6 CllAP'rER I ( .. WIND RIVER DPERIMENTAL FOREST HISTORY Bay R. Silen In 1933, a 10,310 acre block of the Columbia National Forest (now Gifford Pinchot National Forest) in aouthwestern Washington was formally designated as the Wind River Experimental Forest. an historic event. This action was more a name change than Although a formal history might concentrate on events after that date, Wind River had long before beccme the focal point of forest research in the Douglas-fir Region. Wind River is aptly called the •cradle of Forest Research in the Pacific Northwest. • No other site in the west provides so concentrated a capsule of forestry frcm its beginning to the present. First experiments at Wind River began when a 5-acre spot in the valley was . cleared in 1909 for a nursery and eastern white pine was included among the first seed lots sown. In most years since, sane experimental savings have been done for experimental purposes. By 1913 part of the expanded clearing was formally called the Wind River Forest Experiment Station, and research activity continued to expand there through the next two decades. In 1908, a year prior to the first nursery clearing, Forest Service acSministration in the Northwest was reorganized. '1'he Bureau of Forestry was decentralized with establishment of a District Forester's Office (later called Regional Office) in Portland. Two aeetions of this office were involved with research at Wind River for a number of years. The Section on Planting under Julius F. JCunmtel founded the nursery and conducted nursery and regeneration studies. These were ..inly informal studies that were reported annually in ' I' 7 nursery reports which still are aaintained Nursery. Rive as a ccmplete file at the Wind 'l.'here are relatively few articles in scientific journals that . reflect this work at the Wind River Hursery. 'l'he Section on Silvie& under 'l'hornton T. Munger conducted formal as well as informal studies. The Douglas-fir Heredity Study and Wind River Arboretum, both started in 1912, were formal projects still maintained for their long-te:cm research value. This section eventually evolved into the present Experiment Station. Organizationally, Forest Service research activities were offically located at Wind River in 1913 on a site ajoining the nursery. Following a pattern in other western regions, the site was called the Wind River Experiment Station. Administrative control remained with the District Forester's office until 1919, then research jurisdiction shifted to Washington D. c. office. The Director of the Wind River Experiment Station from 1913 to 1924 was Julius v. Hoffman (he used the initials •J.v.• presumably to distinguish from Julius ICulllmel) . The northmost dwelling-site building in the present row of structures on the site was erected as the Experiment Station headquarters. When the Pacific Northwest Forest Experiment Station was established in 1924, the Wind River Experiment Station became an important branch location. By then a 160-acre tract adjacent to the nursery in the Wind River Valley was devoted exclusively to experimental use and dotted with many studies. In 1932, Leo Isaac was assigned to rec011111end on additional lands in the area that would be suitable for experimental forest designation. •The committee on experimental forests and ranges decided in 1930 to and natural areas set aside an experimental forest for the Douglas-fir (sic) type on the COlumbia- National Forest in the . -· . .... .,. , . . 8 vicinity of the Wind River Branch Station at Stabler, Washington. .. In accordance therewith, two areas of the !NZ'rounding country were examined and maps and a preliminary report prepared during the past year." Be looked for old growth stands, young stands, and cutover or burns to provide ample area for experimental plots in each of these forest conditions. The 10,300 acres set aside in 1932, including the original 160 acres, were in two blocks or •Divisions•--Trout Creek and Panther creek. The Trout Creek Division contained 6,500 acres of mature and overmature forests, burns and cutover lands. It included a natural area of 280 acres, set aside in 1926, which was later expanded to 1,180 acres. Be stated that--•by including the non-timbered sections between the area and the Wind River Station, the present experimental reservation and the proposed experimental forest will become a continuous ccmpact unit and will include Douglas fir (sic) land in several natural stages, i.e, remnants of fire-killed old growth, 30-year burn with varying degrees of restocking, non-restocking brush, and fresh burns partially replanted.• The Panther Creek Division was primarily second growth Douglas-fir forest. This specific point was made regarding Panther Creek: •The Panther creek Division constitutes the even-aged young growth (90 year) Douglas fir (sic) area. Since there was no area near this age class in the Trout creek watershed and it is a type in which much private cutting is now being done and a size at which Douglas fir (sic) will probably be cut under management, it was decided to include this area in the experimental forest." In total, stands on the experimental forest are considered typical of the cascade Range hum.id forest in the cascade Range. ( Formal establishment of an Experimental Forest assured control over ccmpeting forest uses that might jeopardize long-term study plots. �------ ------·-·· 9 Since 1933 there bas been no official change in the area's status as an experimental forest, except for ainor boundary adjustments, that brought the acreage to 10,815 as of January 1, 1983. Research activities and emphases have undergone changes every decade in response to national and regional events. The period up to world war II -s characterized by gradual expansion of progr ams and facilities at Wind River by research pioneers in the Forest Service. During the 1930's, several residences and a headquarters building were constructed under the CCC program and enrollees also helped with research studies. Research activities ebbed during world War II but expanded after the war with emphasis on timber harvest and regeneration problems. 