) in Montana. Distri- . bution IJ Species Game Grayling Kokanee salmon Brown trout Rainbow1trout Eastern brook trout Dolly varden trout Rygmy whitefish Mountain whitefish Others Common white sucker Eastern longnose sucker Columbia largescaled sucker Mountain sucker Carp Longnose dace Columbia River chub Squawfish Redside shiner Black bullhead Burbot Pumpkinseed Yellow perch Northern sculpin Slimy sculpin Torrent sculpin Thymallus arcticus Onchorhynchus nerka Salmo trutta Salmo. gairdneri Salveiinus fontinalis .Salvelinus alpinus Prosopium coulter! Prosopium williamsoni W EM Catostomus eommersoni E Catostomus catostomus ■ E Origin E/ N I : I .I I N ■N N E EM EM EM ■ Catostomus macrocheilus . ¥ E Pantosteus platyrhynchus Cyprinus carpio EM EM Rhinichthys oataractae Mylocheilus caurinus ¥ ¥. Ptychocheilus oregbnense ¥ Gila balteata E. Ictalurus melas EM Lota lota ¥ Lepomis gibbosus Perea flavescens EM 1 W Cqttus bairdi ¥ . Cottus cognatus. ¥ Cottus rhotheus N ■ N N I N N N N I N I I N N N i/ E - east Continental Dividej W - west Continental Divide| EM - both sides of Divide. S/ N - native 5 I - introduced. r 5/ St. Mary’s Drainage, east Continental Divide. , V" . ’ dominant species in the Talley streams of the cutthroat trout range. Eastern brook trout (Salvelinus fontinalis) was introduced in l 8pb and now occupies many of the small valley brooks and mountain headwater.creeks as well as a considerable number of mountain lakes. These exotic species . have gradually replaced the cutthroat trout in the lower parts of its original range. The native strains of cutthroat trout are now limited to a few remote areas of the State. Hybrids between rainbow and cutthroat trout have appeared in practi­ cally all drainages where rainbow trout were introduced. These hybrids are numerous in most places which makes identification of the cutthroat trout and the determination of its present range extremely difficult. The effects of hybridization on the future of the cutthroat trout are not known. This study of cutthroat trout has two primary objectivess to de­ termine the distribution and abundance of pure cutthroat trout stocks? and to secure information on influencing factors. In addition, observations were made on taxonomic differences between the various native strains of cutthroat trout and on the prevalence of hybrids. This study may prove useful in future management of these species. Time did not permit the writer to determine the complete range of the cutthroat trout for the whole State. east of the Continental Divide. Investigations were concentrated However, all the available information on this species in Montana has been reviewed and included. Field collections were made and surveys conducted during the summers of 1957 and 1958 (June to September). ' • '■ ' ' ■ ™ - I - ■ . . • ■ ' ' The writer extends thanks to the following persons and organizations for their assistance during the study. Dr. G . J. D. Brown gave technical supervision and aided in the preparation of this manuscript. Nels A. Thoreson suggested,the problem,and rendered valuable field assistance 5 other Montana Fish and Game Department personnel aided in collecting Edward Nevala 5 Quenton Stober specimens and furnished distribution data. and James Galkins assisted in the stream surveys. Service supplied maps. The U. S. Forest The Montana Fish and Game Department financed the field work under Federal Aid to Fisheries Restoration Project F-^-R. DESCRIPTION OF THE STUDY AREA The present distribution of the cutthroat trouta east of the Conti­ nental Divide in Montana 5 is confined to parts of most major primary tributary drainages; in the Missouri River from Three Forks to the mouth of the Musselshell River 5 and in the Yellowstone River from the Wyoming boundary to the mouth of the Big Horn River. This species is rarely found in the main stem of the Missouri River 5 however, it does occur frequently in the Yellowstone River for a distance of about 90 miles down stream from Yellowstone National Park. The' major primary streams of these two large rivers have vast net- • works of secondary and tertiary tributaries draining the east slope of the Rocky Mountains in Montana. Remnants, of pure cutthroat trout are mostly confined to the small headwater streams. These drains steep mountain slopes, which are generally covered by coniferous forests; mountain valleys where grasses, sedges and willows ,predominate; valleys at low -8elevations characterized "by sagebrush and bunchgrass. "These streams are £ - 20 feet in width (av, approx, depths usually less than two feet, 9 feet) and have They originate at elevations from 6 5OOO to 8,000 feet above' sea level. The lowest elevation at .which cut­ throat trout were collected in streams was ij.<,S>00 feet, however, a few specimens were taken in ponds and reservoirs at lower elevations. Esti­ mated gradients of streams presently occupied b y cutthroat trout were usually from ^O to 25>0 feet per mile, but there were extensive stretches of cascades and falls where gradients were "higher-. Summer stream ve­ locities of I - 3 feet per second were characteristic of riffle areas. Ve­ locities taken during early spring run-off in the more precipitous areas were approximately double those of summer. Beaver dams occur frequently on the streams and have a ’tempering effect on the velocities. In general, bottom materials (based on visual estimates) were composed of about 10 per­ cent boulders, 13 percent rubble, and detritus. 60 percent gravel and 15 percent sand Exceptions to the general composition were in areas of beaver activity and mining dumps where silt became a major component. Summer water temperatures (June 20 to Sept, 2It) varied from ItS to 6S° E. and the total alkalinity (methyl orange) range was 13,S - 22? ppm. The principal stream bottom organisms were stoneflies (Plecoptera), caddisflies (Trichoptera) and mayflies (Ephemeroptera). Algae were common but vascular water plants were rare. Mining, logging, and livestock are the major industries found in the area of cutthroat trout distribution. U. S. Forest Service and other access roads are present in some forest areas, however, about 75 percent of the cutthroat trout streams are still inaccessible by road „ FIELD SURVEY METHODS ■■ ' The lack of roads along mountain streams made the use of an. electric fish shocker impractical» Most collections were made by angling or by using eresol, however, other fish toxicants and dynamite were employed to a limited extent, An attempt was made to test the effectiveness of sampling by angling„ Six miles of a stream were selected which had an approximate average width