USDA FOREST SERVICE G ENERAL T ECHNICAL REPORT PNW- 58 1977 TINNING N YI LD 0 G I .. CONCEPTS AND SOME ESTIMATES· OBTAINED BY SIMULATiON DONALD L. REUKEMA DAVID BRUCE PACIFIC NORTHWEST FOREST AND RANGE EXPERIMENT STAT ION U.S. DEPARTMENT OF AGRICULTURE FOREST SERVICE PORTLAND. OREGON Effects of thinning on yield of Douglas-fir: Concepts and some estimates obtained by simulation Abstract Observation and research on yi lds of managed Douglas-fir stands have given us many bits of knowledge, but no unified picture. We assembled these fragmented bits of knowledge into a conceptual model of stand structure and development, which provides a basis for extrapolating results of studies to conditions which have not been directly observed. A stand simulator (DFIT) provides details on number and size of trees and distribution of yield over time. Details and operation of DFIT are described in a companion paper. This paper discusses concepts of stand development and effects of thinning, provides guidelines for thinning, and summarizes current expectations. We describe structure,mortality, and growth of natural stands, these attributes as a basis for projections. not been thinned, been an it would have and use We assume that had the stand "average" natural stand; also discuss effects of some variations around this average. but we The major source of gain from commerical thinning is harvest of merchantable trees otherwise lost to mortality. The two major sources of gains from precom­ mercial thinning are growth onto trees that will reach merchantable size growth. ( 1 ) concentration of and ( 2 ) improved height The latter increases with progressively poorer site·quality. We recommend that precommercial thinning be done when leave trees are 10 to 15 feet tall and 10 to of improved height growth, 15 years old. facilitate and minimize possible subsequent This will maximize the advantage selection of high-quality leave-trees, amage to trees, there are still worthwhile opportunities, If this stage has passed, but gains will be less, Average spacing of these leave trees depends on desired tree size at the first commercial entry; that are left, the larger the required size at that time, If first commercial entry will 8 inches in d.b,h., with a minimum merchantable precommercial thinning should 'leave d.b,h, the fewer trees when trees average of about inches, 6 about 400-well-spaced trees per acre. J The first commercial thinning stands, be made (or harvest) in precommercially thinned should be when average d.b,h. of trees reaches the intended size. To maximize merchantable yield of stands not precommercially thinned, first commercial thinning should be made trees begin to die. the about the time merchantable size Just as number of trees left after precommercial thinning depends on desired tree size at the next entry, so does the number of trees left after the last commercial thinning depend on desired tree size at final harvest, But rather than reducing the stand to this number of trees at a single commercial thinning, subsequent thinnings. Thinning interval, lesser extent, excess trees may be carried for removal at level of growing stock, type of thinning may be balanced and, to over a wide range of com­ a binations which will give the desired end product, Thinning shifts the trends of mean annual increment measure of volume, over time. Thus, relativ depend on the age at which comparisons are made. tion of m.a.i., at culmination, greater. greater, (m.a.i.), by any differences among treatments At ages prior to culmina­ relative gains from only commercial thinning are less than whereas those from precommercial thinning are a little After culmination, relative gains from commercial thinning are whereas those from precommercial thinning are a little less. The proportion of total usable production remaining for final hal'vest varies with thinning regime, but will likely be close to 70 percent of the total merchantable cubic volume produced; the other 30 percent being removed in thinnings. The proportion of board-foot production left for final harvest In stands receiving only commercial thinning, will be a little greater. this final harvest volume will likely be no more than about 85 percent of the volume that would have been harvested from the stand if it had not been thinned. The average d . b . h . o f trees cut at the f i nal h arve s t in t h i nned s t an d s wi l l l ikel y b e f rom about 5 t o 3 5 percen t l arger t h an t h a t o f aZZ merch an t ­ Cons i derin g ab l e trees cut a t f i n a l h arvest o f comparab le unt h i nned s t an ds . t h e s ame n umbep o f l argest t rees , however , t h o se cut at f inal h arves t o f st ands wh i ch rec e i ved o n l y commer c i a l t h i n n i n g wi l l b e about t h e s ame s ize as those in unth in ned s t a n d s ; in s t an ds t hat were a l s o p recommerc i a l l y t h inned , t hey wi l l be about 1 0 t o 1 5 percent l arger . I n creased growth o f res i dual trees i s o f f set b y remov a l i n t h i n n i n gs o f s ome l arger-t han - averag e t rees t h at wou l d o herw i s e be present at f in a l harves t . Gains in vo l ume produ c t i o n from o n l y commerc ial t h i nn i n g incpease wi t h i n c reas i n g s i t e qual i t y . The a d di t io n a l gain from precommerc i a l t h i nn i n g , however , is greates t on s i t e I V a n d decpeases with i n cre a s i n g s i t e qual i t y . Rel at i ve gains from commerc i al t h i n n i n g decrease when l e s s o f the t o t al vo l ume i s mercha n t ab l e; t h u s , t he g a i n s are less in t erms o f bo ard - f oot vo l ume t han as c ub i c vo l ume . The t ren d for gains from precommerc i a l t h i n ­ n in g i s jus t t he reverse-- greater i n t erms o f b o ard - f o o t vo l ume t h an as c ub i c vo l ume . Rel a t i ve g a i n s on d i f ferent s i t e q u a l i t ies a l s o vary w i t h t he measure o f v o l ume . KEYWORDS : Th i n n i n g ( t ree ) , t h in n in g ( precommerc i al ) , precommer c i a l t h in n i n g , t h i n n i n g ( co mmerc i a l ) , commer c i a l t h i n n i n g , growth ( f orest ) , i n cremen t , y ie l d ( forest ) , vo l ume i n cremen t , i n crement ( vo l ume ) , growin g s t o c k ( - i n crement / y iel d , mort al i t y , s t an d devel opmen t , s t an d s t r u c t ure , s t an d s ( even - aged ) , Dougl as - f ir , Ps e udots uga menziesii. CONTENTS P age INTRODUCTIOX . 1 NATURAL STAl-,TDS Descript ion o f A verage S t an ds St and Stru c t ure and Devel opment 10rt a l i t y . . . . . . . . Var i at ion Aroun d Average 1 2 4 6 6 SOURCES OF GAIN . . . . . Commercial Th i n n i n g Cap t ures Merchan t ab le Mo rt a l i t y Commer c i a l Th i n n i n g Impro ves Growth o f Res i dual Trees Commerc i al Th i n n i n g Reduces C ub i c V o lume Growt h Per Acre . Pre commer c i a l Th i nn i n g Concent rat es Growth on Mer chan t ab l e Tree s . . . . . . . . . . . . . Precommer c i al Thi n n i n g I mproves Hei gh t Growt h and S i te I n dex . . 7 7 8 8 THI 8 9 10 10 11 I NG GU I DEL I NE S . Precommer c i al Thi n n i n g . Commer c i a l Th i n n i n g Reg i me . RESVLTS OF THINNING . umber an d S i ze o f Trees . Vol ume Yield . . 15 15 20 EXTRAPOLATION TO OTHER STAND COND I T I ONS . Un i forml y Under s t o cked S t an ds Over s t o cked S t an ds . . . Non constant S i t e Index . I rregular S t an ds . 29 29 31 31 31 L I TERATURE CITED 31 APPEND I X . G l o s s ary . Vari ables and Rel at i o n s h i p s Examined . . . . S amp le DF I T P r i n t out . . 33 33 34 35 . . . Introduction I nt ens ive management o f commer ­ c i a l f o r e s t acreage provides many oppo rtun i t i es to increase the amount of u s ab l e wood produce d . To t ake advan t age o f these opportun i t i e s , we mus t e s t imate how s t ands wi l l deve lop under var ious man a gement regimes . The g r e a t e s t need i s f o r p r e d i c t ions wh i c h app l y t o t h e next generat ion : youn g s t ands t h at have not yet p a s s e d t h e p r e comme r c i a l t h in n i ng s t age and s t an d s t h at wi l l be c r e at e d wh en exis t in g s t ands are h arve s t e d . The s e a r e t he s t ands wi t h the gr e a t e s t oppor t u n i t y f o r i n t ens ive management and max imum product ion . A s e condary need i s t o help set man ageme n t r e g imes for t h e r ema ining l i f e o f current s t an ds wh i ch may be comme r c i a l l y The concern here is t o p r o ­ thinned . v i d e for e f f i c ient t r ans i t ion f rom t he current forest t o t h e new forest . Th e s e pract i ces wi l l , in some cases , be qui t e d i f f e rent f rom tho s e f o l l owed in s ub s equent forest rnt a t ion s . In b o t h cases , cho i c e o f anagement r e g imes mus t invo lve both economi c and s o c i a l cons i de r ation s . To e s t ima t e managed s t an d yie l ds , we f i rst assemb l e d our f r agment e d b i t s o f knowl edge i n t o a concept ual mode l , with as sumpt ions regarding s t ruc t ur e an d deve l opme n t o f natural st ands and t he resu l t an t e f f e c t s of t h in n i n g. This provi ded a basis for ext r apo l at i n g resul