'l'he first long-term growth, vegetation, and 1110rtality plots in a Northwest natural area were soon established and the dynamics of rodent populations in clearcuts and adjacent stands were studied. Harvest cutting trials· were installed in both old-growth and second-growth stands on the Experimental Forest. Wind River's role as a prilllary research location declined after the mid-SO's as programs at other field locations were developed and as national emphasis transferred scientists away frCllll field headquarters to scientific laboratories. After 1960, Wind River Experimental Forest was no longer manne d and Experiment Station buildings were transferred to the Wind River Ranger District. Since then existing and new studies have been assigned to be administered by projects headquartered at Forestry Sciences Laboratories in Olympia and Corvallis. An effort in the late 1960's -s made to bring the old growth stands on Trout Creek Hill under a management plan. Under the direction of District Ranger Beal, in 1974 Jon s. BumStead ccmpleted a Troiit Creek Hill Management Plan. It specified that the primary objective of Research and National Forest I / I lO Administration -s to regenerate the 2,800 tract in successive entries. With • this first research entry of 8 clearcut an.its in 1974-1979 for forest residue reduction by Portland Scientists was initiated on 'l'rout Creek Jlill. Methods of ' treating residue and levels of utilization -re studied. 'l'his -s followed by Ol.ylllpia Scientists installing spacing studies, mixed species plantations, and related regeneration growth studies on the clearcuts. Methods of regeneration differed fran those envisioned in the 1974 plan. 'l'he impact of the Wind River Experiment Station and Experimental Forest on aound forestry practices in the Douglas-fir Region has been very great. Al.though most of the Station's research is now centered elsewhere, Wind River always has l>een and remains the focal point. Nursery practices everywhere are still founded on pioneer findings learned at Wind River Nursery. Silvicultural concepts and cutting practices.developed out of the trials, e ven mistakes, of the Wind River experience. Re£orestation methods, especially concepts concerning natural regeneration, data from studies conducted at Wind River. still rely heavily on '1'he genetic studies still in progress at Wind River continue contributing important information. of many leaders in the Forest service, both in research were molded and and Careers administration, by their research experience gained in the Wind lliver valley. ,; 19 CHAP'l'ER IV FINDINGS AND DY RESEAJCH ROy R. Silen INTRODUC'l'ION '1'he lull in research activity during World War II provides a convenient point in time to separate research at Wind River Experimental Forest into two logical time periods. '1'he scientists who dam.inated forest research in the Northwest prior to World War II were winding down their careers by then. A • surge of new Station scientists associated with Wind River soon after World war II have, by now, completed their careers. It was these two surges of effort that contributed the majority of research at Wind River. Research climate of the two periods were quite different. Forest Service emphasis of the first was mainly on protection· and assessment of the resource. In the second, harvesting and its associated problems was the dominant emphasis. Research in the first period was mainly controlled observation featuring broad ranging, pragmatic, ecologically oriented goals. There were few true specialists. In the second period specialization and sophistication became dominant at the loss of research broad questions. acme capacity to effectively 'l'here were few true generalists. Administration of research in the first period was low key, local, or by scientists in Portland who were still active in research at Wind River. In the post World War II era local administration flourished briefly, then the Experimental Forest administration became more and more distant as it shifted to the Olympia canter, and diffused to projects of the Forestry Sciences Laboratories at Olympia and Corvallis. The research findings are divided into contributions frcm each period. Where single studies contributed significantly these are highlighted. r A more I ! 20 comprehensive coverage was attempted 1'y listing lifetime achievements of a group• of the main scientists who worked there. 