Curriculum Vitae Jill Lynn Silverman Professional Address: Department of Psychiatry and Behavioral Sciences University of California Davis School of Medicine Room 1001B Research II Building 96 4625 2nd Avenue Sacramento, CA 95817 Telephone 916-734-8531 FAX 916-734-5089 jill.silverman@ucdmc.ucdavis.edu jsilverman@ucdavis.edu Home address: 9370 Willow Pond Circle Elk Grove, CA 95624 jandmsilverman@gmail.com Education: 2007-2010 National Institute of Mental Health Laboratory of Behavioral Neuroscience Bethesda, MD Postdoctoral Training in Behavioral Neuroscience 2000-2007 University of Maryland, School of Medicine Program in Neuroscience Baltimore, MD Ph. D. Neuroscience 1994-1999 Rutgers University Rutgers College New Brunswick, NJ B.A. Biology and Psychology Honors: Psychology Professional Experience: 2012-present University of California, Davis Medical Center Medical Institute for Neurodevelopmental Disorders (M.I.N.D.) Department of Psychiatry and Behavioral Sciences Sacramento, CA Assistant Adjunct Professor • Manage strategies, experimental design and daily workload of postdoctoral fellows, postbaccaulareate fellows and student interns in complex tasks of expertise designed and executed by our translational research laboratory. 1 | JL Silverman Vitae • Educate, assign and evaluate a large group using rodent behavioral testing to evaluate novel pharmacological compounds to forward medicines development for the core symptoms of autism spectrum disorders. • Design and apply emerging technologies to investigate genes regulating complex behavioral traits involved in autism spectrum disorders. 2010-2012 National Institute of Mental Health, Laboratory of Behavioral Neuroscience Bethesda, MD Senior Researcher Technical Laboratory Manager • Manage strategies, experimental design and daily workload of postdoctoral fellows, postbaccaulareate fellows and student interns in complex tasks of expertise designed and executed by the Laboratory of Behavioral Neuroscience. • Perform administrative tasks, training sessions, and general laboratory maintenance and organization. Prepare and maintain the Laboratory of Behavioral Neuroscience animal care and use protocols. 2001-2007 University of Maryland, School of Medicine Baltimore, MD Graduate Research Assistant • Accrued technical knowledge and skills to participate in projects in several laboratories with emphasis in neuroendocrinology, neuroanatomy and molecular psychiatry • Originated and executed a novel research project in order to fulfill requirements for the degree of Doctor of Philosophy. 1999 – 2000 Progressive Living Units and Systems of NJ (NJ-PLUS) Absecon, NJ Cognitive Retraining Therapist • Clinical experience that included the planning, organization and execution of individual and group therapy sessions with an emphasis on cognitive skills to adults with traumatic brain injury as a lead supervisor of a community based rehabilitation program. 1996 – 1999 Rutgers University Department of Psychology Piscataway, NJ Research Assistant • Honors research project with an emphasis in neurotoxicology that assessed pharmacologically manipulated rodent behavior using complex behavioral paradigms and neurochemical data assessment by chromatography. 2 | JL Silverman Vitae Peer Reviewed Publications: Silverman JL, Babineau BA, Oliver CF, Karras MN, Crawley JN (2012) Influence of Stimulantinduced hyperactivity on social approach in the BTBR mouse model of autism. Neuropharmacology: Special Issue on Neurodevelopmental Disorders. In press Silverman JL, Oliver CF, Karras MN, Gastrell PT, and Crawley JN (2012) AMPAKINE enhancement of social interaction in the BTBR mouse model of autism. Neuropharmacology: Special Issue on Cognitive Enhancers. July 16 [Epub] Woehr M§, Silverman JL§, Scattoni ML, Turner SM, Harris MJ, Saxena