t s o f studies t o con d i t ions not d i r e c t l y observe d . Resul t s of s t u d i e s were balanced against deve l opment o f natural st ands , and against one ano t h e r , to derive r e l a t ionsh ips amon g var ious s t and p ar amet e r s and t o i n s ure cons i s t ency . The variables an d re lat ionships examin e d are l is t e d in t h e appen d i x . ll appendix f o r samp l e DF I T p r i n t ou t . W i t h i n t h e l im i t a t ions o f our mode l , e s t imat e s o f me rchan t ab l e y i e l ds ove r a r an g e o f s i t e qual i t y , merchan t ab i l ­ i t y l imi t s , a n d t hi n n in g r e gimes appear c o n s i st e n t a n d provide good e s t imat e s o f r e l a t ive g a i n s f rom t h in n in g . To t he e xt e nt t h at y i e l ds w i t hout t h i n ­ n i n g may d i f f er , so wi l l y i e l ds w i t h th inning . S i n c e no s t ands in t h e P ac i f i c No r t hwest have b e e n t h i n n e d re gu­ l ar l y for 60 t o 80 years , t h e y i e l ds e s t imat e d by t h e s imul a t o r are e xt r a p o ­ l at e d f a r b e y o n d t h e ava i l ab l e dat a base . The e x t r ap o l at i ons s e em r e a­ s o n ab l e t o t he aut hors a n d sev e r a l u s e r s , b ut t h i s i s no demo n s t r at ion of t h e i r val i d i t y . Th e growth r a t e s and o t h e r f actors a f f ect i n g y i e l d wi l l n e e d revision wh en r e s u l t s of a c urrent cooperat ive s t udy of mana g e d Dougl as - f i r y i e l ds are p ub l i s h e d . 1 Th e purp o s e o f t h i s p ap e r is t o c l ar i f y concepts r e gar d i n g s t an d We t r y t o s t r u c t u r e and devel opmen t . e xp l a in g a i n s f rom t hinn in g . W e f ir s t d e s cr ib e as sumed charact er i s t i cs o f natural s t an ds , s o urces o f g a i n f rom t h in n i n g , and e f f e c t s o f t h i n n i n g on From t h e s e t he s e s o ur c e s of gain . w e provide g u i de l ines f o r t h i nn in g . We t hen i l l u s t r a t e and d i s cu s s r e su l t s o f t h i n n in g , a s e s t ima t ed b y DF I T . We e s t imat e t r ee s ize an d vo l ume y i e l ds f o r s e l e c t e d s i t es, me r c h an t a­ b i l it y s t an dards , and t h in n i n g r e g imes ; we supp l ement t h e s e est imat es w i t h b r i e f d i s cu s s i o n s o f e f f e c t s o f a l t e r ­ n a t e t h i n n i n g r e g imes . Final l y , w e d i s cuss ext r ap o l at in g t o o t h er s t an d conditions . The p aper b y Bruce e t al . ( 1977) provides the i n f o rmat ion n e e de d t o u s e the s imul a t o r . L i t t l e o f t h e informat ion i n c l ud e d in t h i s report has been pub l ished . We then const r uc t e d a s imu l a t o r o f man aged s t and y i e lds ( DF I T--Dougl as­ f ir I nt e r im Tab l e s ) wh i ch e s t ima t e s exp e c t e d s t an d deve lopment and y i e l d , wit h i n spec i f ied l im i t a t ions about s t a n d charac t e r i s t i cs and t r e atment See r e g im e s ( Bruce e t a l . 1977). 1/ Sciences Supporting data on file a t Forestry Laboratory, Olympia, Washington. Natural Stands To prov i d e a b as e f o r evaluat i n g e f f e c t s o f t h inn i n g on s t an d dev e l o p ­ ment , w e des c r i b e t h e development o f " average" n at ur a l s t ands . A l l our / Some a s s umptions and resulting estimates may require revision when a coop­ erative study of managed stand yields is completed. This study is being conducted by Robert O. Curtis, Pacific Northwest Forest and Range Experiment Station, Portland, Oregon, and James Arney. Weyerhaeuser Company. Centralia. Washington. project i ons o f s t and devel opment ith t hinn i n g ar e made with t h e a s s umption. that , had the s t and not b e e n t h in ed , it wo uld h ave had these ave r a g e n atur a l st and at tribut e s . Some char ac­ t e r i s t i cs of aver age nat ura l s t ands are summar ized in the f o l l o w i n g dis ­ cus s i on and i l lustrat io n s . s l i gh t l y l ar g er on good s i t e s t han As t r e e s b e come on poor s i t e s . l ar ger (older ) , t he trend i s r ever s ed; f o r a g iven h e i gh t , D g i s sma l l er o n good s i t e s t h an o n poor s i t e s. . D E S C R IPTION O F A V E R A G E S TA N D S The eff e ct s of s i t e and age on ( Dg ) / and on number o f t r e e s and b as a l ar e a Der acre are s hown in f i gur e l . i/ Dg incr e a s e s w i th age. number o f t r e e s de cr e a s e s as Dg i n creases, and t h e r e sul t in g basal area p e r acre i n creases with age ; t h e b et t er the s i t e qual i t y , t h e mor rap id t he rat e o f chan g e . he re suZting r e l at ionship b e tween D g , numb e r , and b as al ar ea i s shown i n f i gure 2 . F o r l ack o f c l ear evidence to the contrary , we assume t h i s r e ­ l at i onship : 0 b e independ ent of s ite and age ; th i s de fines a s t and Th e den s i t y index (Reineke 1 9 3 3 ) . s t and s t a r t s out with several t hou­ s and t r e e seedl i n gs per a c r e . As t h e t r e e s increas e in s iz e , t h e sma l l e r t r e e s are crowded o u t o f t h e st and b y t h e ir mor e v i gorous n e i ghbors. By the t ime the D g of t r ee s reaches 24 inches ( 6 1 cm ) , o n l y about 100 t rees p e r acr ( 24 7 p e r ha ) r emain . The major e f fect o f s i t e i ndex is on the r te o f s t and deve l opment; t h e b e t t er the s i t e , the e a r l i er t h e age at wh i ch a given Dg i s r e ached . The r e l at i on s h i p b e t we e n h e i ght o f domi n an t t r e e s a n d o t h er st and at t r ibut es do es not appe a r to be independent of Si t e , h owever , as has o f t en b e en a ssumed (f i g . 3 ) . Our expe r i en ce has shown t hat Dg-­ and thus vo lume per acr e - - at a g iven dominant h e i gh t var ies w i t h s i t e index . Wh en t r ees ar e sma l l , D g correspondi n g t o a given h e i gh t i s I A s used throughout this paper, refers to average d.b.h.; i.e., a tree of mean basal area. il Dg 2 .§. w w 'r "I 170 20 a; ... 18 a: 16 a; 14 w ... Q iii "l 110 80 10 8 6 4 2 0 0 5000 140 10 20 SITE INDEX B 30 40 50 60 70 So 90 30 40 50 60 70 80 90 100 8R;:e N ___ 4500 w II II <Jl ... a: :; 4000 3500 3000 2500 2000 1500 1000 500 0 0 10 20 100 S . " " III ..J 140 110 2S0 ! U " a: 170 300 80 > f 200 150 100 <Jl SO 0 0 10 20 30 40 50 70 60 AGE (ye.rs) d.b.h. of Derived by reanalysis of data on which USDA Technical Bulletin 201 (McArdle et al. 1961) was based, plus additional data for young stands. 13 28 2B 24 80 90 100 Figure l.--Descriptors of natural stands by age and site: A, Dg of all number of trees per acre; per acre. C, trees; B, basal area ... NUMBER OF TREES PER ACRE F' gure 2 . --R e1 ationship aroong Dg, number of and basal area per acre i n natural trees, stands. 170 20 40 slo' , I, so , I , 100 o 140 I HEIGHT OF 00 MINANTS AND COOOMINANTS Figure 3 _ . - U:e ;:) SITE INDEX 100 I ' 150 zoo -Relationsh' p becween Dg and hei gnt ' of dom nant and codominant trees i n natural stands. 3 STAND STRUCTURE A N D D EVEL O P M ENT We have bro ught D g , n umber o f trees, and basal area t ogether in t o a single graph , a s i n f i gure 2 , t o f a c i l i tate examination and d i s cu s s i o n o f how stands deve l op over t ime and t he effect o f t h i n n i n g on that developmen t . We wi l l b uild upon t hi s f i gure in s u b s equent ill u s t ra ­ t io n s . In f i gure 2 t he number o f t rees decreases f rom l e f t t o ri ght so t hat t he t ime sequence of stan d development pro gres s es in t he c o n ven ­ t io n al manner , from left t o ri ght . The di agon a l l ines , connect i n g poin t s o f equal Dg , show t he mat hema t i cal rel at ionship amo n g t he t hree variables . For any comb inat ion o f two of t hese vari ables , t he correspond i n g third vari ab l e can be read for t ha t p o i n t on t he graph . There is a wi de range in s ize o f l i vi n g rees at an y t ime . Thus , the largest t hird of the trees i n a s t an d in c l udes about 55 percent o f the b a s a l area per acre and t he sma l le s t t hird i n c ludes o n l y about 15 percen t of t he t o t al basal area ( f i g . 4A ) . The l argest third is mo s t l y domi nant s a n d co dominan t s t hat are growin g wel l ; the sma l le s t third i s mo s t l y s uppres sed trees that h ave nearly Development o f t he s t opped growing . s t an d over t im e i s shown in f i gure 4 B , <OO c • 80 S· 70 ;:; • 0. 1J :; E ..J 0 ... i : I I I I I I I I I so 60 ..J !D A INTERMEDIATES " : SUPPRESSED 50 40 30 20 10 �������J-�J-J-J-J-�J-� o 10 20 30 40 50 60 70 80 90 100 TOTAL NUMBER OF TREES (cumulaced percent] Figure 4.--Structure and development of natural stands: A, Cumulated contribu tion of successively smaller trees to number and basal area per acre; B , successive curves of cumulated contribu tion at progressively older ages. 400 300 ...., j,) Q) Q) "- Q) c.. ro ::J IJ" IJ) '-' W IT U <:{ 200 100 80 so IT W D<:{ W IT <:{ ..J <:{ (f) <1: m 40 30 20 2.000 '.000 SOD 400 300 NUMBER 200 100 OF TREES 80 PER so ACRE 40 30 20 10 s hown b y a v e r t i c al l ine acro s s t hi s s h e af o f cur v e s ( fig. 5A ) . Thus , for examp l e , while Dg of t h e t o t a l s t an d incre a s e s from 3 . 