of nine feet, a depth of eight inches and a velocity of two feet per secondo The stream was then divided into six 1-mile,sections= A 300 foot portion of each mile section, selected in favorable -trout habitat,• was shocked ( H O volt A.C.), The fish recovered were counted and returned to the area in which they were taken« Each one mile section was then fished using flies (wet and dry), mately two hours of fishing. A distance of one mile was covered in approxi­ While the number of fish taken b y angling was considerably less than by shocking, angling appeared adequate to show the'range and relative abundance of trout (Table 2). In actual practice, the length of streams fished ranged, from 3 ’^ 8' miles a Usually two fishermen sampled alternate parts of a stream from lower to higher elevations. Success was generally good, possibly because of low fishing pressures in these areas.' Angling was considered sufficently .successful to determine the range and relative abundance of trout in 80 of the 100 streams surveyed. Relative .abundance estimates were probably more accurate on .small streams where fishing was more intense. , -10Table 2« Sections Angling and shocking' success on test stream. . Shocking. C p e r J O O t) .Angling (per mile) No. Fish No. Fish Species of trout I Cutthroat trout U8 22 2 Cutthroat trout 36 Ih 3 I/ Eastern brook trout Cutthroat trout Rainbow trout ■ 98 # •I 2k 6 0 sa 17 . ,h 2 0 U 5 6 Eastern brook trout Cutthroat trout , „ Rainbow trout . Rainbow trout . •Eastern brook trout Eastern brook trout Rainbow trout Cutthroat trout 2 26 ; : ■ 11 8 W 2/ 18 .6 8 I 5 .0 3/ Bainbow x cutthroat hybrids were presents but at this early date of the investigation no definite identification was made. &/ Twenty-nine of these were less than 3 inches in length. Oresol was used where angling success was low. An estimate of the stream volume was made by using .a^-veldci-ty-vhead. irodv ’ Gresol was applied- at the rate of one gallon per four cfs for each 100 yards of the stream (Wilkins, 1955), Cresol was spread over the upper half of the sample area when velocities were less than one foot per second. When velocities were greater, it was applied in a narrow band across the stream, usually “XXat the head of a pooX„ Incapacitation of trout and seuXpins in the faster streams was aXmost immediate after appiication and the effect was onXy momentary. from I? - In the sXower streams the incapacitation time varied 8 minutes and recovery from £ - 20 minutes» AXX sizes of fishes were affected b y the treatment, A smaXX amount of mortaiity occurred as a resuXt of fish thrashing about and becoming beached, FIELD SURVEY RESULTS FieXd surveys were made on XOO streams east of the GontinentaX Dividej, 73 of which contained cutthroat trout. Fifty-five (?5 percent) of these had onXy popuiations of cutthroat trout above barriers 5 nine had exotic trout pXanted above fish barriersj two had cutthroat trout planted into existing exotic trout populations; five had cutthroat populations which were seriously effected by pollution or dewatering; two had popu­ lations of cutthroat and exotic trout with no barriers separating them. Forty-six (Si+ percent) of the barriers which had only cutthroat trout above them were either natural falls, high gradient areas or beaver dams. Natural falls (Fig. l) varied in height from I* - 30 feet and no exotic trout were found above them in most instances. areas (gradient High gradient !?00 - 1,^00 feet per mile) varied in length from 330 - 1,320 feet (Fig, 2). The bottom materials in these areas were pre­ dominantly large boulders and rubble with numerous dead falls and other debris, Beaver dams formed barriers either singly or in series. dams were usually old and ranged from 6 - 1 2 feet in height. Single Even low beaver dams were barriers if a large enough number occurred in a series -12- Fig. I. Natural falls fish barrier (Hellroaring Creek, Beaverhead River Drainage)» Fig. 2. High gradient area fish barrier (David Creek, Big Hole River Drainage) . -13- (Fig. 3)• The stream in such places was often diverted into numerous channels which covered the entire flood plain. In a few instances beaver dams occurred in conjunction with irrigation diversion dams to form fish barriers. Fig. 3. Low beaver dams form fish barrier (Deep Creek, Missouri River Drainage). Exotic trout when planted above barriers were invariably predominant„ An example of this was found in Tenderfoot Creek (tributary of Smith River) which had a pure cutthroat population above a 30 foot natural falls prior to stocking with rainbow trout in 1955• was made in the immediate area above the falls. This rainbow trout plant In 1958 a preponderance of rainbow trout occurred for three miles above the falls, followed by a —ll].— two-mile section where cutthroat# rainbow, and hybrid trout (rainbow X cutthroat},were present in approximately equal numbers» trout were found above this section,. Only cutthroat "Wherever cutthroat and rainbow trout were found together in a stream, hybrids were present. Another example was found in Highwood Creek (tributary of Missouri River) which had eastern brook trout planted in 1938 above a series of beaver dams and a natural falls. Mhen surveyed eastern brook trout were'predominant and cutthroat trout were rare in the entire stream. The West Gallatin River did not have a barrier separating cutthroat from exotic trout. Below the mouth of Speciman Creek, brown, rainbow, cutthroat and hybrid (rainbow X cutthroat) were found. Approximately one mile above Specimen Creek hybrids and cutthroat trout were present. The main river above this area as well as one tributary in Montana and three in Wyoming had cutthroat "treat only. Cutthroat trout are presently restricted to the headwaters of streams which originally were entirely inhabited by them. The major factors limiting cutthroat trout distribution are; stream habitat changes, compe­ tition with exotic species and hybridization with rainbow trout. Practi­ cally all pure cutthroat trout population presently existing in streams, are above barriers. TAXONOMIC CONSIDERATIONS The widespread introduction of rainbow trout into cutthroat trout waters, as well as the indiscriminate stocking of various cutthroat trout strains, along with the fact that these species readily hybridize, has -15>created serious taxonomic difficulties. Taxonomic considerations are based upon the;examination of 3h$ cut­ throat trout (126 from streams that had never been stocked with rainbow trout)s rainbow trout and 88 rainbow I cutthroat trout. The taxonomic characteristics used (Table 3j Figs. It - 7 ) 3 with minor exceptions, are those described by Miller, 1950. No single characteristic was found to be adequate for identification but when used in combination satisfactory separation of fish over it.O inches total length was achieved. Lateral' line scale counts are often used to separate cutthroat and rainbow trout, but in the specimens studied there was much overlapping and this characteristic was not used. CUTTHROAT TROUT DISTRIBUTION The distribution of cutthroat trout .