R, Crawley JN (2012) Developmental delays and reduced pup ultrasonic vocalizations but normal sociability in mice lacking the postsynaptic cell adhesion protein Neuroligin2. §Equally Contributing First Authors. Behavioural Brain Research. In press. Brielmaier J, Matteson PG, Silverman JL, Senerth JM, Kelly S, Genestine M, Millonig JH, DiCicco-Bloom E, Crawley JN (2012) Autism-relevant social abnormalities and cognitive deficits in Engrailed-2 knockout mice. PLoS ONE. In press. Silverman JL, Smith DG, Sukoff Rizzo SJ, Karras MN, Turner SM, Tolu SS, Smith DL, Ring RH, Crawley JN (2012) Negative allosteric modulation of the mGluR5 receptor reduces repetitive behaviors and rescues social deficits in mouse models of autism. Science Translational Medicine. Apr 25;4(131):131ra51. Yang M, Abrams DN, Simon H, Katz AM, Zhang JY, Harris MJ, Saxena R, Kalikhman D, Scattoni ML, Wöhr M, Roullet FI, Silverman JL, Bozdagi O, Sakuri T, Buxbaum JD, Crawley JN (2012) Behavioral phenotypes of mice with an ankyrin repeat domain deletion of the Shank3 gene. Journal of Neuroscience. 2012 May 9;32(19):6525-41. Yang M, Abrams DN, Zhang JY, Weber MD, Katz AM, Clarke AM, Silverman JL, Crawley JN (2012) Low sociability in BTBR is independent of partner strain. Physiology and Behavior: Special Issue on 20 Years of IBNS. Jan 8 [Epub ahead of print] Silverman JL, Turner SM, Barkan CL, Tolu SS, Saxena R, Hung AY, Sheng M, Crawley JN (2011) Sociability and motor functions in Shank1 mutant mice. Brain Res 1380:120-37. Papaleo F§ , Silverman JL§, Aney J, Barkan CL, Chadman KK, Crawley JN (2011) Working memory deficits, increased anxiety-like traits and seizure susceptibility in BDNF overexpressing female mice. §Equally Contributing First Authors. Learning and Memory. 18(8):534-44. Silverman JL, Yang M, Turner SM, Katz AM, Bell DB, Koenig JI, Crawley JN (2010) Low stress reactivity and neuroendocrine factors in the BTBR T(+)tf/J mouse model of autism. Neuroscience 171:1197-1208. Silverman JL, Yang M, Lord C, Crawley JN (2010) Behavioural phenotyping assays for mouse models of autism. Nat Rev Neurosci 11:490-502. Silverman JL, Tolu SS, Barkan CL, Crawley JN (2010) Repetitive self-grooming behavior in the BTBR mouse model of autism is blocked by the mGluR5 antagonist MPEP. Neuropsychopharmacology 35:976-989. 3 | JL Silverman Vitae Bozdagi O, Sakuri T, Papapetrou D, Wang X, Dickstein DL, Hagahide Takahashi N, Kajiwara Y, Yang M, Katz AM, Scattoni ML, Harris MJ, Saxena R, Silverman JL, Crawley JN, Zhou Q, Hof PR, Buxbaum JD (2010) Haploinsufficiency of the autism-associated Shank3 gene leads to deficits in synaptic function and reciprocal social interactions. Molecular Autism. 1:15. Silverman, JL, and Koenig JI (2007) Evidence for involvement of ERβ and RGS9-2 in 17-β estradiol enhancement of amphetamine-induced place preference behavior. Hormones and Behavior 52(2):146-55. Sharifi, JL, Brady, DL, Koenig JI (2004) Estrogen modulates RGS9 expression in the nucleus accumbens. Neuroreport 15(15):2433-2436. Halladay, AK, CoyneT, Sharifi, JL, Seto, J, Wagner GC (2000) Avoidance responding following amphetamine induced dopamine depletion. Pharmacology and Toxicology 87(5):211-217. Invited Chapters: Yang M, Scattoni ML, Chadman KK, Silverman JL, Crawley JN (2011) Behavioral evaluation of genetic mouse models of autism. Invited book chapter, Autism Spectrum Disorders, Eds. David G. Amaral, Geraldine Dawson, Daniel H. Geschwind, Oxford University Press. Yang M, Silverman JL, Crawley JN (2011) Automated three-chambered social approach task for mice. Invited Unit, Current Protocols in Neuroscience, Unit 8.26 Manuscripts submitted: Lin L, Genestine M, Yan Y, Sonsalla PK, Matteson PG, Silverman JL, Crawley JN, Millonig JH, DiCicco-Bloom E (2011) Absence of autism associated gene Engrailed-2 (En2) produces