0 t o 7 . 0 inc h e s ( 7 . 6 t o 1 7 . 8 cm ) - -wit h a lo s s of about 1 , 650 t r e e s p er ac r e ( 4 , 080 p e r h a ) --Dg of t h e 400 lar g e s t t r e e s p e r a c r e ( 9 90 p e r ha ) incre a s e s from about 4 . 3 to 8 . 0 in c h e s ( 10 . 9 t o 20 . 3 cm ) . where cu rves of t hi s cumulat ed con­ tribution of t r e e s at successively old e r ages are plo t t e d relative to t h e t o t a l numbe r of living t r ees . The six curve s p l o t t e d are for ages when the t o t al numb e rs of trees in t h e s t an d are 3 , 200 , 1 , 600 , 800, 400 , ?OO, and 100 , r es p e ctively . The 100-percent curve ( s t r aigh t line on l o garit hmic s ca l e ) is the s ame curve shown in figure 2; t his curve is t raditionally c a l l ed ORMAL in t h e Unit e d S t a t e s . The port ion of the s t an d wi thin each crown class is the s ame as in figure A. A s t h e s t an d grow s , t h e number of t r e e s lar ger t h an a s p e cified minimum d . b . h . inc r e a s e s fo r a time as more t r e e s grow to t h a t size ( fig . 58 ) . Eventually the smaller of t h e s e t r e e s b e come su p p r e s s e d and sub s e quent l y die . Thus , t h e r e Th e d e velopment of any numb er of lar g e s t t r e e s in t h e stand is w U q J: w 11 <! w [ q ill <:' ID :1 c 3C 2Cl A 1,000 8CXJ 3= 2.= III :; Figure 5.--Development of stand relative to cotal of largest trees; specified d. b. h. stand: B, omponents A, ."ixed number trees lar er than !! :11 600 4= 600 4= 300 200 ,= 200 6t/) UJ D q [ w 1 q w J: <! --' q iJJ <! ell 100 so so 5:) 40 1,000 soo NUMBER OF 3CJCJ TREES 2C)(J PER ACRE ,= is a maximum number of tTees that will reach any specified d.b.h. At the point where this maximum is reached, trees larger than this d.b.h. comprise about 63 percent of all trees in the stand and about 80 percent of the total basal area. We call this 63-percent curve MAXIMUM. Its importance will become more apparent in subsequent dis­ cussion. The portion of the cumu­ lated-stand curve which is above MAXIML represents suppressed trees. MO RTALITY An average natural stand loses about 1.5 percent of its trees for each 1 percent increase in Dg. Dg of trees which die each year is about two-thirds the Dg of the stand. We have divided this mor­ tality into two components--trees smaller than the specified minimum merchantable d.b.h. and trees larger. The latter comprise the part of mortality that can be sal­ ',,-aged or preven ted by commercial thinning. When the merchantable component of a natural stand reaches· 1AXIMUM (fig. 5B), many trees which are still alive have not yet attained the minimum merchantable size--and most of these submerchantable trees never will. ost of these sub­ merchantable trees are suppressed and will die sporadically over an extended period. On the average, the rate at which they die is a function of the rate of stand growth; the number of live trees smaller than any specified minimum d.b.h. is a fUnction of Dg of all trees in a natural stand (fig. 6). On good sites, most submerchantable trees will die before the time of final harvest. On poor sites, many of them will still be alive at final harvest. Volume of these small trees is similar on all sites; as a proportion of total volume produced during a rotation, such submerchantable volume is relatively minor on good sites but is substan­ tial on poor sites. Once the maximum number of trees larger than a specified d.b.h. is reached, merchantable trees in natural stands die at a rate which complements the submerchantable 6 ",7 . 3CO 20Cl 0 J cr !. w a: u <i a: w Q. <i W a: <i ..J <i II) <i ;n 100 90 60 50 40 1.0Cl0 900 Figure 600 6. --Remaining number of submerchantabl e and merc.':a.ntabl e trees in na t:.Jral stands when mini.:num merch.antable d.b.h. is 6.0 inches. mortality (fig. 6). This merchant­ able mortality in natural stands is related to declining tree vigor. In thinned stands. most of these trees should be cut before they die or, at least, they should be salvaged after they die. With more growing space, tbe life and growth of some trees may be extended. VA RIA TIO N A R O UND A V E R A G E I n reality, there i s wide varia­ tion around this average natural stand. On plots from which this average was derived, number of trees at a given Dg varied from about 65 to 150 percent of the average; variation was similar for all sites and for the range of Dg included. Other plots have been observed where number of trees with a given Dg have exceeded this average by more than 50 percent, but such extreme density is not common. The reasons for the variation around th€ average differ from one stand to another, and each has a different effect on stand structure and development. Eflects of some of these differences in s and structure will be alluded to .in the section regarding extra ola­ tion to other stand conditions. 440 Sources of Gain Thinnings provide wo s o u r c e s o f g a i n in us able y ield : t he harv e s t o f mer chantable r e e s whi ch have died or s oon would d i e and t he imp r o v ement o f gro w h ra e of t re e s t hat are l e f t . We f i rst di scuss g a ins ( and lo s s e s ) a s s o cia t ed w i t h commercial hinning-­ which is commonly a p a r t of r e g imes " b e g inning wit h p r e commer c i al t hi nning . Q/ We t hen discuss ga ins as s o c i at ed w i h precomme r c i al t hi nning--which we consider dded g a ins r elat i v e t o commercial hinning alone . M E R CHANTA B L E M O R TA LI T Y The maj or source of gain f rom comme r c i al t hinning i s har ve s t o f me r chantable r e e s t ha t would o t he rwi se be lo s t t o mo rt ali t y . The larger t he s i z e to whi ch t r e e s are grown , the g r e a t e r i s t h e los s t o mort ali t y ( f i gs. 5 , 6 , and 7A ) . Thus, loss o f mer chant able produc­ t ion without commer c i al t hinning- ­ and po ent i al gain f rom comme r c i al t hinning- - i s much g r e a t e r on good I f com­ s i t e s han on poor s i t e s . mer c i al t hinning i s st art ed e arly enough , virt ually all mer chant able mo r t ali t y can b e c ap t ur ed . When thinning is dela y ed , part of t he me r chantable mo rt al i t y occurr ing prior t o thinning will b e lo s t ; but p art can be s alvaged when t h e s t and is thinned . Likewise, when an unt h i nned st and is harve s t ed , some r e c ent merchantable mor t al i t y c an be s al­ vage d . Thus, mer chant able y i eld har v e s t ed from a nat u r al s t and i s a l i t tle gre a t e r t h an net me r chant ­ able y ield , and t h e gain f ro m thinning is a l i t t le le s s t han t h e di f f erence b e t w e e n t o t al me r c h ant ­ able production in t he t hinned st and and net me r chant able y i eld in t he unt hinned . / If trees are spaced widely 360 320 280 240 200 MERCHANTABLE NET YIELD IN UHTHINNED STANO, "".5 6.0+ /fiches In d.b.h. 160 121J J i C O M M E R CIAL THINNING C A PTU R E S A 400 §, " • 80 40 ���-L���-L��LO ��-L� 0 2 6 a to t2 14 16 18 440 B 400 360 320 280 240 200 MERCHANTABLE NET YIELD flees 6.0+ IfiChas 1(1 d.G.h. 160 IN UNTHINNED STANO, 120 80 40 00 OF UNTHNI">.IED STAN:) [JI"CI"oeaj Figure 7.--Sources of gain from thinning, relative to normal stand Dg: A, Capture of merchantable mortality with commercial thinnings B, concentration of growth onto merchantable trees with precommercial thinning. enough, precommercially thinned stands may be carr ied to final harvest with no commer­ cial thinning. 7 C O M M E R C IA L THIN N IN G IM P R O V E S LOO GRO WTH O F R ESID UA L T R E E S Comme r c i a l t h in n i n g al so impro ves growth rat e s o f i n d i v i dual The g a i n in s i z e r e s idual t r e e s . o f t h e s e t r e e s , howeve r , usua l l y d o e s not l e a d t o a sub s t an t i a l i n c r e a s e in s iz e o f t r e e s at t ime of f inal h a r ve s t . Thi s gain is l ar g e l y o f f s et b y removal of many l ar ge r - t h an - av e r age t r ee s in t h in ­ n in g s ( Reukema 1 9 72 , Reukema and P i en aar 1 9 7 3 ) . C O M M E R C IA L TH IN N IN G R E D U C ES C U B I C V O LU M E G R O WTH P E R A C R E Tot al cub i c vo l ume growth p e r a c r e i s reduc ed b y mo s t t hi n n i n g s b e cause t r ees do n o t comp l e t e l y o c cupy t h e area a f t e r t h i n n i n g . A c t ual t r e n d s o f growt h over t ime wi l l vary w i t h t h e t h i n n i n g r e g ime; gr eat e s t r educ t ion s , r e l a t i ve t o p o t e nt i a l , wi l l b e imme d i at e l y a f t er e ac h t h inn in g . W i t h i n f a i r l y w i de l im i t s , however, t h in n i n g r e g ime appe a r s t o have l it t l e e f f e c t on t h e average p e r c e n t age r e du c t ion in growth over an extended p e r i o d ( Reukema 1 9 72 ) . I f r e s i dual s t o ck ­ i n g i s carr i e d at a h i gh e r l e ve l , growth may b e r e du c e d a l i t t l e l e ss; if at a lower l ev e l , growt h may be r e du ce d even mo r e . Th i s d e p e n ds , howe ve r , not o n l y on s t o c k i n g l e v e l b u t a l s o on whioh trees ar e cut . For l ack o f b e t t er i n f o rmat ion , we e x p r e s s this r edu ct ion in growth as a fun c t ion of s t and age at t im e o f f i r s t c omme r c i al t h i nn in g ; t h e o l d e r t h e s t an d wh en t h inn i n gs a r e b e gun , t h e gr eat e r t h e r e l at iv e r e duction in g r o w t h ( f i g . 