1908 TO WORID WAR II The main research activities during this period involved nursery, planting and 11eeding, natural regeneration, precOlllr llle cial thinning, provenance trials, species trials, spacing, silvicultural, stand development and monitoring natural areas. • Nursery Studies Nursery studies were included with the first aowings in 1909. By 1911, c. P. Willis produced a separate section of the annual Wind River Nursery report on regeneration studies. , { Scme idea of the scope of the 1911 effort is provided from the annual production of over a million seedlings. p0rt questions addressed in this single annual re Research -re drill versus broadcast saving, seed covering techniques to promote more even germination, irrigation schedule trials, thinning densities of 1-0 stock, sulphuric acid treatment for damping off, late summer transplanting, trials. and gang dibble versus planting board Similar scope of trials or research -re COlllDIOn to each subsequent annual report. It is safe to say that most of the practical questions of nursery and planting practices received some research at Wind River between 1908 and World war II. For many years thi• was the only nursery in the Region. However, there waa aCllll8 trading of information initially with George C. Sawyer of the Silverton Nursery near Granite Falla, Washington, a Forest Service nursery that operated between 1909 and 1914. ( The study on Douglas-fir 11eed reported on in 1915 was the largest seed study ever done on the species. __....... . 1'y Willia and Hoffman, It covered seed yields 21 per tree, ber bushel, per cone, and per pound, and variation by tree age as well as by parent. Much of the data are •till applicable and useful. el was long sustained toward Research into nursezy practices under upgrading practices at Wind River Kursezy. This covered nursery design, cultivation, fertilization, bed design, 80Wing methods, seed holding, weed and pest control, root pruning, culling, grading and packing methods. These early decades of experiments and trials have undoubtedly merged indistinguishably into the technology of all modern forest nurseries. The only documentatio for much of this is in the publication, •Forest Planting in the Douglas-fir Region• by ICumme l, Rindt, and Meager in 1944. Planting and Seeding Studies The earliest seeding study was a trial of eastern white pine at Warren Gap in 1910, resulting in a complete failure. Some of the most imaginative work at Wind River resulted from the early finding that mice consumed virtually all the seed in sowing of cutovers. One activity by c. P. Willis in 1916 discusses his frustrations from studies of concealment, burying, and chemical protection of the seed. Despite coatings of such repulsive items as red lead, tar, belladonna, and mink urine the mice located and efficiently consumed the seed even under 2 inches of soil. Planting methodology was developed to considerable sophistication from Seedling size, time of planting, planting methods, planting planting studies. tools, care in transport, and storage methods were standardized. Some indication of the success of the methodology is provided by the 1915 and 1916 planting of the Douglas-fir Heredity Study. were planted as far away as Seedlings grown at Wind River Mt. Bebo and Mt. Baker with 12 separate plantings nearly all of which averaged over 90 percent first year survival. Much of this technology was reported in •Forest Planting in the Douglas-fir Region• by . . . --· _, -:--··.· - 22 1tU111111 el, Rindt, and Munger in 1944. However, like the nursery research, many of the trials and studies were undocumented. I Natural Regeneration Studies Large burns that were in every part of the Region when Wind River nursery began was the main reforestation concern of the period. Planting projects prior to world War II such as Yacolt, Cispua, Bebo, Breitenbush, Sanntiam, and Still Creek required a sustained production of over 2 million seedlings by 1914. Although natural regeneration usually followed the forest fires, planting had to be done on areas repeatedly burned. • A systematic look into the principles of natural regeneration was started by J. v. Hoffman. A transect of 4 milacre plots across Wind River Valley was established in 1918 to record every seedling from natural seeding. ( Wind River Valley was logged from a railroad which resulted in a cutover of the valley bottom and up each slope to the limit of skyline yarding. The transect of the plots at right angles to the valley permitted estimation of the effectiveness of seeding at various distances downslope frcm the two timber edges, one on a north and the other on a south slope. Within 5 years over 1, 000 natural seedlings per acre, mainly Douglas-fir, had reforested the north facing slope l/4 mile from the timber edge. The co=eaponding south slope was stocked with about 100 seedlings per acre. The conclusion, which has been the keystone of natural regeneration methods in the Northwest, is that on favorable slopes openings should be less than l/2 mile across, preferrably l/4 mile. These plots have been examined at intervals to date. The record constitutes the only long-term detailed sampling of natural regeneration ( through an entire rotation of Douglas-fir. However, the conclusions of the early years are now temperated with quite contrasting long-term results. valley is now entirely stocked to conifers except near the river. The The originally well-stocked plots within l/4 mile of timber edge are now a lower I 23 value stand so suppressed as to provide mainly small stems and many openings fran snow damage. The south-facing •lope, considered a failure for several • decades, has maintained a stocking of about 100 trees per acre. The south •lopes •tand is now of ccmmercial •ize and more valuable per acre than the overstocked north •lope, which for decades was considered a forestry Buccess. Now the most valuable stands are the ones on north slope more than 1/2 mile fran