abnormal forebrain monoamines and structures and depression-like behaviors. Submitted. Journal of Neuroscience. Manuscripts in preparation: Silverman JL, Gastrell PT, Karras MN, Solomon M, Crawley (2012) New touch screen technology for evaluating cognitive flexibility in mouse models of autism. In preparation Silverman JL, Barkan CL, Tolu SS, Crawley JN (2012) Comprehensive characterization of the BDNF-tg overexpressing transgenic mice. In preparation Silverman JL, Yang M, Crawley JN (2011) Automated three-chambered mouse social approach task for the evaluation of therapeutics. Invited Unit, Current Protocols in Pharmacology, In Preparation Invited Lectures: April 2012, Workshop on Mouse Ultrasonic Communication, Institut Pastuer, Paris, FR, Invited Speaker, “Pharmacological Rescue Using Assays of Social Communication in the BTBR” 4 | JL Silverman Vitae November 2011, Society for Neuroscience Satellite Symposium, Washington DC, Social and Communication Behaviors, Invited Speaker, “Mouse Assays of Social and Communication Behavior in the BTBR” March 2011, German Neuroscience Society, Gottingen, Germany, The social brain in health and disease symposium, Invited Speaker, “Pharmacological intervention in autism-relevant phenotypes in mouse models of autism” September 2009, Karolinska Institute, Stockholm, Sweden, Developmental Cognitive neuroscience 2 week postgraduate course, Invited Speaker “Phenotyping of mouse models of neurodevelopmental disorders” Abstracts and Meeting Presentations: P.T. Gastrell, M.N. Karras, M. Solomon, J.L. Silverman, J.N. Crawley (2012) New touch screen technology for evaluating cognitive flexibility in mouse models of autism. International Meeting for Autism Research 11th Annual Meeting, Toronto, ON, Canada, Submitted for May 2012. J.L. Silverman, D.G. Smith, S.J. Sukoff Rizzo, M.N. Karras, D.L. Smith, K.R. Fonesca, R.H. Ring, J.N. Crawley (2012) An mGluR5 negative allosteric modulator improves social deficits and decreases repetitive and stereotyped behaviors in mouse models of autism. International Meeting for Autism Research 11th Annual Meeting, Toronto, ON, Canada, Submitted for May 2012. J.L. Silverman, S.M. Turner, M.N. Karras, S.S. Tolu, S.J. Sukoff Rizzo, D.K. Bryce, D.L. Smith, D.G. Smith, R.H. Ring, J.N. Crawley (2011) mGluR5 blockade reverses autism-relevant phenotypes in the BTBR mouse model of autism. Society for Neuroscience 41st Annual Meeting, Washington D.C., November 2011. J. M. Brielmaier, J.L. Silverman, L. Lin, P.G. Matteson, S. Kamdar, J.H. Millonig, E. DiCiccoBloom, J.N. Crawley (2011) Autism-relevant behavioral phenotypes in Engrailed-2 mutant mice. Society for Neuroscience 41st Annual Meeting, Washington D.C., November 2011. M. Yang, D.N. Abrams, J.Y. Zhang, H. Simon, A.M. Katz, M.J. Harris, R. Saxena, M. Wöhr, F.I. Roullet, M.L. Scattoni, J.L. Silverman, O. Bozdagi, T. Sakurai, J.D. Buxbaum, J.N. Crawley (2011) Evaluation of phenotypes relevant to autism and Phelan-McDermid Syndrome in Shank3 mutant mice. Society for Neuroscience 41st Annual Meeting, Washington D.C., November 2011. L. Lin, M. Genestine, Y. Yan, P.K. Sonsalla, P.G. Matteson, J.L. Silverman, J.N. Crawley, J.H. Millonig and E. DiCicco-Bloom (2011) Absence of autism associated gene Engrailed-2 (En2) produces deficits in forebrain monoamines and structures and depression-like behaviors. International Rett Syndrome Foundation Meeting, Boston, MA May 2011. J. M. Brielmaier, J.L. Silverman, L. Lin, P.G. Matteson, S. Kamdar, J.H. Millonig, E. DiCiccoBloom, J.N. Crawley (2011) Selective social deficits in Engrailed-2 mutant mice. International Behavioral and Neural Genetics Society Annual Meeting, Rome, Italy May 2011. M. Yang, M.L. Scattoni, A.M. Katz, D. Abrams, M.J. Harris, R. Saxena, H. Simon, J.Y. Zhang, M. Wöhr, F.I. Roullet, J.L. Silverman, O. Bozdagi, T. Sakurai, J. D. Buxbaum, and J.N. Crawley (2011) Evaluation of phenotypes relevant to autism and Phelan-McDermid Syndrome in Shank3 mutant mice. 1st International Phelan-McDermid Syndrome Symposium, New York City, NY March 2011. 