8 ) . Th e ab i l i t y o f t r e e s t o respond t o r e l e a s e i s r e l at e d t o t h e i r v i go r wh i c h , i n turn , i s l o o s e l y corr el at ed w i t h age . At a given age , t r e e s on good s i t e s have r e l a t i v e l y s h o r t e r crowns t han t h o s e on poor s i t e s , but t h e t re e s o n g o o d s i t e s a dd t o t he s i ze o f t h e i r crown s more r ap i dl y . 8 88 86 a a: .J z UJ fa a. 84 82 8 ua 88 fz UJ u a: 86 UJ a. 84 82 80 w---L---L---L-�L-�--�--�--� 20 2S 30 3S 40 4S 50 55 80 65 AGE AT FlRST COMMERCIAL THINNING iyears) Figure 8.--Rela tive reduction in growth between first commercial thinning and final harvest, b y age a t firs t commer­ cial thinning. P R E C O M M E R C IAL THIN N IN G C O N C E N TR A T E S G R O WTH O N M E R CHA N TA B LE TR E E S Precomme r c i al t h i n n i n g i n c r e a s e s growth b y e l im i n a t i n g exce s s compe t i ­ t i on a n d c o n c e n t r at i n g a l l growth o n t re e s t h a t wi l l b e come m e r c h an t ab l e ( f i g . 7B ) . I n a n at ur a l s t a n d , as crop t r e e s i n c r ea s e i n s i ze t h e y gradua l l y crowd out t h e i r weaker n e i ghbor s . Crop t r e e s do not com­ p e t e s t ron gl y w i t h one another unt i l t h e i r Dg app r o aches MAX IMUM ( f i g . 5B ) ; howeve r , a sub s t an t i a l p ar t o f t h e t o t al s t an d p r o duct ion h a s b e e n on t r e e s t h at n e v e r r e ac h merchan t ab l e s i ze , resul t i n g i n r e du c e d growth on t r e e s t h at do . In a p r e c ommer ­ c i al l y t h i n n e d s t and , o n l y c r o p t r e e s ar e l ef t . The s e t r e e s do not comp e t e st ro n g l y wi t h o n e ano t h er unt i l t h e i r Dg appro ach e s MAX IMUM, Rnd t h e y reach t h i s s i z e sooner be cause o f t h e e l iminat ion o f addi t i o n a l com­ p e t i t ion ; the e ar l i e r the p r e commer­ c i a l t h in n in g , t h e gr8ater the gain . . If two comparable stands are preco ercially thinned to different spacings, trees grown at the wider spacing will eventually reach a For a while, trees in larger size. both stands will grow at about the same rate; but as the trees increase in size, competition retards growth sooner in the stand with more trees. For example, by the time a stand with 400 trees per acre ( 9 9 0 per ha) reaches the MAXIMUM stocking level, trees in a stand with only 29 0 trees per acre ( 715 per ha) will be somewhat larger (fig. 9 ) . --.---- .- • .• -r ";". - ...-- > •••••• __ .,- , '' ..... ... . .... ,. T" '-" - ."""" . • - - '. -,"-- !".'r-.,;,' '; " '• trees per acre, the period to stand closure is longer on poor sites than on good sites; thus, relative improvement in height growth is greater on poor sites than on good sites. Likewise, the period to stand closure depends on stand density; the fewer the residual z Q f- 1 a 1= A ,10 .... « ,= t!J f- ill I so 80 « J z Z 70 <I. .J « .J a: 0 so Z 2 sec 40C 300 "JUMBEq Figure 2CC ::= 9.--Effect 100 TPEES PER SO 80 ACHe of spacing of attained d.b.h. and on basal 50 40 trees on area per ac=-e. u. 0 .... Z w (.J a: w il. 50 8 10 P R E C O MM E R CIAL THIN N I N G IMP R O V E S 12 14 16 18 20 AGE 22 24 {yeersl 26 28 3D 32 34 36 HE IGHT G R O WTH A N D SITE IN D E X Precommercial thinning also improves height growth. Substantial improvement is common on poor sites (Reukema 1970 ) : apparent short-term improvement has been observed on better-than-average sites. Spacing apparently has little effect during the period when rate of height growth is rapidly increas­ ing; however, in stands in which the crown canopy has not yet closed, this period of rapid height growth is prolonged. Thus, height growth in open stands culminates at a later age and at a higher rate than that in the average natural stand (fig. lOA). 'Iii th a given number of HEIGHT OF OOMINANTS ANO COOOMINANTS Figure lO.--Gains with precommercial A, [teet] thinning: Trends over time in height growth of dominant and codominant trees; B, ment in site index. 9 . ".,-,-- improve­ timing o f th i n n i n g , average spac i n g o f l eave tre e s , and s e l e ct i o n of l eave tr ees. To h e l p a manager s e l e ct des i rab l e th i n n i n g r e gimes , we d i s c u s s each o f th e s e factor s . We do not try to de f i n e a " b e st" th i n n i n g r e g im e , nor do we d i s c u s s al l factor s whi ch t h e manager should c o n s ider i n determi n in g what i s b es t f o r him . trees , the longer th e p e r iod . CUl ­ mination o f h e i gh t growth wi l l not b e d e l a y ed inde f i n it e l y on open­ g rown tr e e s , so oth e r f actors must be l imiting at wide spacings . By the time th e stand c l o s e s , h e i ght growth w i l l h ave dec l ined to th e n o rmal pepoentage rate and w i l l f o l l ow the n ormal h e i ght growth patt ern th ere afte r . Th is has the e f f e ct o f improv i n g s ite index up to th e t ime o f stand closur e , and th e r e a f t e r of maint a i n i n g it at the h i gh e r l ev e l ( f i g . l OB ) . Th i s improvement i n s it e index i s n e gl i­ g i b l e on good s i t e s b ut very s ub­ stan i al on poor s i tes , wh e r e th e stand may be r e l at i v e l y open f o r many year s . A f t e r stand c l o s ur e , the potent ial rate o f cub i c vo l um e gr owth p e r acre appears t o b e about th e normal rate f o r current s ite index and age , r e gardl ess o f wh eth e r or n o t a stand w a s p r e commercial l y th inned ( f i g . 1 1 ) . h us , the inherent p roducti vi ty of the s ite e x ce eds its capac ity to produce under norma l l y den s e stand condi­ tions . P R E C O M M E RCIAL THINNING On l y a f ew h i gh l i ghts regar d i n g p r e commerc ial t h i n n i n g w i l l b e s um ­ mar i zed h er e . We suggest that readers r e vi ew " Guide l in e s f o r p r e c omme r c i a l th i n n i n g o f Douglas - f ir" ( Re uk ema 1 9 75 ) f o r mo r e detai l . We r ecommend that p r e commer c ial th i n n i n g b e do n e whe n l e ave tr e e s are about 10 to 1 5 y ears o l d and 10 to l 5 f eet ( 3 to 5 m) tal l ( f i g . 1 2 ) . Thi s wi l l maximize the advantage o f imp r o ved 35 30- 30 28 t=2.8 ­ 24 r::.. 22 t- 25 20r :. s Poorest site E- t ., ; 20 12 I­ :.0 C8­ st: [ !:!:. l­ X 88 !.OO 120 INITIAL SITE INDEX 140 :'80 (100-year-ba",) 200 Fi gure 11. --Relative improvement in si t e index and current gross cubic vol ume growth wi t h precommercial thinning. Thinning Guidelines The e f f e cts o f th inning on stand deve l opment depend on th e p ar t i c u l ar th inn i n g r e g ime f o l l owed . For both p r e commerc ial and comme r c i a l th i n n in g , maj o r cons ide rations are: 10 UJ X 10 ______ _ 5 20 25 TOTAL AGE (Years) Fi gure l 2 . --Recommended time for precommercial thinning, rela tive to height and age of l eave trees. h e i gh t g rowt h ( f i g . 10), f a c i l i t at e s e l e c t ion o f h i gh - qu a l i t y l e av e t r ee s , and min i m i z e p o s s ib l e sub s e qu e n t damage t o t r e e s . I f t h i s i d e al s t a g e h a s p as s e d , t he r e a r e s t i l l wor t hwh i l e oppor tunit i e s f o r p r e commer c i a l t h i n ­ n in g ; but g a i n s wil l b e l e s s . The average s p a c in g o f l eave t re e s d e p e n ds on how l a r ge t h e t r e e s are expe c t e d t o b e wh e n t h e f ir s t commerc i a l t h inn i n g ( o r h a r v e s t ) i s made; t h e l ar g e r t he t r e e s a t t h a t t ime , t h e f ewer t h e t r e e s t h a t are l e ft . The numb e r of t re e s to b e grown t o a s p e c i f i e d s iz e i n a p r e ­ comme r c i al l y t h i n n e d s t an d i s about equal to the maximum numb e r t h at reach t hat s iz e i n t h e a ve r a g e n at u r a l s t an d ( f i g . 5B ) . With the spacing c o r r e s p o n d ing t o t h i s numb e r , a l l t re e s a r e e xp e c t e d t o r e a c h m e r ch an t ­ ab l e s i ze . When t h e d e s i re d t re e s i ze i s r e a c h e d , mos t t re e s w i l l s t i l l b e growi n g we l l a n d wi l l r e sp o n d we l l t o r e l e as e . Pro p e r s p a c i n g i s m o s t c r i t i c a l on p o o r - qua l i t y s i t e s . P r e c omm e r c i a l t h in n i n g at t h e r e commen d e d t ime s h o u l d l e av e t r e e s o f f a ir l y un i fo rm s i ze . I f p r e com­ mer c i a l t h in n i n g is de l a y e d a p p r e ­ c i ab l y , it m a y b e n e ce s s a r y t o l e ave a wider range o f s i z e s . Larger t r e e s c l o s e t o mer ch an t ab l e s i z e shou l d g e n er a l l y b e r e t a in ed i n t h e s t an d un t i l t h e f i rs t comm e r c i a l t h i n n ing. C O M M E R CIA L TH I N N I N G R E G IM E Jus t as t h e num b e r o f t r e e s l e f t a f t er p r e c omme r c i a l t h i n n i n g d e p e n ds o n d e s i r e d t r e e s iz e a t t h e next e nt r y , s o d o e s t h e number of t r ee s a f t e r t h e Zast commer c i al t h inn i n g d e pe n d on d e s i r ed t re e s i z e a t f in a l h a r ve s t . But r a t he r t h an r e du c i n g t h e s t an d t o t h e d e t ermin e d numbe r o f t r e e s a t a s i n g l e commer c i al t h in n in g , e x c e s s t r e e s may b e l e f t f o r r emov a l a t sub s e quent t h in n i n g s . Th i n n i n g i n t e rval , l ev e l o f g r o w i n g s t o c k , and , t o a l e s s er e x t en t , t ype o f t h inn ing may b e b a l an c e d o v e r a w i d e range o f comb i n a t i o n s wh i c h wi l l g i v e t h e d e s i r e d e n d p r oduct . Cho i c e o f commer c i a l t h i n n i n g r e gime , w i t h i n wi d e l im i t s , h a s l i t t l e e f f e ct on t o t a l p rodu c t ion ; i t d o e s a f f e ct t im i n g o f y i e l d and t r ee s iz e d i s t r ib u t i on. De s ir ed t im i n g o f t h i n n i n g s , r e s i du a l s t o c k i n g l ev e l , a n d t y p e o f t h i n n i n g a r e so int e r r e l at ed t hat d i s cus s in g o n e w i thout t h e o t h e r s i s d i f f i cu l t . S in c e t h e y cannot a l l b e d i s cus s e d a t o n c e , we wi l l r e l at e e ach , i n d i v i dual l y , t o a s t o c k i n g gu i de ( f i g . 