timber edge with normal volumes of com.mercial size stems. these were considered understocked. For decades Barly •tand density differences appear to be the main factor for present stand characteristics. • The experiment has not been reported formally since 1951. After 1918 Hoffman continued to •tudy the factors of successful regeneration, but wrongly concluded that enough seed was stored in the duff to regenerate a clearcut. The implication that no other seed source was needed, was enthusiastically received and continuous clearcutting was considered as a scientifically acceptable practice. Hoffman became a Professor of Silviculture at North carolina State University and outstanding leader in Southern forestry after World war I. Isaac studied natural regeneration after 1924 as one of his major interests. Bis approach was a ccaplete one involving detailed study of the physical factors of light, temperature, moisture, competition, and of dam.aging factors. He produced many •cientific papers over a long period which are •till regarded as the best information available on the topic for Douglas-fir. Bis •Reproductive Habits of Douglas-fir• published in 1943 9ummarized his life work in reforestation research. It was timely. The heavy cutting in the Region following World war II finally had a sound •ilvicultural body of knowledge. Both oregon and Washington •oon passed sound forestry laws 24 l_ base upon this work. on the national forests his •patch cutting• or •staggered setting• proposal became the standard cutting practice. Although Isaac had regeneration plots over the region, he used the Wind River Valley clearcut, and adjacent Yacolt burn as his ma.in study area. Several reburns had kept many openings in the Valley floor until the mid 1930's as convenient research sites. Virtually all of his plots involving instrumentation were near the headquarters site. • Bcoloqical Studies Thornton Munger's career was strongly slanted toward ecological studies. He observed the natural succession of species in all the timber types. Following Munger's suggestion, Judd, who had a. short career in research at Wind River, perhaps was the first who deduced the true role of forest fires as dominating the ecology of the Region. his later publications. Munger elaborated upon this theme in These papers had served as the basis of the intensive ecological studies today. The establishment of the natural area in the Trout Creek Division was only one of the efforts by Munger to provide for the long time study of ecology. He was the prime mover in starting other Experimental forests, each usually providing a natural area. He extended this thrust to formal setting aside of natural areas in each unique forest habitat, the start of today's extensive ayat- of natural areas. ( Silviculture Information needed to maximize productivity of established stands through harvest was addressed in several early studies. 25 P,ew studies in Douglas-fir have produced more persuasive impact on stand management practice than a apacing trial installed by Isaac in 1924. its design ahortccmings, such as Despite lack of replication over a range of site qualities, it has constituted a striking demonstration as well as producing auch data on the consequences of overdense early stocking. Anyone who has visited the area comes away convinced that there can be a severe value loss without control of tree density. The plot provides a fairly clear • demonstration that while gross volume of a closed stand has been nearly equal at the range of spacing frcm 4' x 41 to 12' x 12', there is an opportunity to distribute this wood to fewer large rather than many small stems, with perhaps little growth loss frcm underutilized apace. An unexpected outcome of the study was that height as well as diameter reductions occurred with increasing stand density. A clear indication of increasing snow damage with increased stand density was verified on a larger scale on the Wind River Transect Study. The study has been regularly reported on in recent years by Don Reukema and Robert eurtia. While other Wind River plots have not been as famous they have established important concepts. Considering the year 1919, a study of precommercial thinning of a 9-year-old stand would not have been expected. Its installation by Hoffman has provided an unusual insight of an inte%play of trends experienced over most of a rotation. in 1910 was thinned to 8' as x The dense reproduction following logging 81 density on two plots and a third left unthinned a control when the trees were about 6 feet tall. Thia single early thinning succeeded in shifting a large portion of aubsequent growth to produce larger stems. It has been the -in demonstration in the Region to pr wideapread practice of precommercial thinning. sophisticated question. te the The study addressed another Ia it better to thin to an exact 8' x 8 • spacing or ' 26 , to le.ave an equivalent naaber of dominant trees instead? it -s clearly best to leave the dalllinants. to exact spacing is the better. For about 30 years By DDW, however, the plot thinned The reason appears to he an indirect one • .Many of the smallest trees that fell on the exact s• x s• spacing did not develop, hence the plot for practical pu:<poses developed as if it were at wider spacing, even though it took many years for the good trees on the plot to express this advantage. once the crowns closed, Gross growth on the 3 plots appears about equal just as in the spacing study. • Walter Meyer installed a series of