(Figs. 8 - 13) was determined from: I - field surveys made by the writer; 2 - collections at Montana State College; 3 - records of fisheries biologistffS and creel census, Montana Fish and1Game Department. The,distribution, of cutthroat trout is listed along with that of other game fishes and the presence of barriers (Tables It - 8). The streams and lakes are arranged by drainages beginning at the mouth and going up stream. In the tabular data, the major tributary streams of the Missouri, Yellowstone, Flathead, Kootenai, and Clarks Fork of the Columbia Rivers are underlined. The primary, secondary, and ter­ tiary tributaries of these drainages are indented to show their relation. ship to one another. " Streams in parentheses have no records of cutthroat Table 3. Characteristics of cutthroat, rainbow and rainbow S cutthroat trout. Diagnostic Characteristics Cutthroat trout Rainbow X Cutthroat trout Rainbow trout Bentary mark Always present 5 orange to blood-red; weaker on juveniles Usually present;"often lighter than on cut­ throat trout Usually absent; rarely indistinct yellow Hyoid teeth Usually present Present or absent Absent Dark Usually milky-white Conspicuously milkywhite Range 1.3 2.3 "(usually 1.6 - 1 .9)I maxillary extends beyond eye Range 1.6 - 2.1 (usually 1.8 - 2 .0) Range 1.8 - 2,1 (usually 2.0 - 2 .l); maxillary not extend­ ing beyond eye Scales usually more exposed; visible without magnification Scales exposed; visible without magnification Spot distribution Usually concentrated above lateral line and on caudal peduncle Usually concentrated along lateral line Usually spotted over entire body ■ Spot size and shape Usually large; margins regular Usually large; margins irregular and contigu­ ous Usually small; margins irregular Shape of head JLong5 pointed and conical Similar, to either cutthroat or rainbow - Short=, blunt and rounded Shape of body Usually slender and compressed' Similar to either cutthroat or rainbow Usually deep and robust . v- Ventral border of anal fin Maxillary length in head length Scales deeply embedded; Scale distinction■ hardly visible with­ out magnification -17- hO hO faO hC . L. . . 6. . 7. Cutthroat trout from Missouri River Drainage. Cutthroat trout from Yellowstone River Drainage. Rainbow trout from Missouri River Drainage. Rainbow X cutthroat trout from Missouri River Drainage. “18trout and are listed only to show the relationship of other streams or lakes which do have cutthroat trout„ A series of symbols devised for expressing tabular information follows under appropriate headings s Cutthroat relation to other game fish. -The categories below are estimates of abundance in relation to other game . f i s h Ho information was secured on the actual abundance of cutthroat trout in the streams and lakes considered* e.g,« .The actual number of cutthroat trout in a d-stream might well be greater than that of an a-3 b - 5 or c-stream* abcde- cutthroat trout only game fish present3 or when used with' barrier, cutthroat trout only game fish above cutthroat trout more abundant than any other game fish cutthroat trout second in abundance to any other game fish cutthroat trout presents but third or less in abundance to any other game fish cutthroat trout collection record only Other game fish. REBLDGKW- Symbols used for other game fishes. rainbow trout eastern brook trout brown trout lake trout dolly varden grayling kokanee salmon mountain whitefish Barriers. Symbols used for .fish barriers. BD- beaver dam ' ID- irrigation diversion dam HF- natural falls or high gradient areas Information source. of the latest record. CSDM- $3>, Symbols used for sources of information and year creel, census' records ■ . field survey records biologist records of Montana Fisheries, Division collection records, Montana State College, etc.,- year of last collection. Area No.* I (Figs = 8 ^ 9) ° This area is in the extreme northwestern I H O I Area Boundary State Boundary — — C'n n t i -.1 r\.... Figo 8. Cutthroat trout range and area designations in Montana= — part of Montana, 20 ~ Qn the eastern slopes of the Continental Divide it in­ cludes the headwaters of the St, Mary's, Milk, Marias and Sun Rivers and on the.western slopes it includes the primary drainages of the Flathead, and Kootenai Rivers and the Clarks Fork of the Columbia River below the mouth of the Flathead River.' ; Cutthroat trout were recorded from 218 streams and 93 lakes but were predominant (only game fish present or ranked first irl-.relation to any other game fish) in only 112 streams and 62. lakes. Cutthroat trout records from the lower Milk River (Bear Paw Mountains) drainage were in­ cluded in this area. This trout was not native here but was planted in 1879 (Lucke5 1958) b y soldiers of a nearby army fort who secured fish from the Sun River5 west of Great Falls, Area No, 2 (Figs. S 5 10). This area is in west central Montana, entirely west of the Continental Divide, The primary drainages are the Blackfoot and Bitterroot Rivers and that portion of the Clarks Fork above the mouth of the Flathead River. Cutthroat trout were recorded from predominant in only ... 19h streams and I4.8 lakes but were 76 streams and 27 lakes. Area No, 2 (Figs, S 5 l l ) . the Continental Divide, This area is in central Montana5 east of . It includes the Missouri River and its tribu­ taries from Three Forks to the mouth of the Dearborn River and the head­ water streams of certain primary tributaries of the Missouri River from the mouth of the Dearborn River to the mouth of the Musselshell River. Cutthroat trout were recorded from 9h syreams and 21 lakes but were ST MARYS — — Area Bound ary N ^ State Boundary Continental Divide Records — streams ® lakes 9 5 10 miles Fig. 9* Cutthroat trout distribution in Area No. I (northwest area) I ro ro ^ Area Boundary — — State Boundary ------ Continental Divide Records — streams ® lakes tL U P miles Fig. 10. Cutthroat trout distribution in Area No. 2 (x^est central area) N Area Boundary ------ Continental Divide Records — streams © lakes I ro V Fig. 11. Cutthroat trout distribution in Area No. 3 (central area). predominant in only 23 streams and 13 lakes» Area No, L (Figs. 8 ? 