1 3 ) . u w 2300 • L J IT 200 5 150 w <! [ W n. <! 100 W [ <! 80 .J <! UJ <! 60 OJ 800 400 300 NUMBER 200 OF 150 TREES PER 100 80 60 ACRE Fi gure l3 . --A thinning regim e maintaining stocking level between RECOMMENDED RESIDUAL and MAXIMUM / wi th maximum interval between thinnings . AXIMUM, as de f in e d p r e v ious l y , i s t h e appro x im a t e maximum s t o ck i n g l e v e l t o wh i ch a g i ven number o f m e r c h an t ab l e t r e e s should b e grown in a man a g e d s t a n d . RECOMMENDED RES I DUAL i s t h e m in imum s t o c k i n g l e ve l t o wh i c h t he me r c h an t ab l e p or t i on o f t h e s t an d wi l l b e r e du c e d at e ach t h inn i n g ; f o r t h i s i l l us­ t r at i on , RECOMMENDED RES IDUAL is s e t at a b a s a l a r e a l ev e l o f about 7 5 square f e e t p e r a c r e ( 17 m 2 / h a ) b e l ow MAXIMUM . Tim in g of Thinn i n g F i r s t Th inn i n g T o maximize us ab l e cub i c vo lume y i e l d , one shou l d make t h e f i r s t t h in­ n i n g at about t h e t ime m e r c h an t ab l e­ s i ze t r e e s b e g in to die . There f o r e , 11 f i r s t pos s ib l e t h i n n i n g is s e t at t h e a g e when t h e mer chan t ab l e port ion o f t h e s tand reaches MAXIMUM . For st ands r e c eiving only comm e r c ial t hinn ing , t h i s age is a fun c t i o n o f s i t e index and s pe c i f i e d min imum me r chan t ab l e d.b.h. For s t an ds wh i ch have b e en p r e comme r c i a l l y t h in n e d , t h i s age i s also a func t i o n of age when the s t and was pre comme r c ial l y t h inned t o a spe c i f i ed numb e r of t r ees; t h e ear l i er t hat s t o c k i n g dens i t y is cont r o l l e d, the younger t h e age o f t re e s a t f i r s t comm e r c ial t h in n in g ( f i g . 14 ) . ;::::::'-' ::0 40 30 :: o 80 A ! ! 90 The f ir s t t h in n i n g o f s t an ds t h a t are only comme r c i al l y t h i n n e d may b e de l ayed b e y o n d t h i s f ir s t p os s ib l e age f o r t h i n n i n g . Thi s may b e e i ther a m i s s ed oppor t un i t y or t h e r e s ul t o f a man a gement ob j e c t ive other t han maximizin g t o t al usab l e y i e l d . I f t hi n n i n g i s delaye d , mer chan t ab l e t r e e s wi l l d i e out o f the s tan d at t h e natural s t and rat e ( f i g . 5 ) . Th e first com­ mercial t h innin g of prec ommercia Z Zy t h inned s t ands shoul d not be de layed app rec iab l y b e yon d the f ir s t p o s s ib l e t h inn i n g age . Th e s e s t an ds are mo r e un i f orm t han natural s tands a n d are not l ik e l y t o t h in t h emse l ve s e f f e c ­ t i ve l y; espec ia l l y ori poorer s i t e s . I n s t ead , t hey may r e t a in an excess i ve numb e r o f trees f o r a pro longed period an d t e nd to s t a gnat e; e ventu­ a l l y , t h e r e l ik e l y w i l l be heavy mor t al i t y ( f i g . 1 5 ) . I . 100 ! I 120 ! ! I 140 ! I I 160 180 160 180 ! ! ! ! 80 70 30 50 40 30 20 10 0 80 B 100 120 140 I I INITIAL SITE INDEX (10Dwyeal"'-baSls1 I I 200 Figure l 4 . --Age at first possibl e commercial thinning, by ini tial si te index and mini­ mum merchantable d . b . h . (dm) . A, when dm is 6 . 0 inches i B I when dm is 7 . 5 inches. Las t Th inn ing 300 Tim i n g o f t h e last t h i n n i n g s houl d depend o n t h e d e s i r e d a g e an d s t an d charact e r i st i c s a t f inal harvest . General l y , it wou l d appear des irab l e to leave as much vo lume . as prac t i cal for t h e f inal harvest-­ rat h e r t han remo v i n g mor e t r e e s than n e c e s s ar y in th i n n i n g s . A r easonab l e ob j e c t ive wou ld b e t o have t h e s t an d a t MAX IMUM s t oc k i n g l e v e l a t t ime O n c e t h e s t an d of f i nal harvest . h a s b e en r e duced t o t h e n umbe r o f t r e e s that are e xp e c t ed t o r e a c h MAX I MUM l e v e l a t a des i r e d age , ano t h e r t h inn ing i s unn e ce s s ar y . :; 2' • 200 c • J cr .J!!. 150 w II U " II w a. " w II " -' " !]) " III 100 80 60 50 40 Th inn i n g I n t erval I f s t o ck ing i s ma i n t a i n e d b e tween o u r RECOMMENDED RES IDuAL 1.000 800 600 500 400 NUMBER OF 300 TREES 200 PER 150 100 ACRE Fi gure l5.--Effect of delaying commercial thinning of precollvllerciall y thinned s tands . 12 I 200 __ , . . _ 1._.. • \ • . ,.... " _!Jo'. a n d MAXIMUM levels (fi g . 1 3 ) , t h e int erval between t h i nn i n g s i s t h e t ime required t o grow about 7 5 s qu are feet per acre ( 17 m 2 per h al i n b a s al area. I f a s t an d is t hinned more f requen t ly , it will not b uild u p t o t h e MAXI MUM level , a n d each t h i n n i n g w i ll remove les s volume . I f t h i n n i n g s are. made less f requen t ly , the g r ow i n g st ock level at t a ined before t h e next t h i n n i n g I f gr0w i n g will exceed t h e MAX IMUM . s t o ck i s allowed t o exceed t h i s level b y ver y much , i n d i v i du al t ree growt h may be severely ret a r ded . l--Cornnel"aiaZ thinning ages oO'lTBsparuiing t<' suggested heigr.::­ g!'O'lJth 'intervaZ and adjusted fol' aompar;ibi Zir;y /Jith l'ota::-::.7. aqeJ by treatment and mininn.on merchanr;able d.b.h. Table maxim u m Th i s maximum i n t er val can be clo sely app r o x imated b y f i xed h e i ght g rowth i n t ervals , where the i n t er val var ies b y s i te qual i t y - - f rom about 2 4 feet ( 7 . 3 m) on s i t e I to 15 feet ( 4 . 6 m ) on s i te V ( f i g . 1 6 ) . The c o r respon d i n g t ime i n t er v al between t h i n n i n g s var ies sub s t an t i ally w i t h s i te q u ality a n d s t an d a ge- - f rom about 6 years in youn g s ite I s t an ds to more t h an 30 years in s i t e V s t an d s ( t able 1 ) . Thi s i n t erval c an be mo d i f i ed t o make it comp at ible with desi red t im i n g o f the last t h in ­ n in g ; t h e r i ght -hand p o r t ion o f t able 1 shows an example o f such a mo d i f i c at ion . With suggested ; nches J 6.0 7.5 . inches inches Adjusted for compatibil ity interval + PCTY CTli alone 6.0 oH 25 32 40 51 65 (85) 30 38 48 61 (eo) 11 22 28 36 45 57 (74) 30 38 49 64 (86) 36 46 60 (80) 26 33 43 55 (72) I 6.0 inches inches Site l CT alone CT 7.5 I (85 PCT 7.5 6.0 J incoes '.5 ; nches inches 30 37 44 53 65 22 28 34 42 52 65 36 44 54 67 26 33 42 52 67 45 55 68 31 40 52 69 : .. 5': 6S 59 65 38 51 69 ':; 5 5? years)lI 25 32 40 51 65 24 31 38 48 62 (81) + C 2! J: JS 5D : Site II (90 years)lI 29 37 47 61 (82) III (95 Site 36 47 63 (86) 45 60 (82) 31 40 52 (70) 47 65 (95) 59 (84 ) 38 51 69 68 (110) (89) 51 (74) IV (100 43 58 (80) Site V (105 59 (87) J5 -- 2; 53 56 years)lI 35 46 60 (81) Site Jj y 1I 11 30 39 51 67 36 49 68 years)lI 47 66 years)lI 68 59 51 70 Conmereial thinning. Preeo"",ereia 1 thinning. Site class (rotation age). Numbers in parentheses are ages at completion of next thinning period. .IH Res i du a l Stock i n g Level (Average Spacing) Figure l 6.--Hei ght growth i n terval which will maintain s tocking l e 7el approximatel y between R ECOMMENDED RESIDUAL and MAXIMUM , by si te index. At t h e RECOMMENDED RES IDUAL s t o ck i n g level , b as al area r an ges f rom b o ut 1 0 5 s q u are feet per acre ( 24 m per hal f o r la- i n ch ( 2 5 - cm ) t rees t o 2 2 5 square feet ( 5 1 m 2 ) f o r 3 0 - i n c h ( 76 cm ) t rees ; corres­ pon d i n g aver age s p a c i n g s of t rees are about 1 5 an d 3 1 feet ( 4 . 6 an d 9 . 4 m) . We f u r t her recommen d t hat no more t han one- t h i r d of t h e mer ­ ch an t able basal a r e a b e r emo ved in a t h i nn i n g . I t would be reason able t o set res i dual s t o c k i n g at ot her levels for cer t ain s i t uat ions . For example, i f the t ime req u ired f o r trees to grow fr om RECOMME DED R E S I DUAL to MAX I MUM ( f i g . 1 3 ) i s lon ger t h an the t h i n n i n g 13 i n t e rval d es ired f or a part icular s t and , res i dual s t ocking could be s e t at a h i gher level compat ib le wi th the short er thi n n i n g i n t erval ( f i g . 