replicated thinning and spacing plots in 1934. Because of sto:i:m damage to sane of the series their results have been clouded, and their contribution to practice has been small. In the post world war I period a main thrust of the Station was inventory of the Region. A part of this effort .by Richard McArdle and Walter Meyer was the development of the yield tables for Douglas-f.ir. used were on the Wind River Experimental Forest. A portion of the plots The publication •The Yield of DOuglas-fir in the Pacific Northwest• in 1930 may have been the Station's most important contribution to forestry, since it provided estimates of volUllle growth rates for any DOuglas-fir forest. Studies of the improvement in stem quality frcm pruning have elucidated the major principle that pruning may improve growth if continued up the stem into the lover live whorls. Apparently the shaded branches in dense stands require enough photosynthate frCllll the upper crown to add detectable volUllles of I stem growth if removed in pruning. However, the ability to select the right ' \ tree to prune in a young stand is difficult. Studies of crop tree pruning reviewed two decade• later tlbowed only about 70 percent of the pruned trees were fated to aurvive caupetition. .. ·· - ·· . --... ..... - - .•. 27 Considerable insight -s gained by pathologists into the extent of stem rots detected on the basis of external indicators such as fruiting bodies and 11t• ...,.lling. scme of the early work by John s. Boyce vas done at Wind tiver, but his coverage -s regionvide of pathological problems of all western species. The use of these studies vas in estimating the llJllOunt of cull degrade in cruising or scaling. • Species Trials Forest species frcm eastern Ollited States were represented in early aovings at Wind tiver, presumably because such seed vas ca:mnercially available. Scme planting of each lot behind the Experiment Station headquarters was a common practice. River Arboretum. These formed the nucleus of the Wind Soon, the trial of forest tr••• frcm all parts of the world became a vigorous project under Thornton Munger. Meanwhile forest plantings of introduced species was having mixed results. Summer drought was usually too severe for species that came from regions like eastern United States vith ample summer rains. animal damage was also a major problem. Undoubtedly By 1924 the Wind River scientists had already concluded that introduced hardwoods were generally a failure even in the arboretum, and henceforth the emphasis of introduced trees would be confined to conifers. The main trials of introduced species after 1924 were of conifers at the Arboretum, or on Experimental Forests such as Cascade Head if Wind River climate vas considered too harsh. Arboretum record. Nearly 200 species trials are in the Initially a great interest vas generated by a clearly 28 •upera.or early growth of Buropean and Asiatic larches. Up to 20 years several exhibited a SO percent 1113periority over Douglas-fir in the Arboretum. l'Ortunately this enthusiaan 4id :ay age SO, all the introduced not carry over into many ccmmiercial trials. larches were fading, and any •uch ccamerical plantings might now be a 4i•aster. A few individual species, like Norway spruce, continued to compete with Douglas-fir through three or even four decades, but by now all are outstripped by Northwest conifers. • '?he Wind River experience with species introduction has thus provided two priceless lessons upon which a sound forest genetics program can be based. 11e droughts are more ••vere than most other major The first is that our SU111r commercial forest regions of the World whose trees, particularly hardwoods, are probably not adapted or useful. '?he •econd is that for each genera, our native Northwest conifers may start more slowly, but in their grand period of growth they outgrow their counterparts fran other telllp8rate zone forest regions of the world. Northwest species will probably be used on a quarter of the forest sites of western Europe for this reason. These two lessons haw permitted the region to confidently devote its limited efforts in forest genetics to native species. Provenance and Family Genetic Studies Although only two •tudies were installed at Wind River with a genetic -Uvation, both have had an impact far beyond the original study plans. were initiated by 'rhornton T. Munger. ( r Both 29 '1le Douglas-fir Heredity Study of 1912 -• unusual for .its time. Basically .it i• a study of 120 .individual parent trees, even though thirteen local.it.iea frcm which the parents originated are highlighted .in subsequent Racial studies -re aore usual elsewhere in the world. reports. But clearly the questions asked had to be approached with studies of .individual parent trees. Are diseased trees, young trees, suitable parentage .in reforestation? or trees growing on poor soils Although the seed crops of 1912 was • listed as moderate, the crews sent out by Munger to collect cones along the logging railways were successful .in gathering seed samples of individual trees covering moat of the Douglas-fir Region. known, like E. J. Hanzliak, c. Hames, which later became widely R. J:raebel, R. w. Weidmann, R. Ames, and C. P. Will.is -re involved in the collections. .involved, it .is small wonder that c. R. Tillotson, A. With such talent embellishments appeared .in the execution of the study that were ahead of their time. A mimeographed parent tree description fo:i:m recorded the location and there was a sketch of every