12). This area is in southwestern Montana, east of the Continental Divide, and includes the drainage of the Missouri River above Three Forks, The Jefferson, Madison and Gallatin Rivers are the primary tributaries, Cutthroat trout were recorded from 100 streams and predominant in only 19 kl lakes but were streams and 28 lakes. Area No, £ (Figs, 8, 13), This area is in southern Montana, east of the Continental Divide and includes the Yellowstone River drainage from the State boundary to Billings, also included in this area. The headwaters of the Bighorn River are The primary tributaries of the Yellowstone River are the Shields, Boulder, Stillwater and Clarks Fork Rivers. Cutthroat trout were recorded from 63 streams and 25> lakes but were predominant in only 23 streams and 12 lakes. Several farm ponds on the lowep Yellowstone River (not included in this area) have had cutthroat trout planted in them. These records were not included. Including all of Montana, cutthroat trout were recorded from 669 streams and 2Itit lakes but w ere,predominant in only 2^3 (38 percent) streams and lit2 (£8 percent) lakes, Cutthroat trout records, west of the Continental Divide, include 378 streams and 133 lakes with cutthroat pre­ dominating in.102 streams and 83 lakes. Records east of the Continental Divide include 291 streams and 111 lakes with cutthroat predominating in 71 streams and £9 lakes. Fig. 12. Cutthroat trout distribution in Area No. It (southwestern area). m ile s ♦ AreaBoundary ------ State Boundary ------ Continental Divide R ecords. — s tre a m s (S) la k e s I ro CN I YELLOWSTONE NATIONAL PARK F i s . 13 Cutthroat trout distribution in Area No. (southern area). -27Table Iu Cutthroat trout records in Area No. I (Fig. 9). PRIMARY DRAINAGE and Tributaries ST MARY’S RIVER Kennedy Cr. So. Fk. Kennedy Cr. Beaver Cr. Beaver I. Duck-Lo Lower St Mary’s L. MILK RIVER Clear Cr. (Wind Cr.) Ross Reservoir Beaver Cr. So. Fk. Milk R. Mid. F k . Milk R. Livermore Cr. No. F k . Milk R. MARIAS RIVER Sec. I (Teton R.) No. F k . Teton R. (Muddy Cr.) (No. Fk. Muddy Cr.) Cow Cr. Deep C r. ■ Mid. F k . Teton R. W. Flc. Teton R. Tiber Reservoir Cut Bank Cr. Sec. I Cut Bank Cr. Sec. 2 Lower Mission L. Willow Cr. Ray L. (Two Medicine R.) Birch Cr. Dupuyer Cr. No. Fk. Dupuyer Cr. So. Fk. Dupuyer Cr. Blacktail Cr. Swift Reservoir Cutthroat Other. relation ■ game ,..Barriers to other fish '■ game fish NF C RWD HD b R BD-a C a a Information source 857,028 ■ 057 055 S57 857,058 057 058 d R RWL d RE C51i d e d d RE RE 058 M57 057,857 . RE c58 C RE d RW .053 057 C R 057 d HEW C58 C C b C C C d .d C RE RE . E . E BD-a 056,857 c5ii C56 D52 057 C5L 05k 058 055 056 D5k,058 056 055,857 857 D5k 058 RE REW RE .R a a - C RE RE ID E NF-a E RE RK ID-a d b b d d -28Table h, continued. Cutthroat 'Other relation . game to other fish ■ game fish PRIMARY DRAINAGE and Tributaries Mid. Fko Birch Cr. Big Badger. Cr. Four Horn L, (Limestone Cr.) .Cooper L= (Two Medicine Cr.) Little Badger Cr. KiyoL. So. Fk. Two Medicine Cr. (Deep Cr.) Dog Gone I. No. Fk. Two Medicine Cr. Railroad Cr. R RE REB a =** C a C C d d SUN R i m Sec. I No. F k 0 Sun R. "Willow Cr. Nilan Reservoir Cobbs L. Pishkun Reservoir Gibson Reservoir Big George Cr. (Open Cr.) Lalre Levale So. Fk. Sun R. (Smith Cr.) Wood L. CLARKS FORK COLUMBIA R i m Cabinet Gorge Reservoir Bull R. E. Fk. Bull R. Rock Cr* E. Fk. Rock Cr. Rock Cr. L. Noxon Rapids Reservoir Swamp Cr. Buck L. "Wanless L. Martin Cr. Sb. Fk. Martin Cr. c d d d d d . C d d d C Barriers Ihformat: source #8 #k,C# ID c& C$8 E . RE NF-a C$7 S$7,C$8 E EEW EEW NF NF C$6 S$7 C$3,S$7 c$h RBGW REWB RE RE RE RG REK B, C$8 c$h D$2,C$8 C$8 C$lt C$8 C$lt ■ a d ,b “ '■ RBGW 7 " ■ R C$7 C$lt C$8 ' d d b b b a ' d b a a b a , .REBW HEKD D E R * REBW E - W . ■ C$$,D$7 C$8 C$8 C$8 C$8 C$6 .1^8 C$7 C$8 C$8 D$7,C$8 C$8 -29Table !4.5 continued. PRIMARY DRAINAGE and Tributaries Vermillion R, Cataract Cr. Sims Cr. Willow Cr. Beaver Cr. White.Pine Cr. Big Beaver Cr. Deep Cr. Graves Cr. Prospect Cr. Dry Cr.. E. F k . Dry Cr. Khox Cr. Cooper Gulch Evans Gulch Glidden Gulch Cherry Cr. Thompson R« Sec. IThompson R . S e c . 2 W. F k . Thompson R. (Four Lakes Cr.) Cabin I. Fishtrap Cr. Beatrice Cr. (Mantrap F k 0 Fishtrap Cr.) (Radio Cr.) Fishtrap I. Little Thompson R. Little Rock Cr. ■ Big Rock Cr. (Twin Lakes Cr.) Twin L. McGregor Cr. McGregor L. . Lower Thompson L. . Boiling Spring Cr. Mid. Thompson L. Slimmer Cr. Davis Cr. Upper Thompson L» Buffalo Bill Cr. Cutthroat relation to other game fish b a b a b b c a a .b b a a a a Information source C£8 c# ci?8 C^6 C^7 C53 C^8 c# C^7 RED —' D R RE E - b C$1 c$h c$h C$3 C$3 .c$$ c$b C$8 C$7 C$7 b C b R RED D . C$6 C$7 C$8 E. , REDH — ED C$7. .. C$8 ^$8 C$6 , a d C .9 d . a b -a a • Barriers RED EB — — .BEDU E ■ E a - Other game fish C57,M£8 - —'. REL A d EDH : — a ' EDKH .d . E ' > e KW b b E . . C^6 c$b C$8 M?2,C^65D57 BD n$2,-D$i c$lj$7 M$2,V$7 B$6 056 c$$ -30Table it, continued. ’ P R H A R T DRAINAGE and Tributaries TIiATHEAD RIVER Sec. I Flathead R. Sec. 2 Revais Or. Jocko R. (Valley C r .) Hewolf Cr. Finley C r . . M d . - F k . „Jocko. R. Lower Jocko L. Upper Jocko E. Post C r . Mission Cr. Crow Cr. (Mud Cr.). Lake on Mud Cr. No. F k . Crow Cr. Little Bitterroot R. (Warm Springs 'Cr.) (Dry Fork Cr.) Dry Fork Reservoir Briggs Cr. Flathead L. Lake Mary Ronan Dayton Cr. Swan River Sec, I Swan R. Sec, 2 (Mud Cr.) Mud L. (Birch Cr.) Birch L . Bear Cr, Swan L. (Hall Cr.) Hall L. (Bond Cr.) Trinkus L. ,.Lost Cr. S o , .Fk. Lost Cr. No. Fk* Lost Cr, Cilly Cr. Cutthroat relation to other game fish d .d b d C C Other game fish Barriers .RIMf RDKM E RE Information source 027 D2.7,C28 C22 D27,G28 . E RE b HjJPRE D E C22 C28 C33 D26 D26,C28 . 027 C22 C23 d b d RE E RE c22 C26 D275C28 a e d b C ■ ID • •<: C2k a G d d a E RDKM RE d C .REDM RED W a b d d b a .b b . b C Q2.8. D27,C28 m 026 . D27,c27 c28 023 .023 D27 C27 R .ED RDK R ED v D R E ' c'22 ,'Cg .026. 023 c23 . 022 —31Table It5 continued. PRIMARY DRAINAGE and Tributaries (Cedar Cr,) Shay L 0 Fatty Cr, Fatty Cr. Reservoir Rainbow L. (No. Fko Cedar Cr.) Lower Fish L= Upper Fish L. Goat Cr. Lion Cr. Piper Cr, Piper L . Mid. Piper L. (Jim Cr.) Jim L. (Pony Cr,) Pony Lo Dog Cr. (Condon Cr.) Smith Cr. (Cold Cr.) (No. Fko Cold Cr.) 