5 | JL Silverman Vitae J.L. Silverman, S.M. Turner, M.N. Karras, and J.N. Crawley (2010) Partial Rescue of Social Deficits in the BTBR T+tf/J mouse model of autism by AMPA receptor modulator CX546. Society for Neuroscience 40th Annual Meeting San Diego, CA November 2010. S.M. Turner, M.J. Harris, R. Saxena, J.L. Silverman and J.N. Crawley (2010) Comprehensive behavioral phenotyping of Neuroligin 2 mutant mice. Society for Neuroscience 40th Annual Meeting San Diego, CA November 2010. M. Yang, M.L. Scattoni, M.J. Harris, A.M. Katz, R. Saxena, S.M. Turner, L. M. Woldeyohannes, F.I. Roullet, J.L. Silverman, O. Bozdagi, T. Sakurai, J. D. Buxbaum and J.N. Crawley (2010) Selective deficits in social interactions in Shank3 mutant mice. Society for Neuroscience 40th Annual Meeting San Diego, CA November 2010. A.M. Katz, L.M. Woldeyohannes, M. Yang, J.L. Silverman and J.N. Crawley (2010) Neonatal and juvenile behavioral phenotypes in mice carrying a deletion of the Neuroligin-4 gene. Society for Neuroscience 40th Annual Meeting San Diego, CA November 2010. F.I. Roullet, M.L. Scattoni, J.L. Silverman, M. Wöhr, M. Yang, and J. N. Crawley (2010) Autism-like behavioral phenotypes in BTBR T+tf/J mice. International Behavioural and Neural Genetics Society 12th Annual Meeting, Halifax, Nova Scotia, Canada May 2010. L. Lin, P. Sonsalla, P. Matteson, J.L. Silverman, J.N. Crawley, J.H. Millonig and E. DiCiccoBloom (2010) Deficiency of Engrailed 2 (En2) produces abnormal development of forebrainprojecting, monoamine neurotransmitters systems and depression-related behaviors. International Meeting for Autism Research 9th Annual Meeting, Philadelphia, PA May 2010. J.L. Silverman, C.L. Barkan, S.S. Tolu, S.M. Turner, R. Saxena, D.D. Diagne, A.Y. Hung, M. Sheng and J.N. Crawley (2010) Behavioral phenotypes of Shank1 mutant mice. International Meeting for Autism Research 9th Annual Meeting, Philadelphia, PA May 2010. S.M. Turner, D.D. Diagne, M.J. Harris, R. Saxena, J.L. Silverman and J.N. Crawley (2010) Comprehensive behavioral phenotyping of Neuroligin 2 mutant mice. International Meeting for Autism Research 9th Annual Meeting, Philadelphia, PA May 2010. J.L. Silverman, S.S. Tolu, C.L. Barkan, S.M. Turner and J.N. Crawley (2009) MPEP, an mGluR5 antagonist, blocks repetitive behaviors in the BTBR mouse model of autism. Invited Guest, American College of Neuropharmacology 48th Annual Meeting, Hollywood, FL. J.L. Silverman, S.S. Tolu, C.L. Barkan, and J.N. Crawley (2009) Pharmacological intervention of autistic-like phenotypes in the BTBR mouse model of autism. Society for Neuroscience 39th Annual Meeting Chicago, IL M. Wöhr, F. I. Roullet, J. L. Silverman, A. Y. Hung, M. Sheng and J.N. Crawley (2009) Reduced ultrasonic vocalizations in mice lacking Shank1. Society for Neuroscience 39th Annual Meeting, Chicago, IL C.L. Barkan, S.S. Tolu, A.Y.Hung, M. Sheng, J.L. Silverman and J.N. Crawley (2009) Comprehensive behavioral phenotyping of Shank1 mutant mice. Society for Neuroscience 39th Annual Meeting, Chicago, IL 6 | JL Silverman Vitae J.L. Silverman, S.S. Tolu, C.L. Barkan, and J.N. Crawley (2009) Pharmacological intervention of autistic-like phenotypes in the BTBR mouse model of Autism. NIMH 13th Annual Scientific Retreat Gettysburg, PA. J.L. Silverman, C.L. Barkan, S.S. Tolu, and J.N. Crawley. (2009) Behavioral effects of acute administration of risperidone and MPEP in the BTBR mouse model of autism. International Meeting for Autism Research 8th Annual Meeting, Chicago, IL. C.L. Barkan, J.L. Silverman, S.S. Tolu, and