parent tree plus the record of germination on its seed. As another .innovation, the study was replicated at a.ix locations, and over two planting seasons, a practice that -s not reinstituted until the 1930's. Every tree -s tagged with a c:uatcm made stamped aluminum tag which numbered its row and position. And a set of 12 long rows of 100 progeny per fllJllily, each a separate fllJllily, ran the length of the plantation specifically planned to assess with.in-plot a.ite variation. Unfortunately randcmization for statistical purposes was at.ill unheard of, so the study violates randcmness, the major assumption of modern statistical analysis. pants. Fisher -s still in Jenee 30 orpe study quickly demonstrated genetic differences at the racial level. It also quickly llhowed large family differences in the family rows of 100 trees, although the significance of the latter observation -s DOt highlighted until after 1950. 'l'hrough the period until world war II the viewpoint of each analysis was that certain seed sources, like Darrington, Washington, should serve as seed sources for general planting because of demonstrated superior growth rate on most of the five remaining planting sites. 'l'he sixth site was destroyed in a 1917 forest fire. • Eventually the overriding iJllpac t of survival differences began to suggest different principles in the intexpretation of results. High elevation races began to show superior survival, growth and stand forming characteristics at high elevation sites. Northernmost races began to shaw superior development at the northe rnmost site. The coastal race, which failed at four sites, showed excellent growth at the lower than sites. the test site. coastal site despite orginating from l,000' cascade seed sources •howed up best at the cascades A broad brush view of seed source specificity was evolving. Family differences within each race were always large, but began to dominate after the stand& closed weaker progeny. and Today any forester suppression reduced fami.ly numbers of can rank the parent performance without statistics by simply observing large differences in growth and survival -lking from one family row to the next in any of the five plots. The impact of this study on tree improvement in ( very great. the Northwest has been first practical progrUIB were started by Northwest landowners, they ccpied European progrllll.lB I In the l950's when the and southern United States These poorly fit Northwest conditions because plus tree selection ll nccmpatible of -s 41fficult, and Douglas-fir -s aong the most graft-i species. A cc:apletely different type of program -s proposed by Silen for J>Ouglas-fir. It -s based priaarily upon large scale testing to substitute for plus-tree selection, upon highly localized progrlllllB to fit theae study results, and seedling orchards to avoid grafting problems. . Heredity study -s over SO-years old at that The Douglas-fir The new program could be proposed with little risk since the use of wind-pollinated seed was already • known fran Northwest experience to provide an adequate test of parent trees. The Ponderosa Pine Regional stuc!Y established in l926 was based on a range-wide collection of seed fran 10 sources. This was outplanted at 6 locations both east and west of the Cascade Mountains. were alreac!Y large in the nursery, and in 1962 developed most of the findings. Racial differences have widened with time. A publication Inherent growth of ponderosa pine is best in southwest Oregon and northern California and drops off clinally eastward, northward, and southward in response to assured moisture availability in spring. Needles were shown to be longest in western sources and are inherently shorter in a clinal pattern that roughly follows the inherent growth pattern. 'l'he western races have 3 or more needles. east of the continental divide have 2 needles as a rule. Those Winter cold susceptibility is also clinal with west coastal races most s usceptible. There are also large inherent differences in damage by frost and animals. The ponderosa pine study has one plantation at Wind River. There is a local strain of ponderosa pine of which a few individuals are on of this plantation. the boundary Hone. of the 10 planted races approach the growth rate of the Douglas-fir surrounding the plantation. However, the ponderosa pin e native to the valley is actually taller than Douglas-fir of the same age. 1( l- 1 I 32 Afte the visitor views the problems with the 10 planted races, a visit to the nearby elUU!lples of local ponderosa pine provide a lasting demonstration of the illportance of correct seed source. I Like the clear demonistration of density effects in the nearby 1924 spacing study, the Wind River plot of the Ponderosa pine race study is a convincing genetic demonstration of genetic differences. The study, when canbined with similar data from a contemporary study at • Priest River, Idaho, has provided the main racial variation data for the .species. Monitoring Natural Areas The setting aside of 160 acres of natural area for future studies, and its ( increase to 1,100 acres soon after the Wind River Experimental Forest establishment emphasizing the desire to gain data and information on c!ynamics of natural stands. Measurements of growth and mortality of plots in the Natural Area established that in general old grovth stands were virtually static with mortality losses balancing or exceeding growth. In older stands this balance represented a shift from high value old growth Douglas-fir to lower value often decadent hemlocks, firs, and cedar. It was this known trend toward stand degeneration that became the focal point the controversy over selective logging as the official policy of Region 6 in the mid 1930's. Munger and Isaac saw in the practice a potential to degrade the Regions National Forests by hastening the succession to an all aged, low value forest heavy to cull trees. 