1 7A ) . On the o t her han d, s t ocking could b e mai n t ain e d a t a lower average level (f i g . 1 7B ) . I n some ins t an c es , i t would b e reasonab le t o t h i n a s t and t o a s t o ck i n g l e v el b elow the RECOM­ MENDED l evel--w i t h a corresponding redu ct ion i n grow h p e r acre . 300 " .." Type o f Thinning (Selection of Leave Trees) Type o f t h i n n i n g is d e f i n e d by t h e diD rat io , whi ch is Dg of cut t rees Cd ) relat i ve t o Dg of merchan t ­ able t rees b e f ore t h i n n i n g CD ) . A re aso n able trend o ver t ime can b e e s t ablished b y relat i n g d iD rat i o to Dg o f merch ant able t rees (D ) and the minimum merchant able d . b . h . ( dm ) (f i g . 1 8 ) . We sugges t that d i D at t he f irs poss ible thinn ng should be be tween 1 . 0 and 0 . 9 . §/ When dm is 0 . 4 D or less, d iD should be about 0.8. lCo 80 6- w [ U 60 [ w 0. [ 300 -1 In 200 Figure l 8 . --Regimes of diD ratio rela tive to mimimum merchan table d . b.h . (dm) and , current Dg of merchan tabl e trees (D) . ill 50 1 00 80 60 600 400 300 NUMBER 200 OF TREES 100 PER 80 60 ACRE Fi gure l 7.--Maintaining residual stocking l evel by frequen t thinnings: A, At hi gher a verage l evel; B, at lower a verage l evel . When thinning is done at the " f i rs t poss ible " age , mos t merchan t ­ able t re es will b e capable o f respond­ ing t o releas e . There fore , cut trees c an b e proport ionately dis t ribut e d across t h e ran ge i n merchan t able s i z e If largest t rees are ( d iD 1 .0) . not cut , d i D may drop t o about 0 . 9 . On the other hand , the smaller mer­ chant able t rees could b e l e f t to grow ; it is more import ant to pro perly space des irable large t rees than t o remove t h e s e smaller trees . Leaving thes e smaller t r e e s wo uld push d i D back t oward 1 . 0 . I n y o u n g s t ands , = §./ It has not been uncommon for diD to exceed 1.0 at first thinning on experi­ mental plots. 14 - ,.. .. . t h e r e are opporcun i t i e s t o r emo ve "o ugh dominant s ; t h e r e fo r e , diD on goo d s i t e s ( younger ) cou l d be a l i t t l e great e r t han on poor s it e s ( o l de r ) . When f i r s t c h i n n i n g is d e l a y e d , me rchan t ab l e t r ees wi l l i n c lude supp r e s s e d t r e e s , n o t capab l e of r e sp o n d i n g t o r e l e as e . A l so , n e i gh ­ b o r s o f l ar g e t r ees wi l l be l e s s ab l e t o r e spond t o r e l e a s e . Th e r e f o r e . t he s i z e d i s t r ibut ion o f cut t r e e s s ho u l d b e skewed t owar d t h e sma l l er trees . L ikewi s e , succe s s ive t h i n ­ n i n g s wi l l h a v e lower d i D rat i o s b ec ause t h inn ing wi l l g e ne r a l l y i n c l ude i n c r e as i n g p r o po r t ions o f lower crown c l a s s e s . The min imum p o s s ib l e d i D rac io for a given p e r ­ c e n t c u c is dec erm i n e d by s t an d s t ructure ; f o r examp l e . r emoval o f o n e - t h i r d o f t h e b a s a l ar ea b y cut t in g t h e smc : l e st t r e e s i n an average natur al s t an d wou l d r e s u l t i n a d i D rat io o f about 0 . 8 . Results of Thinning b . Terminat e s at an age compat ­ i b l e w i t h f in a l -har v e s t age , s o t ha t merchan t ab l e port ion o f s t an d w i l l b e at app r o x i ­ m a t e l y MAX I MUM s t o c k i n g l ev e l a t t ime o f f in a l harve s t . c . I s done at t he l o n g e s t int e r ­ v a l s comp at ib l e w i t h o t h e r e l emen t s o f t h e r e g ime ( t ab l e 1 ) . d . R e du c e s me rchan t ab l e port ion of the s t and , a f t e r e ac h t h innin g , t o t h e RECOMMENDED RES IDUAL s t o c k i n g l ev e l o r t wo - t h i r d s o f i t s b as a l area b e f o re t h in n in g , wh i chever i s great e r . e . Fol lows t he d i D r a t i o r e g ime in f i gure 18 in wh i c h d i D r a t io i s 1 . 0 at t h e f ir s t t h i n n in g . N U M B E R A N D S I ZE O F T RE E S In the f o l low i n g d i s cus s ion o f e f f e c t s o f t h i nn in g , a s est imat e d by D F I T , o n n umb e r a n d s ize o f t re e s harve s t e d an d o n vo l ume product ion , we emphas ize re sul t s app l ic ab l e t o s e l e ot e d t h inn i n g re g im e s a n d g i ve d e t a i l e d re sul c s p e r t inent c h e r e t o . We i n c l u d e s upp l emen t ar y d i s cus s io n s o f some e f f e ct s o f a l t e r n a t e r e g imes . A l l r e sul t s obc a i n e d from DF I T app e ar t o b e compat i b l e i t h wh at we h av e o b s e r v e d on exper imen t a l p l ot s . Charact e r ist i cs o f t h e s e l e c t e d r e g ime s are : 1 . Pre comme r c i a l t h in n ing , i f don e , i s done at age 1 0 , 1 1 , 1 2 , 1 3 , an d 15 y e ars on s i t e s I t h ro ugh V) resp e c t ive l y . 2 . Minimum merchan t ab l e d . b . h . 6 . 0 i n c h e s ( 15 . 2 cm ) . MAXIMUM s t o c k i n g l eve l ( fig . 1 3 ) . is 3 . S t ands are c l e ar cut at age 85 , 90 , 95 , 100 , and 1 05 ye ars o n s it e s I t hrough V , r e s p e c t i ve l y . 4 . Commer c i al t h i nn in g : a . I s b e gun a t t h e f i r s t p o s s ib l e age ; i . e . , when merchan t ab l e port ion o f s t an d r e ach e s R e l at ion B et w e en Num ber and S i z e Th e r e l at ion between numb e r an d D g , t o ge t h e r w i t h how it i s a f f e c t e d b y t h in n in g , is i l l u s t r a t e d i n f i gu r e 1 9 . Th is p art i c u l a r examp l e i s f o r a s it e I s t an d . The e xamp l e , however , app l i e s to a l l s i t e s ; o n l y t h e a g e s a n d t h e number o f i n t e r va l s wo u l d vary app r e c i ab ly . Th i s s t and h as been t hi n n e d f i ve t ime s , b e g i n n i n g a t age 25 , a n d carr i e d t o a ro t at io n age o f 85 years . Numb er a n d Dg o f me r c h a n t a b l e t r ee s , at t ime o f e ach t h i nn in g , are shown for both t h e t h i n n e d a n d unt h in n e d s t ands . Also shown a r e t h e appr o x imat e cumu l a t e d s t an d curves ( s ee f i g . 4 ) . Not e t h a t at t h e e n d o f e ac h p e r io d , Dg in t h e t h inned s t an d is great e r t h an t h at o f a l l merchan t ab l e t r e e s i n t h e unt hi nn e d s t an d . At t h e en d s o f t h e f i r s t two i n t e r v a l s ( ag e s 3 2 and 40 ) , howeve r , t re e s in t h e t h in n e d st and are sma l l e r t h an t h e s am e numb e r o f larg e st tre e s i n t he un t h i n n e d s t an d . At t h e end o f the t h i r d i n t e rva l , equal numb e r s of the l ar ge s t t re e s are n e ar l y the s ame s i ze in t he two s t an ds . Th ere a f t e r , equal numb e r s o f the l ar ge s t t r ( ", s are a l i t t l e l ar g e r in t h e t h i n n e d s t an d t h a n i n t he unt h in n e d s t an d . 15 THINNED STANO UNTHINNED STANO 400 300 " OJ OJ .... 200 W 100 OJ (. In J crIII rr [j <J: 80 w SO rr Q. <J: w rr <J: .J <J: {f) <J: ill 40 30 20 SOD 400 300 100 2CC 80 SO 50 40 NUMBER OF TREES PER 30 20 10 8 ACRE Fiqure 19.--Rel a tive development of thinned and un thinned stands, by number and si ze of rees a t specifi ed a ges . Num ber of Trees The number of t rees wh i c h a t t a i n a s pec i f ie d min imum d . b . h . i n average n a t ural s t a n ds i s assumed in dependent of s i t e i n dex ( f i g. 5B ) . In s t an ds p recommerc i all y th i nned to the des i red number o f t rees , t h e number reachi n g a spec i f i ed min imum d . b . h . will b e If t he s ame a s f o r n at ural st ands . we begin commercial t h i n n i n g a t t h e f ir s t po s s i b l e age , all these mer­ chant able t r ees wi l l be harves t ed . Number o f Trees at F i n al Harvest The part o f this total number o f t r ees wh i ch will be carr ied t o f i n al harvest varies w i t h s i te 16 qua l i t y , r o t a t i o n age , a n d s t and For our selected t reatment reg ime . t h i n n i n g reg i mes , number at f inal harvest ran ges from 1 3 to 4 5 percent of t he t o t al i n i t i al number o f merchan t ab le t r ees ( t able 2 ) . Thi s r el at i ve number a t f i n al h arvest increases with decreas i n g s i t e qual i t y an d i s l es s for s t an ds t hat were precommer c i ally t hinned t h an f o r those that were not . Re l a t i ve to unt h i nned s t an ds of t he s ame s i te and age , t h inned s t an ds have about 50 to 75 per cen t as many merchant able t rees to be cut at f in a l harves t . The smallest rela­ t i ve number is in good s i te , p re­ commerc ially t h i n n ed s t an ds , and the greatest i s i n poor s i t e st an ds that were n o t precommer c i a l l y th inned . rab e 2 - -Nwnbe1' IN), avemge d. b . h . IVg) , and basal area ::>el' ac1'e IC) of 'ne1'oncmtab l e tl'ees B. O inches ' and ZaPger in d. h. h. at fina l harvest of thinned st::mds, and their l'e Zc.tionship -;0 tot;al yi'e ld of thinned stands and to live stand ay; ;';,na l harvest of unr;hinned s tand8 S i te Age Current thi nned stand Treatmentll N Og I I n c h es Years I I G Square Re l a t i v e to unthi nned stand Percent of total y i e l d of thi nned stand I I N G .. .. .. .. .. .. N I \ Og G - - Percent - 85 CT al one P C T + CT 60 53 28 . 2 31.0 263 277 15 13 56 55 56 50 122 134 85 89 II 90 CT a l one PCT + CT 78 66 23 . 8 26 . 