J.N. Crawley. (2009) Behavioral phenotype of BDNF-overexpressing transgenic mice. International Meeting for Autism Research 8th Annual Meeting, Chicago, IL. K.K. Chadman, J.L. Silverman, C.L. Barkan, S.S. Tolu and J.N. Crawley. (2008) Comprehensive behavioral phenotyping of BDNF overexpressing transgenic mice. Society for Neuroscience 38th Annual Meeting Washington D. C. J.I. Koenig, J.L. Silverman, D.L. Brady-Bell and J.N. Crawley. (2008) Preliminary assays of neuroendocrine factors that could contribute to repetitive self-grooming in the BTBR T+tf/J mouse model of autism. Society for Neuroscience 38th Annual Meeting Washington D. C. J.L. Silverman, K.K. Chadman, Y. Ayhan, M.V. Pletnikov and J.N. Crawley. (2008) Inducible expression of mutant human DISC1 does not alter mouse social approach behaviors. Society for Neuroscience 38th Annual Meeting Washington D. C. C.L. Barkan, J.L. Silverman, S.S. Tolu, K.K. Chadman and J.N. Crawley. (2008) Preliminary behavioral phenotyping of BDNF overexpressing transgenic mice. NIMH 12th Annual Scientific Retreat Gettysburg, PA. J.L. Silverman, K.K. Chadman, Y. Ayhan, M.V. Pletnikov and J.N. Crawley. (2008) DISC1 mutant mice display normal social approach behaviors. NIMH 12th Annual Scientific Retreat Gettysburg, PA. J.L. Sharifi-Silverman, D.L. Brady, R.L. Neve and J.I. Koenig (2007) RGS9-2 overexpression in the nucleus accumbens shell reverses estradiol enhancement of amphetamine-induced reward behaviors. Society for Neuroscience 37th Annual Meeting San Diego, CA. J.L. Sharifi-Silverman, D.L. Brady, R.L. Neve and J.I. Koenig (2007) RGS9-2 overexpression in the nucleus accumbens shell reverses estradiol enhancement of amphetamine-induced reward behaviors. NIMH 11th Annual Scientific Retreat Gettysburg, PA. J.L. Sharifi-Silverman and J.I. Koenig (2007) A proposed mechanism for gonadal hormone enhancement of amphetamine-induced reward behaviors. Johns Hopkins Bloomberg School of Public Health Conference for the Dissemination of Student Research on Addictions, Infectious Disease, and Public Health Baltimore, MD. J.L. Sharifi and J.I. Koenig (2006) Gonadal hormones modulate place preference performance through reduction of RGS proteins. Sixth International Congress for Neuroendocrinology Pittsburg, PA. 7 | JL Silverman Vitae J. L. Sharifi, D. L. Brady and J. I. Koenig (2005) Effects of estrogen on amphetamine induced modulation of RGS protein expression in the striatum and nucleus accumbens. Sex and Gene Expression. Society for Women’s Health Research 6th Annual Meeting Winston Salem, NC. J. L. Sharifi, D. L. Brady and J. I. Koenig (2004) Nucleus accumbens expression of DARPP-32 is regulated by estrogen. Society for Neuroscience 34th Annual Meeting San Diego, CA . J.L. Sharifi, D.L. Brady and J.I. Koenig (2003) Effects of gonadal steroids on regulators of G-protein signaling in the nucleus accumbens. Society for Neuroscience 33rd Annual Meeting New Orleans, LA. J.L. Sharifi, D.L. Brady and J.I. Koenig (2002) Estrogen modulates intracellular signaling proteins in the nucleus accumbens in response to amphetamine. Society for Neuroscience 32nd Annual Meeting Orlando, FL. J.L. Sharifi (1999) Altered sensitivity of haloperidol on conditioned avoidance response following chronic amphetamine treatment. Rutgers University Psychology Honors Thesis. Teaching and Instruction: 2008-2011, Course Coordinator and Lead Instructor Organize, prepare and execute a training course in animal handling and training for performing behavioral tasks in mice for incoming members of the Laboratory of Behavioral Neuroscience and intramural and extramural collaborators. Organize, prepare, maintain and update manual text accompanying the training course Annual leadership of this important, interactive course involves schedule, design, materialization of each