9/13/12 Figure 50.13 Zonation in the marine environment Life on Earth - Timescale 3.4 BYA - heterotrophs 2.7-2.2 BYA O2 begins to Accumulate 2.2 BYA first Eukaryotes 700 MYA sharp increase in Atmospheric O2. Reached modern levels in Cambrian 570-510 MYA 450 MYA life on oceans surface and on to land Life on Earth • Living things cause change • Living things respond to change • Living things change their environments • Living and non-living components of our Earth interact • Processes like global warming/climate change follow large-scale patterns, but it is the composition of life on earth that can affect those patterns • Ecological systems exist in balance - that balance can be disturbed, and its evolution from there can be difficult to predict. Life - as we know it • Organized internal structure - share a similar chemical composition – Composed of a common organization of atoms, molecules, cells... with emergent properties. • • • • DNA/RNA Respond to stimuli, coordinated growth and development Capture and transform energy from their environment Maintain internal conditions separate from external – Composed of cells • Arise through reproduction • Can adapt through mutations and adaptations • Effect change on their environment Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Organic compounds Photosynthesis/Respiration Photosynthesis: • Plants use carbon dioxide and produce starch and oxygen Respiration • Animals use starch/ sugar and oxygen, and produce carbon dioxide H2O + CO2 = Starch/sugar + O2 Starch/sugar + O2 = H2O + CO2 2-7 1 9/13/12 Photosynthesis • The synthesis of organic compounds (sugars) from simple inorganic compounds (CO2 and H2O) in the presence of chlorophyll using light energy from the sun. – General Formula for PS – The Organs of Photosynthesis - Leaves and Chloroplasts – Chloroplast Structure and Photosynthesis 6CO2 + 6H2O + • The Light Dependent Reactions • The Light Independent Reactions (Dark Reactions) • Coupling the Light and Dark Reactions = C6H12O6 + 6O2 – Photosynthetic Pigments and the Electromagnetic Energy Spectrum Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Photosynthesis/Respiration Photosynthesis: • Plants use carbon dioxide and produce starch and oxygen H2O + CO2 + SUN = Starch/sugar + O2 Energy pyramid Figure 2.15 Respiration • Animals use starch/ sugar and oxygen, and produce carbon dioxide Starch/sugar + O2 + ATP = H2O + CO2 6CO2 + 12H2O + sunlight ---> 6O 2 + C6 H12O 6 + 6H2O 2-13 Source: Data from Howard T. Odum, “Trophic Structure and Productivity of Silver Springs, Florida” in Ecological Monographs, 27:55-112, 1957, Ecological Society of America. Biomagnification Energetics predicts few top predators. Not enough energy at top trophic levels Top predators are also often K-selected (low reproductive rates/output.) Biomagnification is process that concentrated toxins at high trophic levels 2 9/13/12 Two Main Classes of Cells Eukaryotic Cell (animal) Prokaryotic Cell (bacteria) • Prokaryotic (Bacteria and Archaea) – Pro = “Before”; Karyon = “Kernel” – No nucleus, DNA coiled up inside cell • Eukaryotic (Everything else) – Eu = “True” – DNA inside membrane bound organelle inside cell, the nucleus Basic Properties of Molecules in Motion Osmosis- passive transport of water across a membrane • Diffusion: the random movement of molecules from a region of high concentration to a region of low concentration. – Concentration gradients • Osmosis: the diffusion of water… – Osmotic pressure – Isotonic, hypotonic and hypertonic environments for plant and animal cells Osmosis: problems with salt Isosmotic • Osmoconformers - change with outside salinity • Osmoregulators - regulate internal saltiness separate from the outside • Isosmotic: same inside and outside • Hyperosmotic: saltier inside • Hypoosmotic: saltier outside Sea Cucumber 3 9/13/12 Hyperosmotic: fresh water fish Endothermy in fish! ? Hypoosmotic: Marine Fish Reproductive strategies SEX (meiosis): NO SEX (asexual) • Internal fertilization • Fission - parent splits (few or many) • Budding - parent • Nest laying (brooding) develops small growth • Broadcast spawning (mass release) This is mixing genes! This is cloning! Sex? Sexual reproduction • Introduce new genes • Diversity in phenotype and genotype • Resistance to disease • Resistance to environmental change: Adaptation • • • • • Hard to find a mate Picking the right one Mutations/problems Parental investment? Don’t know exactly what you get 4 9/13/12 Sex? Some can do both! Asexual reproduction • Fast • No diversity (easily wiped out) • Know exactly what you get • Problem (if one exists) is reproduced • Can spread your exact • Need a self recognition genes mechanism • Cover an area quickly • No need to find a mate • Combination of sexual and asexual reproduction can bring the best of both worlds • Hermaphrodites: no need to find the right sex! These are organisms that are both sexes at once, or can change from male to female or female to male. Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Reproductive strategies Populations: what affects their size and growth? • • • • • • • • 3-10 Density (#/area) - carrying capacity (K) Natality - birth rate Mortality - death rate Age distribution/sex ratio Spatial distribution Resource availability Species interactions Migration/emmigration Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Population oscillations Predator-prey oscillations Figure 3.18 Figure 3.19 3-7 3-8 Source: Data from D. A. MacLulich, Fluctuations in the Numbers of the Varying Hare (Lepus americus), Toronto: University of Toronto Press, 1937, reprinted 1974. 5 9/13/12 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. J and S population curves Competitive exclusion Figure 3.20 3-9 Major Components of the Environment • Abiotic components, which consist of nonliving chemical and physical factors, such as temperature, light, water, minerals, and air • Biotic components, which include the living factors—all the other organisms that are part of an individual’s environment. Species Interactions (Biotic): Interspecific and Intraspecific • Competition – Competitive exclusion – Resource partitioning • Predation • Symbiosis – Mutualism – Parasitism – Commensalism Zooxanthellae: the key to coral reef productivity 6 9/13/12 Speciation, adaptation, and competition What about analogous structures? • Co-evolution • Convergent evolution What is a niche? Competition for resources leads to: • Competitive exclusion (one ‘wins’) or • Resource partitioning: the resource is ‘shared’ Adaptation by the process of Natural Selection • Survival and differential reproductive success over a period of time • Traits that increase ‘fitness’ are ‘selected for’, and are passed on through generations Ecosystem structure Predators: Sea Otters • Keystone species - is there such a thing? – A species whose ‘role’ or niche has a major impact on the structure of an ecosystem – Some case studies? 7 9/13/12 Sea Otters and Sea Urchins: a kelp forest paradigm Ecosystem change: an example Sea Urchins eat kelp, especially new recruits If kept in check, they eat drift kelp If populations expand, they will eat established kelp Sea Otters eat urchins, especially exposed ones They will keep sea urchin populations in check The Aleutian Island studies Sea Otters as a keystone predator Reduction of coral = increase in Algae. This shifts functional Groups of species, and affects Primary productivity 8