8 240 256 19 16 58 56 60 50 118 133 83 88 III 95 CT a l one PCT + CT 107 84 19 . 3 22.6 216 234 27 21 61 58 64 50 114 132 82 88 IV 100 169 1 17 14 . 4 18. 1 191 210 42 29 66 62 74 51 105 132 81 89 V 105 CT a l one P C T + CT 180 13 . 5 178 45 67 54 1 30 90 PCT + CT N per acre X 2 . 4 7 1 = N per h a ; Metr c conver s i on : G i n feet p e r acre X 0 . 2296 = G i n m2 p e r h a . 11 C T i s cornne rci a l thi n n i n g ; P C T is Og in i nches X 2 . 54 = Og i n centimeters ; preconrnerc i a 1 th i nn i ng . ; OO D i s t r ibut ion o f Tree s Over Time Merch ant ab l e t r e e s not r emain i n g for t h e f inal harvest h ave b e e n removed in comme r c i a l t h in n i n gs ( f ig . 20 ) , Many t r e e s are r emove d at t h e f ir s t t h i nn i n g , an d p r o g r e s ­ s i ve l y f ewer a t e a c h succe s s i ve t h i n n in g . St ands on good s i t e s r e c e i ve more t hinn i n g s t han t h o s e o n poor s it e s ; s t an d s that were pre comme r c i a l l y t h in n e d rece ive mor e t h i n n i n g s t h an those wh i c h were n o t p r e comme r c i a l l y t h i nn e d . The numb er o f me rchantab l e t r e e s in t h e unt h i n n e d st an d i s gre a t e r a t e ach age . D i am et e r of Trees D i ameter at F in a l Harve st As a coro l l ary to numbe r o f t r ee s , t h e D g o f t r e e s at f i n a l harvest ranges f r o m about 5 t o 3 5 p e r c ent l arger in t h in n e d st ands t h an the ave r a g e of all merch ant ab l e t r e e s r ema i n i n g i n un t h i n n e d s t an d s ( t ab l e 2 ) . The g r e at e st di f f e r e n c e i s in st ands t h at h ave b e e n p r e c om­ me r c i a l l y t h i nn e d . When we comp are D g of r e s i du a l t r e e s in s t an d s wh i ch we r e o n l y comme r c i a l l y t h i n n e d w i t h t h o s e o f t h e s am e n um b e r o f l ar g e s t t r e e s i n natural s t an d s , howe v e r , we f in d t hat t he y are s imi l ar ( f ig . 19 ) . I n p r e aomme r a i a Z Z y t h i n n e d s t ands , t h e Dg o f t r e e s 650 SITE I I A \ \ \ \ \Tltll Und \ \ \ 6+ \ \ inchU \ 600 550 500 : \\ 350 300 , ,, , ... 250 200 ... .... ... 1 50 w a: II a: w ... ... ,, 1 00 50 O o 700 [:' 650 f- IJHTHIHHED COfi,4MEflCIAl THINNING ALONE L-L-L10 20 J- 30 B -L 40 -L 50 SITE IV \ \ 600 550 500 450 \ , \ \ \ L-__ 250 200 90 1 00 \ \ ------" " " " " " " ', UHTHIHNED 1 50 COMMERCIAL THIHNIHG AlOHE 1 00 50 80 \ \ \ \ 350 300 70 \Tm! SIM4 6+ 400 80 P'RECOMMERCIAl HIIHHIHG COfi,4MERCIAl THINNING I __LL-L-L- PRECOMMEIICII.L THIHNIHG + COMMERCIAL THIHNING O �L-L-����J--L-L-LI�I�!�I����I�I�!��! o 10 20 30 40 50 T01'AL /J .G E \ 60 {yeers} 70 80 90 100 Figure 20 . --Trends of number of trees over time : A , Si te II; B, Si te IV. 17 !, wi l l general l y be 1 0 to 15 p e r cent l ar ger t h an the D g of the s ame numb er of l argest t r e e s in un th i n n e d s t an ds . I n creased growth o f r e s i d­ ua] t rees is mor e or l es s o f f s e t b y t h e removal o f some larger th an average t re e s in t h i nn i n gs-- t rees wh i ch wou l d o herw i s e b e present at f i nal harve st . Tren ds Over Time The increas e in Dg ove r t ime r e f l e c t s a comb in at i on o f growth o f surv i v i n g t rees a n d l o s s o f t r e e s from t h e s t an d . The r e fore , t h e t r e n d over t ime i n t h inned s t an ds i s qu i e d i fferent f r om that in un t h inned s t ands ( f i g . 2 1 ) . I n an un t h i nn e d s t an d , t h e i n cr e a s e in D g o f merchan t ab l e t re e s progr e s s e s smoot h l y o ve r t ime--on a s t e a d i l y dec l i n i n g numb e r o f t r ees ; t h e i n c r e as e i n D g i s great e r t han t h e d . b . h . growth o f survi v i n g t r e e s . I n t h e t h i n n e d s t and , t h e re i s an abrupt i n c r e a s e in Dg e ach t ime t h e s t an d is t h i n n ed ( i f diD is l es s than 1 . 0 ) ; the i n c r e a s e be tween t h i nn i n gs is the actual growth o f survivors . On good s i t es t h e gain in D g b e c au s e o f commer c i a l t h i n n i n g is gre a t e r than t h e addi t i o n al gain from pre comme r c i a l t h i nn in g . On poor s i t e s , the revers e i s true . The d . b . h . growth rat e with com­ me r c i a l t h i n n i n g alone is ver y c l o s e t o t h e r a t e o f i n cr e ase i n D g o f merchan t ab l e t rees i n t h e n a t u r al s t an d ; -- a l i t t l e b e t t e r on good sites . D i amet e r growt h rate wi t h p r e commerc i a l t h i n n i n g is always a l i t t l e bet t er than with comme r ­ c i al thin n i n g a l on e . SITE " Effects of Al t ern at e Thinning R eg i m es M i n imum Merchan t ab l e D . b . h . ! , ! j ! ) 70 I I 30 1 , 80 I 100 SITE IV 20 18 16 14 The l ar ge r t h e min imum merchant ­ ab l e d . b . h . ( dm ) is , t h e fewer and t h e l arger are the t rees harves t e d ( t ab l e 3 ) . The e f f e ct on t h e f in a l harvest is f a i r l y minor . With only commer c i al t h innin g , a l arger dm t ends t o r e s u l t in s Zight Z y mor e t re e s an d sma l l e r t rees a t f i n al harve s t ; in p r e commer c i a l l y t h i nn e d st ands , t he r e v e r s e is t rue . The maj or e f f ect is on t r e e s removed in t h inn i n gs . With a l a r ge r dm , a much sma l l er number o f t re e s i s remov e d ; a s a r e s ul t " t h e Dg o f b o t h t h e t r ees harve s t e d i n t h i n ­ n in gs a n d al l t r e e s harve s t e d is l ar g e r . 12 10 t 0 D e l ayed First Thi n n in g I 10 I \ 20 t ! 30 I ! 40 • ! 50 ! , 80 I TOTAL AGE ( yeat"'S) [ 70 ! ! ! 30 Figure 21 . - -Trends of Dg over t:ime : Si t:e II; B , si t:e IV. 18 I , 80 I 100 A, The e f f e c t o f de l a y i n g t h e f i r s t commer c i a l t h i n n i n g i s s i m i l a r Fewer l i v e t re es t o t h e f o r e go i n g . wi l l b e av i l a b l e for harvest in t h i n n i n g s ; howev e r , p art o f t h i s l o s s c an be o f f s e t b y sal vage o f r e c e n t merchan t abl e mor t a l i t y Growth rat e s o f mer­ ( t ab l e 4 ) . ch a n t ab l e t r e e s w i l l b e r e duc ed somewh a t dur i n g t h i s de l ay p e r i o d ; IDg) Tab l e 3--/lumbe"l' (N) and ave"l'age d. b . h. of t,.., e8 ha:rvested in e<Xrme"l'cnaZ thi7l1li7l{1B finaZ ha:rvest, by Bite, treatment, and minimum TTl€1"ehantab Ze d . b . h . (dm) S i te I CO/TlJonen t a n d dm II S i te II 1 09 I II I nc h e s Og I I S 1 te III II I I nches S i te V S i te IV l ug I 1 09 N I nches and at N 1 09 I nches I nches ONLY COMMERCIAL TH I NN I NG 6 . 0+ i nches : Thi n n i ngs F i n a l harvest Tota l number of trees 343 60 403 10 . 6 28 . 2 14 . 6 327 78 405 10 . 0 23 . 8 13 . 8 295 107 402 9.3 19 . 3 12.8 235 169 404 11.4 225 64 289 12 . 1 27 . 3 16 . 7 207 82 289 11.5 23.0 15.6 176 113 289 10 . 9 18 . 5 14 . 3 116 172 288 10. I 14 . 2 12 . 7 284 1 17 40 1 9.2 18.1 12.5 222 180 402 8.5 13.5 11.0 176 10 . 9 18.7 161 1 26 10 . 2 14 . 4 287 12.7 8.6 14 . 4 7 . 5+ i nches : Th i nn i ngs F i na 1 harvest Tota l number of trees PRECOMME RCIAL TH I NN I NG + COfotIERCIAL TH I NN I NG 6 . 0+ i nches : Th i nn i ngs F i na l harvest Tota l number of trees 349 53 402 10 . 9 31.0 15.2 336 235 52 287 12 . 8 31.1 17 . 6 222 65 -287 66 402 10 . 4 26 . 8 14 . 4 402 9.8 22 . 6 13.5 12 . 1 27 . 1 205 82 ll.5 22 . 9 III 16 . 7 287 15 . 7 287 318 84 7 . 5+ i nches : T h i n n i ngs F i na 1 harvest Tota 1 number of trees Hetri c convers i o n : II per acre Og X in inches 2 . 471 X 2 . 54 II = pe r h a . Og = 14 . 4 in centi me ters . T a b l e 4--Numbe1" (N) and average d. b . h . (Dg) of merahantab Ze Zive trees 1"emaining if thi.7I1Ii.ng i. de Zayed, and aorrespcmding pe1'i<xi.ic r"hantabZe d. b . h. 1.0 merehantabZe mo1"taLity, when minimum 6. 0 inaMs .... S i te Fi rst po s s i b l e age for thl nnlng HUI!tler of years d e l ayed 0 H 1 09 J H .!2s Years 30 20 10 lXi N Inches I Og H I nches I ll9 I nches LIVE TREES I II III IV V 25 30 36 47 68 403 405 402 403 403 8.0 8. 1 8.0 8. 1 8.0 312 333 362 382 DEAD I II III IV V 25 30 36 H 68 0 0 0 0 0 Ketri c conyers i o n : 11 . 0 10 • • 9.7 9.2 H per a c re X 13.8 12.7 11. 4 10 . 4 176 206 2048 16.4 14 . 8 13.0 85 76 65 46 8.3 7.9 7.4 7.2 50 50 10 . 1 .9.3 11 . 4 TREES 91 72 40 21 lXi i n i nches 227 256 297 336 6.8 6.7 6.8 7.2 2 . 47 1 X 2 . 54 H • • 49 per hectare . Il9 in centllneters . 19 f urt hermore , t h e y wi l l c o n t i n u e t o b e lo we r a f t e r t h i n n i n g than i f t h e s t an d h a d b e e n t h in n e d ear l ier-­ t h e l o n g e r t h e d e l ay , t h e gre a t e r ehe loss . Fr eque n c y and S e v e r i t y o f Thinn in g Redu c i n g t h e average growin g s t ock l e v e l l e ads t o f ewe r and l arger t re e s at f i n al harve s t ; i n c r e a s i n g the average level l eads t o mo re a n d smal l e r t rees at f i n al harvest . R e l at i n g t h i s to t h e n at ur a l s e and curve i l l u s t r a t e s h ow D g ae f in a l h a r v e s t o f t h e s e t h in n e d s t an ds compares wi th t h a t o f t h e s am e n umb e r o f Zarge s t tre e s i n t h e un t h i n n e d s t an d ( f i g . 