'Cold L. (Elk Cr.) (So o Fk. Elk Cr.) Elk L. (Glacier Cr.) Glacier I." Rumble Cr. Holland Cr. Holland L. Upper Holland L. Beaver Cr. , Lindbergh L. Crystal L. (Therriaults Cr.) Bunyan L. Jessup Mill Pond Creston L. Ashley Cr. Cutthroat relation to"other game fish Other game fish a . e a a a a b d b a a — - Barriers Information source CA D£6 CA,D£7 D£7 - D ^ 5C A C58 CA C#,#7 PA D57 PA a■ - C55 a e - CA MA C E EDWTRE - E D57 a 'a a C d d d b. d b .a a b d CA - CA - CA PA CA .057 C58 CA CA B RE ■ RED RB B RDK K — ■ RB RE CA 056 056 CA D57 -32Table h, continued= PRIMARY DRAINAGE and Tributaries (Truman Cr.) mid Bill Cr.' Lake Monroe Lone L= Ashley L= Stillwater River Uhitefish R= Haskill Cr. Whitefish L. Laay Cr. ■Uhitefish Cr, E= Fk= Whitefish Cr= Upper Mhitefish L= W= F k . Whitefish Cr. Spencer L. Logan Cr= Good Cr = Plume Cr. Cedar L. Talley L= Sheppard Cr. Dunsire Cr. Griffin Cr = Sylvia L= Lupine L. Meadow Cr. Martin Cr. Upper Stillwater L= Lebeau Cr. Sunday Cr. South Fork Flathead River Hungry Horse Reservoir Emery Cr. Hungry Horse Cr. Margaret Cr. Doris Cr. Lost Johnny Cr. ■ Wounded Buck Cr. Cutthroat relation to other game fish b e b C d d d b C a a d C b C a a b d C a C b ■a e a d a e b b b • e ' a e a a Other game fish Barriers E C£6 D56 D57 #7,C58 — G . RKW REDW RDKW ■ E LK E — — RD D R REW — — R RE E — E G “* RD RDW RDGW RE * ■- Information source ID ID #7,058 #7 c£6 c# C53 c# C£6 C56,D57 C5£ c58 c58 C58 C58 .C56 jD^7 C$8 C$6 C$8 C$8 C$6 C$8 D$6 C$6 C$8 C$6 D$6 ' D$7,C$8 D$7,C$8 C$8 D$6 C$it. D$6 C$8 D$6,G$8 .. ■33Table k) continued. 'PRIMARY DRAINAGE and Tributaries (Wildcat Cr.) Wildcat I,. Ryle Cr. ' Riverside Cr. Murray Cr. Clayton. Cr. Harris Cr. Felix Cr. Graves Cr. • Aeneas Cr. Handkerchief I. Black L. Logan Cr. Devils Corkscrew Cr. Baptiste Cr. Sullivan Cr. Quintonkon Cr. Soldier Cr. Lower Twin Cr. Twin Cr. Spotted Bear R . Bunker Cr. (Gorge Cr.) Sunburst L. Big Salmon R. Big Salmon Li White R. Youngs Cr. Hahn Cr. Danaher Cr. Camp Cr. Basin Cr. Limestone Cr. North Fork Flathead River Spoon I, Canyon Cr. Big Cr. 'Langford Cr. Mud X. Hallowat Cr. Cutthroat relation ■■ to ,other game .fish Other game fish a e e - E e a b TM: C REDG - 027,M28 - d £6 - c # G . c£8 E NF E — a a - D C a a Informat source '.D57,C£8 D£6 P56 #7 D$6 - C e .b . a b . e. e b e e e Barriers D27 D56 B^6 D^.7 D^6 D^6 .D # .. D26,C28 0^7 C^3 . - a - a mm a a - C^6 - c :31t. - C^8 C53 CS8 C53 , a b M2l,C%8 . c^U D k a - C RDM C E a C e a e . DZ7,C28 c $ h -* , , DW D57 D§6 'M* - 0^6 D56 ~3h~ Table Zt9 continued Cutthroat Other relation game to "other ‘ fish game fish PRIMARY DRAINAGE and Tributaries (Kletomus C r 0) Moose Lc Coal Cr, Cyclone C r 0Cyclone L 0 Quartz Cr. Moran C r 0 Hay Cr. Bowan Gr0 . Red Meadow Cr, Red Meadow L 0 Hawk Cr. Moose Cr. Miale C r » Yakinikak Cr. Tuchuck Cr. Colts C r 0 KOOTENAI RIVER Yaak R„ Sec0 I Yaak R 0. Sec. 2 Kilbrennan Cr. Kilbrennan L 0 Spread C r 0 So. F k 0 Yaak R 0 ’ Final L 0 Huskin L= W 0 F k 0 Yaak Ri , , . D57D D^6 D^7 D57,C28 W= ' DW T&6 == D — — BD D Dir C#,D# D# ■ D57 D^7 D57 D57 G$k » D^7 d C a b a ■ Information source #8 .a e a b e b b e C a a a b d a a Middle Fork Flathead River Crystal C r 0 Stanton Cr, Stanton L 0 Tunnel Cr. (Essex Cr.) (Marion Cr.) Marion L 0 Bear C r 0 (Lake Cr.) . Flotilla Lo Bowl C r . ' Barriers REDK E D27,CS8 C^3 C58 C^8 C56- W a d EDW c# D27,c27 e C D d 56 C^8 d d d c d a ' ’■a a b - a . REDW RE RED , RE .. RE = - R HF. C58 CS?. DS7,CS8 css CS6 CSii CS?. DS7,CS8 GS7,DS7 - CS? -32Table It5 continued Cutttooat relation to other game fish PRIMART d r a i n a g e and Tributaries (Be Fkc Taak Re) (Wnndy 'Cr =) Fish Lakes ' . Star Cr. Lake Cr. (Falls Cr.) Savage L= Keeler Cr. TSo Fko Keeler Cr. Halverson Cr=Benning Cr. Camp Cr, Bull Li. Stanley Cr= Ross Cr. O 8Brien Cr= Quartz Cr. Bobtail Cr. Pipe Cr. Tom Pole L= E= Fk= Pipe Cr. Libby Cr, Big Cherry Cr. Deep Cr= Swamp Cr = (Howard Cr.) Howard L= Fisher R= Wolf Cr. 'W= Fisher R. (Trail Cr.) Bear L= (Lake Cr.) Geiger L= Standard Cr= Pleasant Valley Fisher R. Leon L= Horseshoe L= Bootjack L= E= Fisher R= Silver 'Butte Fisher R. Jackson Cr. Other game fish Barriers Ihformatic source , ■a a c C$8 •'• \ - a b a a •b b d b C C C$8 RE C$8 C$8 CO C$8 OS c$$ E RE DKW E E REDW C$6 C$8 D$7 C$8 C$6 C$8 a ns C E RED b a b d a a d C d a C C$8. REDW C$8 C$7 • # 7 , C$8 c$$ c$$ ™ E REDf W C$8 C$6 (PO UO ■ C$8 C$li RE ■ R REDW C$8 C$8 D$7,C$8 D$7,C$8 os R a C$6. a C d C C C b b a .E RE KW E R RE RE C$$ D$7 c$$ C$6 D$7 c$$ ■ NF . C$3,D$7 C$6,D$7 D$7 “36“* Table It3 concluded. PRIMARY DRAINAGE arid Tributaries Cutthroat Other relation . game .to other ■ fish 'game fish ' Fivemile Cr. Sullivan Cr. . Tobacco R. St Clair Cr. Lost L. (Giffin Cr.) ■ Tlierriault Cr. Fortine Cr. (Lake Cr.) Lake on Lake Cr. Grave Cr. Dickey L. Dodge Cr. Carpenter I. Young Cr. Wigwam Cr. (Bluebird Cr.) Therriauli L. (Weasel Cr.) "Weasel I.. - ■d b '• C " a ■ " ' 'e . ■" , •' ■ ■b a Barriers RE E1 RE ID '1■ ■ : RE Information 'source D57 . #7 C58,M$8 D27,C28 D^6 D26,C58 0^8 < 3 9 C C C E RED K E a e a' - C53 b RKW 058 b D 056 ' b W HF C^7 057,957 C58 957 058 D36 “37Table Cutthroat trout records in Area Ho. 2 (Fig. id). PRIMARY D R A H A G E and Tributaries CIiARKS FCRK C CEUMBIA RIVER Clarks F k . Columbia R. Sec. I Clarks F k . Columbia R. Sec. 2 Clarks F k . Columbia R. Sec. 3 Siegel Cr. StrRegis R. little St Joe Cr. (Ward Cr.) Cedar Cr. Twelvemile C r . Deer Cr. Big Cr. Silver Cr. Randolph Cr. St. Regis I. Dry Cr. Pardee Cr. Cedar Cr. (lost Cr.) Oregon Gulch Lost L. Trout Cr. Ho. F k . Trout Cr. (Hemote Cr.) Miller Cr. Fish Cr. W. F k . Fish Cr. (Cedar log Cr.) Cedar log I. Ho. F k , Fish Cr. Straight Cr. (French Cr.) French L. So. F k . Fish Cr. Petty C r l . Hinemile Cr. Butler Cr. McCormick Cr. Sixmile Cr. Mill Cr. Albert Cr. Cutthroat relation to other game fish d d d a C Other game fish RBW RBDW RB REDB a — b E REW C a b b C b b a C b a b e .a d a b e C a b . c. c C a C a . a Barriers - E E E E EDB' RDBW R DBW — REW B RD — RED RED : ■ . . REBD ■ RE « E . ■ . ■. . - Ihformatii source C$6, C$Q #8 C$3 C$8 C$8 c$h C$8 C$8 C$8 #8 C$7 C$8 V$7,C$8 .0# . V$7,C$8 C38 C58 T>$7,C$8 B$6 C£8 V$7,C$8 CS8 CS8 D36 C$7 C$7 C$8 GS8 C£8 - c$h . C$h C$7 C$6 C$$ ■ . . -38Table 5a. continued PRIMARY DRAINAGE and Tributaries Gutthroat relation to other game fish Other gaine fish - Okeefe G r 0 a Bitterroot River Sec. I Bitterroot R, Sec. 2 O 8Brien Cr. Miller Cr. Bolo Cr. Mill Cr. Graves Cr. "W. F k 0 Lolo Cr. Howard Cr. E. F k . Lolo Cr. One Horse Cr. Eightmile Cr. (Threemile Cr.) ' Ambrose Cr. Bass Cr. Burnt F k . Bitterroot R. Kootenai C r . Big Cr. Bear Cr. Fred Burr Cr. Willow Cr. Mill Cr. Roaring Lion Cr. Skalkaho Cr. Newton GulchBear Gulch Tenderfoot Gulch Daly Cr. ■ Railrpad Cr. . Hog Trough Cr. 'Weasel Cr. So. F k . Skalkaho Cr. Sleeping Child Cr. (Camas Cr.) Camas L. lost Horse Cr. Twin L. 2nd Tin Cup Cr. Rye Cr. d d. a a d e ae REEW REDB C ED D b a a a b C b C b d a C Barriers C57 C58 C58 .056 C53 D5L,c58 D55 C57 D#l C58 c56 . C57 • M5L,D55,c58 RElW - — - — RED REBD RE E RE RE Information source C56 . 