focus session, evaluation of trainees, development and maintenance of a comprehensive manual that accompanies live training and providing aid and mentoring to participants on a continual basis. Past and Present Funding: NeuroTherapeutics Pharma Inc. Industry Award, 2012SPO Project Number 201121854 PI Jacqueline N Crawley, University of California, Davis Health System Co-PI Jill L Silverman A02 Award Project #201121854 Total Direct Costs: $139,128 New Jersey Governor’s Council for Medical Research and Treatment of Autism, 2010-2012 Co-Investigator (unfunded in accordance with NIH Intramural policy) Developmental Role of Engrailed-2 in Regulation of Forebrain-Projecting Monoamine Systems PI Emanuel DiCicco-Bloom, Robert Wood Johnson Medical School, University of Medicine and Dentistry, New Jersey, Piscataway, NJ 8 | JL Silverman Vitae NATIONAL RESEARCH SERVICE AWARD, 2004-2007 Estrogen Modulated Dopamine Signaling. PI F31 DA017027-01A2 Department of Health and Human Services, National Institutes of Health, National Institute on Drug Abuse NIMH Intramural Funds, 2007-2011 Unfunded Co-Investigator on1 ZIA MH02179-24 Animal Models of Neuropsychiatric Disorders PI Jacqueline N Crawley NIMH Intramural Research Program to Laboratory of Behavioral Neuroscience Awards: 2012 NIH Performance Award for Research Excellence 2011 Howard Hughes Medical Institute Student Internship Mentor Award 2011 NIH Performance Award for Research Excellence 2010 NIH Performance Award for Research Excellence 2009 NIH Performance Award for Research Excellence 2009 Howard Hughes Medical Institute Student Internship Mentor Award 2008 NIH Performance Award for Research Excellence 2008 Howard Hughes Medical Institute Student Internship Mentor Award 2006 Young Investigator Travel Award for Sixth International Congress for Neuroendocrinology Pittsburg, PA. 2005 Women’s Health Research Organization Travel Award for Sex and Gene Expression Annual 6th Annual Conference 2003 Graduate Merit Award $2000, University of Maryland Graduate School. 2000 Pre-doctoral NIH Training Grant in Neuroscience, University of Maryland. 1994-1998 Garden State Scholar, Rutgers University. Peer Reviewer: Autism Research Behavioural Brain Research Brain Research Brain Research Bulletin Genes, Brain and Behavior 9 | JL Silverman Vitae Hormones and Behavior Journal of Pharmacology and Experimental Therapeutics Journal of Neuroscience Methods Neurobiology of Aging Neuropsychopharmacology Neuropharmacology Neuroreport Neuroscience Pharmacology Biochemistry and Behavior Physiology and Behavior PLoS ONE Psychopharmacology Professional Memberships: 2010-present 2008-present 2002-present 2002-present 2001-present 2003-2007 2002-2005 American Association for the Advancement of Science International Society for Autism Research Society for Neuroscience Women in Neuroscience Psi Chi National Honor Society of Psychology American Neuroendocrine Society University of Maryland, Baltimore Program in Neuroscience Training Committee Daily Supervision and Mentoring of Post-baccalaureate Fellows: 2011 2011 2010-2012 Gastrell, Phillip, Post-baccalaureate, Duke University Medical School Oliver, Chicora, Post-baccalaureate Karras, Michael, Post-baccalaureate, University of Kentucky, College of Medicine Turner, Sarah, Postbaccalaureate Barkan, Charlotte, Postbaccalaureate, Columbia University 2009-2011 2008 NIH Summer Student Interns: 2010 Tolu, Seda, University of Maryland, College Park, MD Howard Hughes Student Interns: 2010 Simon, Harrison, Winston Churchill High School, Potomac, MD, Stanford University, Palo Alto, CA 2009 Diagne, Dieynaba, Blake High School, Silver Spring, MD University of Pittsburgh, Pittsburgh, PA 2008 Tolu, Seda, Magruder High School, Gaithersburg, MD. University of Maryland, College Park, MD 10 | JL Silverman Vitae