'l'he outcome was establishment by Isaac of a large scale aeries of old growth yield plots on selectively logged National Forest lands. i ' World War II interrupted thi.s effort, but soon afterwards Isaac produced data to show that these concerns were real. Selective logging of old-growth Douglas-fir was then discontinued as the policy in Region 6. --- 33 l'OS'l' WORLD WAR l:I :in prospective, the proportion of total reaearch effort associated with Wind River Experimental Forest appear in this review. ter World war l:l: is much larger than it miqht The decades follovinq the var were one of the more active period& when one considers that such names as Steele, Eversole, Bullard, Stein, Jtrueger, Meaqher, Staebler, Tarrant, Williamson, wriqht, • Childs, Reukema, Curtis, Miller, and Silen then fowid their way into the literature. A fair portion of post war effort is alreaay reported for early studies that have been brouqht \IP to date in the previous section. The new study areas reported in this section cover second qrovth manaqement, harvest cuttinq, alder and fertilizer studies, seed and seedlinq research and general contributions. Second Growth Manaqement--A heavy emphasis of the Experiment station after 1947 was on second-qrowth naqement. An ad hoc reqional •second Growth Manaqement Committee, • sponsored by Philip Brieqleb to rapidly upqrade the informational base of second qrowth, drew heavily on Wind River apacinq and thinninq studies. Publication of the committee report entitled, "Manaqement of Second Growth Stands in the Douqlas-fir Reqion• stimulated research qenerally, and at Wind River led to several new studies. l:n 1952 a series of thinninq plots were established in the century-old stand on the Panther creek division. 'l'Vo objectives are listed, one economic and the other the biology involved with two levels of thinninq. in this stand proved to be slower than in younqer stands. decade there was decreased ,.rovth in both thinning levels. 'l'he responses For more than .a Results implied that removal of expected mortality could be recaamended, but removal of 34 appr,.::iable fractions of the stand at thia age required periods longer than a deeade before growth be9&11 to approach fo:i:mer levels. 'l'heir valuable experience has contributed to a more -ture view of second growth management. Spacing studies also received renewed interest. s. Meagher was established in 1951. competing trees in a 41-year old one study conceived by G. It involved removing O, 1, 2, and 3 stand frOlll around each of 10 designated crop • trees in dominant, codominant and inte:cnediate crown classes. Results after a decade -re clear and consistent in that greatest response was on dominant trees, and with largest number of COlllpetitors removed. However, early response was n on linear with much less response on codOlllinants or intermediates, than one. and a much greater response with two or 3 cOlllpetitors removed The study had wide implications for.young stands in showing requirements for a quick growth response frOlll thinning. A second spacing study was aimed to extend info:cnation beyond the 12' x 12' spacing of the 1924 study. Wider spacings were achieved by thinning a 23 year old plantation frOlll a density of 680 trees to densities of SO, 200, 250, spacing. 100, 150, and 350 trees, the narrowest corresponding to about 11' x 11' The initial result was unexpected. All thinned plots suffered •shock• to the extent that height growth after 5 years was reduced 46 to 69 percent compared to the unthinned stand. occurred. Sunscald and some mortality also Since then the plots have begun to respond to the thinning. AB intended, they are beginning to demonstrate tradeoffs in financial returns in the goal of concentrating growth on fewer larger stems. ! Thus the plots installed in this period round out the early work toward t producing a regional long-term experience with spacing control. 35 cuttings H Another development soon after the world War II was the Station'• research into harvesting of -ture stands. This research centered mainly at the Andrews and Cascade Head Experimental Forests. At Wind River acme B. J. timber sales -re made in the Trout Creek division to provide new age clases for Emphasis began to decline on this activity in the mid 1950's future studies. • • It revived again at Wind River around concepts concerning the conical extinct volcano of Trout creek Hill. The idea that this landmark might serve as an unusual stuey area was conceived by Dr. David M. Smith, Yale University. Smith spent a summer reviewing the Experiment Station Silvicul.tural research program. The suggestion, which appeared in his report, was to start a series of pie-shaped clearcuttings at each of the four quadrants to regenerate a series of new age classes for detailed silvicultural studies. The idea developed slowly as a cooperative thrust by District and Station personnel. The final plan, which called for establishing a new age class on each quadrant at 10 year intervals, has one installment completed. Red Alder Studies The potential role of red alder in-DOuglas-fir management has received primary stud,y at the Wind River Experimental Forest. This species is one of the -in natural sources of nitrogen in the soils of the region. A fortuitious pre-World war I planting of red alder proposed by E. s. Hanzlik as a firebreak strip on the Trout Creek Division provides a striking example of its effect on DOuglas-f ir growth. Isaac first noticed that seedlings within the strip showed llUCh better growth over those just outside. Tarrant related this increase growth to higher nitrogen levels in the soil. He vent on to assess annual contriblltion of nitrogen fraa red alder in these and other . ·.