955,058 C58 c58 C58 955,057 058 ID - CBh RE a - 056 C56 C RED D5M58 951t e e e d • .© e e b d a C c , a C - D5L D51t 058 ' RD- - D . VBh VBh VBh VBh , RED ' 058 - 056 . 058 058 ED R HE 057 056 -39Table 5>5 continued. PRIMARY DRAINAGE and Tributaries Eo F k 0 Bitterroot R 0 Warm. Spring G r 6 . Cameron Cr, Meadow 'Cr = Swift Cr= Dense Cr= Bugle Cr= . ' . Moose Cr= ■ , "■ tick Cr = ■ ; Reynolds Cr= Sign Cr= Cuba Cr. Ripple Lo W= Fk= Bitterroot R= (H q u e t t Cr=) Shelf L= Piquett L= (Boulder Cr=) Dollar L= Boulder L= Neaperce Fk= Bitterroot R. Watch Tower Cr= Blue Joint Cr= Overwhich Cr= Hugheg Cr= Burrell Cr= Lake Cr.Emmett Cr = Woods Cr= Salt Cr. John,s on Cr = Sheep Cr= Rattlesnake Cr, Blackfoot River Sec= I Blackfoot R= Sec= 2 BlacIcfoot R= Seco 3 Gold Cr= Boulder L= Belmont Cr= Elk Cr= Cutthroat relation to other game fish d b b e .e e C$8 C$8 .B$!t .D$ii D$/4 D$1j. . D$^ D . . C REDW a a - d d d !Information source M£2yC58 ™■ C a 1a d a d a e e e e e e e e d Barriers REDW RED RED D C .e e e e e Other game fish 'D$ii • D$li D$i|. D$li b$l,C$8 D$$ D$$ C$8 C$8 .C$8 ED =RED C$7 C$3 C$8 D$lt D$ii D$ii D$L D$lt D$L . D$U • D$it . M$$,C$8 ' REW C REDB REDB REDB RED a d b RD REBD . . C$8 C$8 C$8 .ID C$6 C$lt D$6 D$6,C$8 -ItOTable continued. ' ,PRIMARY DRAHfAGE and Tributaries Clearwater R . Blanchard Cr. No. F k . Blanchard Cr. Harper E.. Salmon L. Owl Cr. Placid E.. ' i /' Finley Cr. Drew Cr. Morrell Cr. Trail Cr. Seeley L. Deer Cr. Fawn Cr. Sawyer Cr. W. F k . Clearwater R. Marshall C r . Marshall L. Lake Inez Lake Alva Rainy L. Cleanfater E. (Bertha Cr.) Summit E. Cottonwood Cr. Cottonwood E. Chamberlain Cr. Pearson Cr. Monture Cr. McCabe Cr. Dick Cr. Dunham Cr. Falls Cr. Warren Cr. No. F k * Blackfoot R. Rock Cr. Coopers L. Spring Cr. E. F k . No. Fk. Blackfoot R. Meadow Cr. Parker I.. Cutthroat relation to other game fish d C C b d C b b 'e d e d , c a e b e cl d C d e e b C a a d b C a a b . C e a a .b b a Other game fish REDW ■ REW E G RKW RED DK RE • RE DK E — ED ■—* ' DKW REDK DKW . — RE R 1— REDW EW RE -*• RE REDW R R - Barriers Information source C58 #k,C28 #7 D£63C$8 ■ C$8 C$8. C$8 c$b D$6 C$8 D$6 D$7,C$8 c$$ C$6 D$6 C$6,D$6 D$6 C$$,D$6 C$8 C$8 C$8 D$6 D$6 C$7 C$$ C$$,D$6 D$6 D$63C$8 C$7 C$7 C$6 C$6 C$6,M$7 .. D$6,C$8 D$6 D$7 D$3 .D$$ C$6 C$8 Table continued PRIMAEY d r a i n a g e and Tributaries Webb I. Wales C r » Y o u m a m e Cr. Nevada Cr. Sec. I Nevada Cr. Sec. 2 Douglas C r . Cottonwood Cr. Chimney Cr. Mud Cr. Murray Cr. Bear Cr. Sturgeon Cr. Nevada Cr. L v Arrastra C r . Stonewall Cr. Beaver Cr. Liverpool Cr. Blue Diamond Cr. Poorman Cr. Humbug Cr. Landers Fork Copper Cr. Heart L. (Hogan Cr.) Keep Cool Cr. Alice Cr. Wallace Cr. Schwartz Cr. Rock Cr. Sec. I Rock Cr. Sec. 2 Ranch'Cr. Stony Cr. Stony L » No. F k . Rock Cr. . Ross Fko Rock Cr. Helm Cr. (Condon Cr.) Medicine Cr. Beaver Cr. W. F k 0 Rock Cr. Mud L. Cutthroat relation to other game fish a c a d' b 'b d C d e e e C C C b c C C C e b b C C C a d d C a b e b e a b b e Other game fish Barriers Information source C£6 RK ' C- REDW . EDW REDW EB ■ E EB - ID ID ID #6 D56 D$6,#8 D56 ID ' ID #6,CZ7 C#3,MSU,D£6 :C # , # 6 C23,D26 D£6 D56, D56 RK ED EB E E E RED RE — RD G REB • .RED E V ' REDB REDB RED . D56,C28 C5? C08 C$7 C$.8 C$& ■ C$6 C$6 D$6 C$8 ■'C$8 . C$8 C$8 ' C$7 C$8 C$8 C$8 D$6,C$7,M$8 ■c.$7 K . REDW - E. . . .REDW - D$6 D$6 c$7 D$6 C$7 C$7 . D$6,C$7 M$3 Table 5>5 continued H t M A R T DRAINAGE and Tributaries Lake Abundance E e Fko Rock Cr. Eo Fko Reservoir Copper Cr. Mid, Fko Rock Cr. (Carp Cr.) ■Carp L. Phyllis Lo Cramer Cr. Harvey Cr. Flint Cr. Sec. I Flint Cr. Sec. 2 Lower Willow Cr. Douglas Cr. Boulder Cr. So. F k . Boulder Cr. (Copper Cr.) Dora T h o m L. Boulder L. Trout C r . Georgetown L. (No. Fk. Flint Cr.) Echo L. Hoover Cr. Millers Cr. E. Warm Springs Cr. Willow C r . Rock Cr. (Rock Cr. L.) Dolus L. Meadow L. 2nd Little Blackfoot R. See. I Little Blackfoot R. Sec. 2 Spotted Bog Cr. Trout Cr. ■ (Carpenter Cr.) Snowshoe Cr. Ophir Cr. Dog Cr. Hope Cr. Mike Renig Gulch Cutthroat relation to other game fish Other game fish a C REEW '-O ED REDW e C b e e d b d RE HEHD REB E ' EB E; . RED C c b b e - C G C ■d b a d d #8 C£8 D£6 #7 . C$7 M$6 D$6 C$8 C$8 C$8 C$6 0$8 .C$3 ■C$8 D$$ c$$ ' C C b b b b HhfOrmat source D$$ e ■a b b C b b Barriers E . REKG C$l| C$8 REB RE RE REBD . E REB C$$,D$6 C$6 B ' ■ BEDB REDB B E EB EB REB EB RE . C$6 C$6. C$8 C$7 . C$6 C$8 C$8 C$8 M$k,C$7 C$lt C$8 C?8.. C$8 .C$8 C$3,M$U Table concluded. PRIMART DRiIMG-E and Tributaries Telegraph Cr. Bryan Cr. Slate Cr. Laribee Gulch Racetrack Cr. Pozega L. Meadow E.. 2nd Fisher L. Little Racetrack Cr. 'Little Racetrack L=. Racetrack L. Lost Cr. Modesty Cr. Dutchman Cr. ('Warm Springs Cr.) Foster Cr. (Twin Lakes Cr.) Fourmile Basin L. Storm Lake Cr. Storm L. Mill C r . ' Clear C r . Beefstraight Cr. Cutthroat relation to other game fish C a a C > a a a e e e . 'c a Other game fish RE i" .E RE “ - 00 REB Barriers Information source .CS? G$6 #6 C5>7 .#baC58 Of# C58 P$7 C# D# . D # C# C RE C$k C$3 C RED C$6 b a C C a b RE C53 C$3 c# C$7 C$6 C$6 R RE E Table 6« Cutthroat trout records in Area No. 3 (Fig» ll). PRIMARY TEAINAGE and Tributaries MISSOURI Missouri Missouri Missouri RIVER Sec. I R. Sec0 2 R. See. 9 R. Sec. 12 Cutthroat relation to other game fish d d d •d Other game fish Barriers RBW RB RB .RBW Information source (#3 C33 c# C58 Musselshell River Sec0 I d REBW (#3 No. Fk. .FlatwIllow Cr. S o o Fk. Flatwillow Cr. (Fish Cr.) Rock Cr. Carters Pond American Fk.-Cr. (Lebo Cr.) Lebo If. Big Elk Cr. L Elk Cr. Reservoir Martinsdale I. So. Fk. Musselshell R. Cottonwood Cr. Mid. Fk. Cottonwood Cr. (Loco Cr.) Sandero Pond Forest I. (Alabaugh Cr.) Castle Lo (Noo F k 0 Musselshell R.) (Flagstaff Cr.) Holiday L. d d REB REB c# D2? C a d . REW C 56 C£8 REB .. REB E RE RB REB RE RE G53 Judith' River Sec= I Judith.S . Sec= 2 (Arrow C r .) Holgate Reservoir Kingsbury I. Dry Wolf Cr, Running Wolf Cr. Spring Cr. Cottonwood Cr. E. Fk= Spring Cr. Yogo Cr. d e d d d d b d a RE ns a C .a a d d „d C b C D^3 D& C$7 d ' d NF-a C53 D£2 ' #8 C53 #7 D^jCgG C^6 . RE C$&,S$6 RE ■ RE ■C$lt.?D$7 C$6,D$7 ** REB RE REBW RE RE RE. C$6 . C$7 C$8 C$8 C$1; C$8 C$3 c$$ =■1^Table 6, continued. PRIMARY DRAINAGE and Tributaries Soo Fko Judith R, Mido Fko Judith R. Lost Fkq Mid. Fk6 Judith Ro Wo Pko Lost Fko Harrison Cr, Weatherwax Cr. Cleveland Cr. Hell Cr. Shonkin Cr. Highwood C r 0 Noo Fko Highwood Cr. S o o Fk0 Highwood Cr= Pohlod Cr. Belt Creek Little Belt Cr. Main Cr. Noo Fk0 Little Belt Cr. So. Fk. Little Belt Cr. Logging Cr. Pilgrim Cr. Tillinghast Cr. Dry Fk. Belt Cr.. So. Fk'. Dry Fk. Belt Cr. Hoover Cr. Harley Cr. Jefferson Cr. Smith River Sec. I Smith R o. Sec0 2 Hound Cr. Eo Fk. Hound Cr. Mid. Fk. Hound Cr. Mid, Fk, Reservoir Dry Fk. ,,Smith R. Tenderfoot tir.1 Rock Cr. No. Fk. Rock Cr. So. Fkq Rock Cr. Eagle Cr. Sheep Cr, Cutthroat relation to other . game fish d d Other game fish RE REK RE O= RE . C a c a c. a d d RE RE RE E E E C C C d d a a .a d b . d ' d C .c , C d d ■ a, d C .BD-a .BP-a NF-a NF-a NF PB PEW. REB C C tm ■a, a d a -? RE • PW C#,S28 CS8,S# ,C#,S28 S^8 #3,858 #7,828 828 S28 b#,D26 C28 827 023 827 NF-a ID~a a b . Information source C22 . 