- ·. · - - - - - - 4.iAA.:• ,/ ( f I 36 aldex> atandS i n the Region. - '1'he finding by Station pathologists that alder also inhibited Phellinus weirii raised the prospect for an important role for the t1pecies in Douglas-fir 111anagement. I Fertilizer Study Effect of nitrogen fertilizer was tested in 1964 on a 35-year-old, Site chlorotic plantation on the Yacolt burn. v, Both diameter and height increased • from application of 200 to 600 lbs of ammonium nitrate per acre. in diameter vas nearly linear, and favored larger diameter trees. The response There was severe snowbreakage of codominant trees with highest application rates, however. The results of the Wind River experiment reinforced conclusions of other scientists that fertilizer application in low .site stands may favor better average stem diameter by encouraging suppression or breakage of lower crown classes. Another result has been to stimulate fertility studies to become a major interest of Olympia Laboratory scientists. Seed and Seedling Studies One stUdY at the Wind River nursery by :Krueger and Trappe contributed much to the concepts surrounding successful forest planting. 'l'Wo seasons of nursery growth were monitered for root and shoot growth as well as levels of food reserves. The finding that shoot and root growth alternated in surges thr0U9h the season contributed a more basic understanding of why planted trees often fail. For example, seedlings planted at the time of shoot elongation would produce few roots. ( 37 ..fa ' '· e cone ther •tudy by Silen U•e&•ed the effect on •eed of prematur _i harvest, and whether the 1-turity could be overcane by artificial ripening. Cone collections up to a JK>nth early could he successfully ripened in cool damp peat moss. However, placing them in a running stream, or in plastic bags proved to be lethal. The studies were carried to camnercial application. The practice of artificial ripening bas become widespread for most western conifer species. Large cooling facilities are incorporated into the Region 6 seed • processing system for purpose of artificially ripening of cones. The General contribution: With Wind River, the total contribution bas been far greater than the sum of its individual studies. simply been its continuity. Its greatest service to northwest forestry has The long term of most of its stdies has repeatedly demonstrated how initial results change over time. •seed Storage The in the Duff,• a seemingly clear early result, was upset a decade later. early notions of what constituted satisfactory restocking on the Wind River harvest study were almost totally at odds with the long-term results. The early notions that larches could outgrow Douglas-fir took half century to correct. What was considered desirable _spacing changed over a few decades frcm 6' x 6' to l2' x 12' or beyond as spacing studies matured. The emphasis for most of the first half century of the Douglas-fir Heredity Study was a •earch for better seed sources, rather than the more modern search for better parents within a local source. The early notions that selective cutting were pertinent for old-growth Douglas-fir required several decades to develop convincing data that best forestry required full sunlight. ' ' ' I 38 Wi.nd River 11erves as the prillle forestry example in the West of pricelessness of continuity of research. Yet t:he forces at work that diaamble such long-term efforts, both unplanne d and acmetimes deliberate, have been strong enough to auc:ceed almost everywhere else. Even at Wind River the question of •Why ahould we continue here?" has aerved to terminate portions and acmetimes threatened all of the continuity. But to those who take the trouble to visit, to study, and perhaps to add to the research, the answer always has come out •1ta too valuable to terminate.• or •sound long-term J>Xperience in forestry is too rare and precioua to give up." Because of the long-term findings, Wind River ia perhaps the single most lasting forestry experience to any visiting forestry newcomer. Where else would one find a complete documented stand history record from seedling to harvest size of trees, or find a demonstration of how superior are the native apecies over apecies from anywhere else in the world, or observe how abject most off-site plantings beccme in half a century, or find a rotation age demonstration of how Douglas-fir families, side by side, compare in growth and survival, or see how really poor most of the stems become in half a century when planting was too dense, or how few stems are really needed to stock an acre, or how over-mature atandS produce fiber at a declining rate, and also deteriorate in value. Such demonstrations tend to reinforce one another to give the newcomer, and even the experienced forester, an unusual insight into why forestry of the Oouglas-fir region is different from any other forest region on the planet. •