827,028 827 827 .827 . NF-a 026,827 ' NF-a 021,827 . NF-a 827 827,028 . 828 NF-a BD-a $27 ' , NF-.a 827 NF-:a. ■ . 827 REBW REBW REBW RE E' a. " ./ NF-a NF BD-a BD REBW RE W « RE , RE RE RE RE. RE RE RE Barriers NF-a NF BD-a BD-a BD-a BD-a BD '023. D2lt 028 .021 ■ 827 ■ 82.7 #3 028,828 .027,828 '$28. ■ . .828 828. .. #1,927 Table 6, continued. PRIMARY DRAINAGE and Tributaries (Black Cr.) Butte Cr. Calf Cr. Little Calf Cr. Moose Cr. Deadmans Cr. Beaver Cr. (Camas Cr.) (Thomas Cr.) Benton Gulch Keep Cool Cr. Keep Cool Reservoir Elk Cr. Camas L. Big Birch Cr. Edith L. Gypsy L= No. Fk= Smith R. Fourmile Cr. Lake Cr. Boundary L. Sutherland Reservoir Sheep Cr. Dearborn R. Mid.F k 0 Dearborn R. Prickly Pear Cr. Canyon Cr. Cottonwood Cr. Virginia. Cr. Trout Cr. Little Prickly Pear Cr. Deadman Cr. lost Horse Cr. Elk Horn Cr. Beaver Cr. (Lake Helena) Prickly Pear ,Cr. Lump Gulch Clancy Cr. Tenmile Cr. Sevenmile Cr. Cutthroat. Other relation game to other. fish game fish ' a ' .b a C e a Barriers . RE — RE - BD-a BD-a — a a b REBW ■ C E b ■■ R C RE RG b E ' b d. REK b ■E E C — a ■ d REKW d REBW C REBW d RE d REB d REB ” a a RE b C RE RE d » a C .E RED d d b C C .C ■ RE RE E RE' E NF-a Information source (#3. C#,S28 SE>8 ■ 85:8 D^lt #3 C$7 C$7 C$7 C$7 C$7 C$7 D$$,C$7 C$7 D$6 ' C$3. S$8 S$8 ■ C$8 . C$8 C$8 . c$$ D$U5C$8 D$$9C$8 c$it G$$ C$3 c$7 HF-a C$6. C$7 S$7 C$7 C$6 C$2 C$6 C$7 C$7 -1,7Table 6^ concluded. PRIHflEY DRAINAGE and !tributaries Trout Cr. "x Hauser E. Canyon Ferry Reservoir Avalanche C r . Wnite Gulch Wilson Cr. Deep Cr. Greyson Cr. Crow Cr. So. Fk= Crow Cr. (Noo Fk= Crow Cr=) No. Fk= Crow Cr. I. Tizer I. Sixteenmile Cr. So. Fk= Sixteenmile Cr. Cutthroat relation to other game fish Other game fish d d d a■ RB RB RBN C C E E REBW RE REB RE d C d. d a C d d Barriers Information source .C£3 c £7 C$6 c$k C$3 C$$ C$3 C$7 C$6 B B 0# C$3 C$$ C$8 ■ ■ C$6 B e E REBtJ RE I I Table 7 e Cutthroat trout records in Area No® U (Fig. 12). ... . PRIMARY DRAINAGE and Tributaries Jefferson River (Willow Cr 7 r ~ ~ . . .Willow Cr. Reservoir (so; wliiow Cr:) Bell L: (No. Willow Cr.) Hollow Top L. So.Boulder R. Sailor I .‘ (Boulder R.) Bison Cr. .,Boulder Cr. Whitetall .Cr . (Big Pipestone Cr.) Delmoe L. Haney Cr. Fish Cr. Hell Canyon Cr. Big Hole River Sec. I Big Hole R. Sec. 2 Big Hole R. Sec. 3 Birch Cr. Pear L. Tub I. Willow Cr. Bond. Cr. Dubois Cr. ,. Cherry L. Rock Cr » Trapper Cr. Camp Cr. Canyon Cr. (Divide Cr.) No. F k . Divide Cr. Jerry Cr, (Tom Cr.) Hamby L. Wise R. . Pattengail Cr. Elk Cr. .. Cutthroat reiatipn to other game fish ,Other game fish d Barriers RBW .. information source ".... c# ' ... d REB C£8 b R 0^6 C C d R RE EB C$'8 . ,.C$6 C$3 .d d b REW REB EB C$8 C$7 b C b . b. REB d d d .0 d a d O a a C b C C d C ,.R RE 1 R ' REBG REBG EEBG E RB RE E -- RE E, E HE ' ■ c$$ C$lt C$7 G$8. G$$ C$6 ' C$8 C$8 . C$7 c$$ C$7 C$3 C$3 C$3. C$3 C$3 , C$3,D$6 C$6 . C$3,D$6 .RE. ' REB C$$ . C$6,D$7 C$8 C$8: C$8 C$1 C E d d REGW REG C B . Table J3continued. PRIMARY DRAINAGE and tributaries •'r Lacy Cr, Bobcat Cr, Mono Cr. David Cr, (Alder Cr,) Johanna L* (Deep Cr,) French Gulch American Cr, (Sevenmile Cr,) Twelve Mile Cr, Ten Mile Cr, Seymour Cr. Lamarche Cr. ('Wo F k „ Eamarche Cr.) Warren I. Fishtrap Cr. Mid. F k . Fishtrap Cr. Pintlar.Cr, , Pintlar L. (Plimpton Cr.) Thompson L . No. Fk. Big Hole R. (Mussigbrod Cr.)’ Mussigbrod E. Johnson Cr, .Tie Cr, (Ruby Cr.) Big Moose Horn Cr. (Trail Cr.) Joseph1Cr. (McVay Cr.) Boot L. (Swamp Cr.) Moose Cr, Steel Cr., Big Swamp Cr. (SIag^a-Inelt Cr.) SXag-a-melt L. Ajax L. Little lake Cr, Cutthroat relation to other game fish C C a b Other game fish EG E RE Barriers BD~a BD-a NF-a NF-a a Information source C£8,S£8 C27,S58 C#,Sg8 S^8 0^6 C a E C?6 CB6 ■o e e C C E RE a C C d d E E RE RE d d REGW REG C$3 C$it e d . C RE E D$6 C$6 . C$7 C ■ E C E d RBW C$7 RE REG E C$8,D$8 C$8 b d e a a C D^7 D57 C56 C$6 no . BD-a C$3 .C$6 S$8 C$6 C$6 - C$6 BD RU ‘ D$7,S$8 C$8 C$7 c$$ E - C$8 Table T3 continued. PRIMARY DRAINAGE and Tributaries (Hamby Cr.) (Englejard Cr.) Englejard L„ Warm Spring Cr. Governor Cr, Berry Cr. Jalanke Cr. Van Hbuten L. Jahnke L. (Dark Horse "Cr.) Dark Horse L= Ball Cr. Beaverhead River Ruby R 0 Sec. I Alder Gulch . Granite Cr. . Ledford Cr. Warm Spring Cr. Romey I. Rattlesnake Cr. Estler Cr. Estler L. Tent Lo Minneopa L. Blacktail Cr. E. Fk= Blacktail Cr. Indian Cr. W 0 Fk. Blacktail Cr, Grasshopper Cr. (Horse Prairie Cr.) (Medicine lodge Cr.) (Dad Cr.) Dad'L, . Bloody Dick Cr. Reservoir I, (Red Rock R.) Sage Cr. Sheep Cr. Deadman C r , Deadman I. Nicholia C r . Cutthroat relation to other game fish a d d C C d ■ a C e d d . .d a C e . a C C C a a C d C C d a ■c C C C C a b Other game fish Barriers Information source C£8 C£83S£8 c£8 C58 C56 C57 ' c# REGW EG EG E ■ RE ■n R E C56 C£8 RBW REB RE ID * RE - RE R R- W. - W REW REB E RE REB G# M523C£6 c# c# c# #7 . P28 . Cf>73S£8 C# C£6 C5>8 c$h ■ C28 C$7 C$6 C$6 - R E c$b C$6 c$$ RE RBW C$3 C$7 B BD 8^8 BD-a C$8 C$6,S$7 - R — 5i°° Table 7, continued FRmmY DRAINAGE and Tributaries (Cabin Cr=) (Indian Cr=) Morrison E. little Sheep Cr. E 0 F k 0 Little Sheep Cr. "Wo Fko Little Sheep Cr. (Willow Cr.) Birch Cr. long Cr. Schultz Pond (Lower Red Rock I.) Odell Cr... (Upper Red Rock L.) Elk Springs Cr. Elk E. Red Rock Cr. Hell Roaring Cr. Madison River Sec. I Madison R. Sec. 2 Madison R. Sec. 3 Madison R. Sec. It Ennis I,. (Meadow Cr.) No. Meadow Cr. Sureshot I. Twin I. McKelvey L. So. Meadow Cr. So. Meadow Cr. L. (Plain Spring Cr.) Axolotl L= Indian Cr. So. Fk= Indian Cr. (W. F k . Madison R.) (Cliff I.) Goose I. Elk R. Hebgen L= Grayling Cr. So. Fk. Madison R. Cutthroat relation to other game fish c C C C Other game fish K E R E G b RE b b d b RE GL REG E d d d d d RBGW REBG REBG RBGW RBG d b b a a b REB E R — R C C R R BD e™ a C d b d Information source C# G# C# C#6 a b a a Barriers 1» RB RBW RB RB c# E#B - #2,0# NF-a S# NF-a G # C# s 5 8 9d # s# c# # c G C 7 # # # 0# C# C# c# c# c# c# NF-a NF-a S58 c#,s# c# c# c# c# c$3 - -52Table 7 S concluded» PRIMARY H?AINAGE and Tributaries Gallatin River Eo Gallatin R e Ross Cr. Hyalite C r . Hyalite Reservoir Emerald I, Bridger Cr= ¥» Gallatin R 0 Sec0 I Wo Gallatin R 0 Sec. 2 Beck and Border Canal Spain Ferris Irr „ Ditch Highline Canal KLeinschmidt Canal Spanish Cr. S o o F k 0 Spanish Cr. Spanish Lakes Lake Solitude (Noo F k 0 Spanish Cr,) Chiquita L. Hell Roaring Cr= (Squaw Cr.) Rat Lo (Beaver Cr.) Beaver Cr= L. (Buffalo Cro) Ramshorn L» Taylor Fko Gallatin R= Speciman C r 0 Bacon Rind Cr, Cutthroat relation to other game fish d d Other game fish C G RBW RBW R . RE REBG RG REB RBW RBW = ca> tao <*n E RE .a a Ca C d b C d d d e e e e Barriers BD - a C R C R NF-a a C58 S58 C57 =■ a C C 58 C58 c5b C58 857,058 c5it C56 c5ii M5i,c5B,s58 m 5i M52 m 5i M5l 058 858 C58 c58 C58 a C Information source R R . BD-a - c58 c55 858 057,858 “53Table 80 Cutthroat trout, records in Area No, 5 (Fig, 13), PRIMARY DRAINAGE . and Tributaries IELEONSTONE Yellowstone Yellowstone Yellowstone Yellowstone RIVER See, 5 R 0--See', 6 R, Sec, 7 R. Sec, 8 R, Sec, 9 Bighorn River Little Bighorn R, Sec, I Little' Bighorn R, Sec, 2 Lodge Grass Cr. Elbow Cr, Rottengrass Cr, Black Canyon Cr, Dry Head Cr. Crooked Cr, Sage Cr, Pryor C r , , Eo Fie, Pryor Cr, Broadwater Drain Clarks F k 0 Yellowstone River Clarks Fk, R, See, I Clarks Fk, R, See, 2 Clarks F k , R. See, 3 Rock Cr, Sec, I Rock Cr, Sec, 2 Rock Cr, Sec, 3 Red Lodge Cr, Cooney Reservoir Willow C r . m i l o w Cr, E, (W. Fk, Rock Cr.) (Basin Cr,) Basin Cr, L« Falls F k , Rock Cr, Broadwater L, Keyser Brown L. First Rock Cr, L, Second Rock Cr, L. Cutthroat relation to other game fish d d d d C d d d b C d d b d d C C C d 0 d d d d C C d b b C C b b Other game fish Barriers RBW RBW REBW REBW RBW REBW REBlf REBW RB REB REB B E RE RE R . E RE RBW E RBW REBW REBW RE R ' REBW RBW E E E E E E Information source C58 C 56 C 58,M$8 C58 C58 C56 C56 c# 058 BD-a NF BD-a ID S58 D57 C58.S58 C58,S58 s58 C55,k57,s58 o5b c53 052,057 052 c22 c$k C$h C56 027 c56 c 2if 022 058 c5L 058 058 057 Table 8S continued PRIMAHT DRAINAGE and Tributaries (Hellroaring Cr0) Hellroaring L 0 Sliderock I 0 Glacier L 0 Hock Island L 0 Margaret L 0 (Broadwater H 0) Kersey L 0 Broadwater L „ Lady of the Lake Cr. Lady of the Lake Ovis L 0 Corner Li Stillwater River Sec0 I S=ETHwaGer H T -S e c T T Stillwater H 0 Sec0 3 W 0 Rosebud Cr. Fishtail C r 0 Fiddler Cr. Mystic L 0 (Eo Rosebud Cr.) Thunder L. (Goose C r 0) Goose L. Bridger Cr. Lower Deer Cr. Sweetgrass Cr. Campfire Lo Boulder River Sec. I Boulder H 0 Sec, 3 W. Boulder R 0 E 0 Boulder H 0 Elk Cr. Fourmile Cr. E. F k 0 Boulder R. (Big Timber Cr.V Swamp Cr. Cutthroat Other relation game Barriers to other fish game.fish C a ' b C E Information source #3 • C31t 058 Cf?3' C58 E E a C RE <{ E C C E RE Ch 7 c# #3 c# C28 #8 a a «0 d d d d d d RBinr RBW REBW REBinr REB REB C R c$U C$8 C R C$1* c# Cf>8 M38SC5U C$8 C$6 a C$1* C E C$6 b d B RB R C$6 C$8 C$6 C d d d d d C C C a REBW RE REBW HBW HBW R H REB HF HF C$8 G$8,M$8,D$8 C$7 C$8 C$3 D$8SM$8 C$8" C$3 c$3 Table 8, concluded Other game fish PRIMARY DRAINAGE and Tributaries Cutthroat relation to other game fish Little Timber Cr. Mission.Cr. Little Mission.Cr. a _ C RB - Shields Rivey Sec. I Shields R. Sec. £ Shields R. Sec. 3 ' Willow Cr. Rock Cr. Brackett Cr. Skunk Cr. Cottonwood Cr. (Horsefly Cr.) Horse Cr. (Cottonwood Cr.) Flathead Cr. Porcupine Cr, So. Fk. Shields R. Mill Cr. Fleshman Cr. Trail Cr. Spring Cr. Mill Cr. Passage Cr. Carpenter L. Sixmile Cr. Dailey L. Big Cr, Donahue Cr, Rook Cr, Tom Miner Or,• Mol Heron Cr, (Cinnabar Or,) Mill Or, Mol Heron L, Bear Cr, Billman Or. Hellroaring Or, Slough Or, Buffalo Fork (Lake Abundance Cr,) Lake Abundance a C d d a C C .b C REB RB EBW . REB REB B REB a - b b a REBW EB C E C E RE RBW R p* . « E RK REB d d C a a b d d a a o' a’ RE R E C R BD BD BD-a c58 C58 S57,C58 C58 C58,s58 C58 . C57 C58,S58’ c57 ED—a R n Information source■ C56 c53 C56 - b a b b a b a Barriers NF-a NF-a ■ 657,058 c57 C57 c57 c57 d57,c58 058 058,858 056 C56 o53 c58 c58 o55 '053 058,858 057 ' '057 057 057 o57 055 057,958 053,858 058 -5>6SBISffiET 1» The distribution of cutthroat trout (Salmo clarki) and some factors affecting it were investigated during the summers of 195>7 and 1958. 2o Distribution records were obtained from the following sourcesj 100 streams surveyed, east of the Continental Divide; 219 records from fisheries biologists and 769 from creel census returns (Montana Fish and Game Department); 35 records from the Montana State College collection. 3. ' ' Tlfty-five (75 percent) of the streams with cutthroat trout had J W populations of cutthroat trout above barriers. The important barriers were natural falls, high gradient areas, and beaver dams. It. Tdherever rainbow and/or eastern brook trout were present in association with cutthroat trout they were predominant. 5. Cutthroat trout are presently restricted to the headwaters of streams which originally were entirely inhabited by them= Influencing factors of their distribution are; stream habitat changes, competition with exotic species and hybridization with rainbow trout. 6. Taxonomic determinations were based upon the examination of 3b$ cutthroat trout (126 from streams that had never been stocked with rain­ bow trout), 51i rainbow trout and 88 rainbow X cutthroat trout. 7„ ho single characteristic was found to be adequate for identifi­ cation but when used in combination satisfactory separation of fish over l+,0 inches in total length was achieved. 8, Cutthroat trout were recorded from a total of 699 streams and -272IiU lakes in Montana. They -mere predominant (only game fish present or ranked first in relation to any other game fish) in 223 (38 percent) streams and lU2 (28 percent) lakes. 9. Records of cutthroat trout are listed and distribution plotted on m aps. / L I T E R A T U R E C ITED i Evermann, Barton W« 1893. A reconnaissance of the streams and lakes of western Montana and northwestern Wyoming. Bull. Ti. S. Fish Comm. 11:3-60. Evermann, Barton W. and Ulysses 0. Cox. 189U. Report upon the fishes of the Missouri River Basin. Comm, of Fish and Fisheries. 20:322-U29.. Jordan5 David Starr. I 889. Cutthroat abundance in Yellowstone Park. Comm. 9sUl-63. Lucke5 A. 1928. History records of Fort Assinneboine. U. S. • Bull. U. S. Fish Havre5 Montana. Miller5 R. R. 1920. Notes on the cutthroat and rainbow trouts with the description of a new species from the Gila River5 New Mexico. Ann ,Arbor5 Michigan. Univ. Mich. Press. Occasional Papers5 M u s . Zool. No. 229 U 2 p p . Wilkins5 L. P. 1922° Observations on the field use of Cresol as a stream-survey method. Prog. Fish Cult. 17(2-):82-66. . MONTANA STATE UNIVERSITY LIBRARIES 762 1001 90 O 136433