2 2A ) . Dg o f t rees remo v e d in t h i n n i n g s i s g r e at e st i n t h e s t an d main t a i n e d at a h i gh aver age l e v e l a n d l e ast i n t h e s t an d main t a i n e d at a l ow average l ev e l , but di f f e rence are smal l . 'i3" Q) .2! ! lJ '" [ ill (L 300 200 50 1 1 :: so li: Og (lncnesj '" ill [ '" Type o f Th i n n i n g The h i gh e r t h e d i D r at i o i s a t t h e f i r s t D o s s i b l e comme r c i al t h i n ­ n i n g , t h e g r e at e r t h e numbe r o f t rees an d t h e sma l l er t h e average d . b . h . o f t r e e s rema i n i n g at any t im e ( f i g . 22B ) . Th e r e l at ive d i f f e r e n c e d e cr e a s e s with e ach succ s s i ve t h in n i n g . Thus , d i f f e r e n c es , both at f i n a l h arve s t a n d ave raged over the r o t a t ion , tend t o de c r e a s e with i n c r e a s i n g site qua l i t y and t o b e l es s i n s t ands that are p recommer ­ c i a l l y t h i n n e d t h an i n t h o s e that are n o t . VO L U M E Y I E LD G ains i n Tot al M er ch an t a b l e Pro d u ction G a i n s in merch a n t ab l e produc­ tion depend on mer ch an t ab i l i t y s t an dards , a s well a s o n t h e age at Th e r e ­ whi c h comp ar is o n s are made . fore , t o i l l us t rat e t h e b as i c e f f e c t s o f t h i n n i ng , w e f i r s t exami ne r e l a ­ t io n s h i p s i n t e rms o f cub i c vo l ume To provide of t o t al s t em ( CVTS ) . p e r s p e c t i v e r e gardi n g t h e e f f e ct of a ge , we compare max i mum me an annual i n creme n t s ( m . a . i . ) , regardl e s s of 20 400 300 200 NUMBER OF TREES 100 PER eo so ACRE 40 Figure 22. --Effects of al terna t e thinning regimes on number and size of trees : A , Al t ernat e s tocking and thinning in terval ; B, al t erna te type of thinning (diD regime) . ages at wh i ch t h i s culmi n a t ion o c cur s ; w e then i l l us t r at e t h e e f f e c t s o f maki n g comp a r i s o n s a t other ages . We t h e n examin e gains at our s e l e c t e d r o t a t i o n ages by various measures of mercha n t ab l e p r o duc t i o n . Culmi n a t i o n o f m . a . i . Wh en d o e s m . a . i . culmin a t e ? A n y p r act i c e o r merchan t ab i l i t y s t andard t h a t i n c r e as e s merch ant ab l e p r o duct ion ear l y i u t h e r o t a t io n . more t h a n t owar d t h 0 e n d wi l l r e du c e t h e a g e at wh i ch culminat i o n o c curs . 330 C o n v e r s e l y , anyt h i n g t h a t i n c r e as e s mer chan t ab l e produc t ion l at e r rather t h an e ar l i er in t h e r o t a t ion wi l l i n c r e as e the age o f culminat ion . Thus , whe reas g r o s s m . a . i . in t e rms of t o t al s t a n d CVTS culminat es at about age 63 on a l l s i te s , mer­ chant ab l e m , a . i . c u l m i n a t e s l at er on p r o g r e s s i v e l y p oo r e r s i t e s --ran g i n g f r o m about 6 5 y e ar s o n s i t e I t o 1 0 5 y e ar s on s i t e V . T h i n n i n g s r e duce the ages at wh i ch mer ch an t ab l e m . a . i . c u lm i n at e s ; t h us , culmin a t i on a g e s i n s t ands r e c e i v i n g e ar l y pre­ comme r ci a l t h in n i n g range f rom about 65 to 85 years on s it es I through V . B o a r d - f o o t m . a . i . culminates at l at er a g e s t h an cub i c - fo o t m . a . i . O n e s h o u l d f ir s t have a c l e ar und e r s t an d i n g o f t h e r e l at i o n s h i p o f m e r chan t ab l e y i e l dZl t o t o t al y i e l d in natural s t an d s ; t h e n one s ho u l d compare m e r oh a n t ab Z e y i e l ds wi t h a n d without t h i n n i n g . At ages of m a ximum m . a . i . , t o t al gro s s y i e l d in unt h inned s t ands i s abo ut 3 8 p e r cent g r e at e r t h an net y i el d . The p o r t ion o f t h i s g r o s s y i e l d con t ai n e d i n mer ch an t ab l e t re e s -­ 6 i n ch e s ( 1 5 cm ) d . b . h . and l ar ger-­ i s abo u t 8 to 33 p e r c ent l e s s t han t o t al g r o s s y i e l d on s i t e s I t h r o u gh V ( f i g . 2 3A ) . M e r ch an t ab l e gro s s y i e l d o f s t an ds r e c e i v i n g o n l y comme r c i a l t h i nn i n g i s l e s s t h an t h at o f unt h i n n e d s t an ds ( f i g . 2 3B ) . Th is is a r e s u l t o f the r e du c e d growth p e r a c r e wh i ch o c c u r s be tween f i rst t h i n n i n g and f in a l harvest . Th i s l o s s i s , g e n e r a l l y , cons i de r ab l y l e s s t h an t h e mer c h an t ab l e v o l ume o t he rw i s e l o s t to mo r t a l i t y , howev e r . Y i e l d of p r e comme r c i a l l y t h in n e d s t an ds is a l so l e s s than t o t a Z gro s s y i e l d of un t h inn e d s t an ds , but n o t as much so . A l l pr e c om­ mer c i a l l y t h i nn e d s t an d s r e c e i ve a l a s t i n g b e n e f i t f rom concent rat i n g growt h o n t o merchan t ab l e t r e e s . 11 I n terms of CVTS , y ield refers trees . to CVTS of merchantable merchantable size A 30e :- 270 t:- 240 f:.. 2 10 [ r "o 1 50 1 20 80 80 W :;: Cub i c Vo l ume o f To t a l S t em r 3 33 0 t " k ,.. t o ' 80 0 [ ; " 270 240 2 10 3 ! ! 100 t b ' 80 ! ! 120 ! ! ! 1 40 1 80 , 1 80 ! 200 MERCHANTABLE GROSS m.•.!. WITH PRECOMMERCIAL THINNING ... COMMERCIAL THINNING I­ :t ! 8 00 ..J 3 z ! ! !-L ! _' 100 !-L ! !-L ! !-L ' ! 120 !-L ! !-L ! !-L ! ! 140 INITIAL SITE INDEX 1 60 ! -L ! 1 80 L ! ! 200 (100-yesl""basls) Figure 2 3 . --Mean annual incremen t of cubic vol ume of total s t em (CVTS) at time i t culminates , when minimum merchan tabl e d. b . h . is 6 . 0 inches : A , Merchantabl e m. a.i . relative to total m. a .i. in un thinned s tands ; B, merchantable m . a .i . wi th commercial thinning rela tive to that without thinning. The add i t ional b e n e f i t a s s o c i a t e d w i t h imp r o v e d h e i ght growt h i n c r e a s e s with d e c r e a s i n g s i t e qua l i t y . At ages o t h e r than culmin a t i o n o f m . a . i . , r e l a t i ve y i e l d s are somewhat d i f f e rent from t h o s e 21 j l; , : I r: 1 [" J I · ; !. di scus s e d above ( f i g . 24 ) . At ages pr io r t o culminat ion , di f f e r e n c e s i n m . a . i . b etween pre comme r c i a l l y t h inn e d s t ands and t hose r e c e i v i n g o n l y comme rcial t h inning a r e a l i t t l e great e r ; at ages l at er t h an culmin a t ion , d i f ferences in m . a . i . Prior t o culmina­ are a l i t t l e l e s s . t ion , d i f ferences in mer ch ant ab l e y i e l d b e tween un t h i nned s t ands and comme r c i a l l y th inned st ands are l e s s ; a f t er culminat ion , d i f f e re n c e s are greater . The rot at ion ages we have chosen for our e xamp l es are about 2 0 years beyond culminat ion o f merchantab l e CVTS in precommer c i a l l y t h i nn e d st ands , b u t sub s t ant i a l l y ear l i er t h an culmin ae ion o f m . a . i . in t erms o f board-foot vo l ume . For t h e remainder o f t h i s s e c t ion , a l l compari sons are b ased o n t h e s e chosen rot at ion ages . Ga i n s b y D i f f e r en t Produ c t ion easures o f Mean annual incremen t s by f i v e me asur e s o f mer c hant ab l e produce i o n are shown in t ab l e 5 f o r t h e s e l e c t e d r e g ime s . Th i s t ab l e is i n t e n d e d t o p r o v i de t h e r e ader w i t h s ome s p e c i f i c e s t ima t e s o f y i e l d , a s we l l a s p r o ­ vi d i n g b ackground for sub s equen t d i s cuss ion . anm...a t inaPemlmt at speoified potation ages, by site, !:reatment, and VariOU8 meaSUl"es of productionJ' trees D . O inohes and "larger i n d . b . h. Tabl e 5 ..-Mean Measure of prodUc!ionY Site t age. and treatment Site r. CVTS Square feet • . ! CY4 Cubic · · feet SV6 IV5 • • Board · feet B5 years: 3 . 65 3.87 5 . 55 5 . 92 215.8 227 , 8 278.4 289 . 3 212.2 224 , 0 270. 1 280.7 1546. 1 1627 . 1 1863 . 5 1947 . 7 139 7 , 1466. 16J6 . 1728. 3.22 3 . 40 4 . 54 5.03 170 . 4 179.2 214.2 228.9 167 . 3 175.9 207 . 1 221.5 1175.9 1231. 7 1376 . 4 1490 . 4 104 5 . 3 109 1 . 5 1 182. ! 1300. Z Unthinned Unthinned plus sal vabl eY Conrnerc ial thinning alone PrecOll111e rcial thinning + cOll111e r cial thinning Site I V , 1 0 0 years: 2.79 2 . 94 3. 75 4 . 21 129. 0 135.2 155 . 8 176.5 125.7 131.6 149. 5 169.9 832 . 0 867.2 939. 3 1095. 3 714.5 741 . 7 i78 . J 931.3 Unthinned Unthinned plus sal vabl "y COll111e rcial thinning a l one Precorrrne rcial thinning + corrmerciaJ thinning 2 . 36 2 . 47 2. 87 3 , 41 91. 6 95. 6 103 . 8 128 . 7 87. 9 91.6 98. 1 122 . 7 528.2 547 . 4 565. 8 744 . 6 427 . J 440 . J 436. 9 607 . 9 1 . 88 1 . 95 2 . 52 57 . 7 59. 6 82 . 4 53.6 55 . 3 7 7. 2 276.8 284. 1 426.9 189. 4 192.9 320.6 Unthinned Unthinned p l us sal vabl eY Canmereia 1 thinning al one Preconmercial thinning + corrmerci a 1 thi nnin9 Site I I , 90 years: Unthinned Unthinned plus salvabl eY COll111ercial thinning alone Precomnercia 1 thinning + canmercial thinning 310 280 Site I l l , 95 years: 270 250 ;; 230 • .!! .Q 2 1 0 .0 B Iz i SITE / I wtln + 1 80 1 70 orecommefCIal HlInnrng com caaJ thinning UNTHINNEO STAND, GAOSS m.a.1 Witn commercIal :hlnnlng only 1 50 Site VI 105 years: Unthinned Unthlnned plus salvabl eY Preconmercia 1 thinnin9 + cOll111e r oia 1 thinning Metri c conyersi6n : 1 30 G in square feet per acre meters per hectare. CV in cubic feet per acre meters per hectare. X X 0. 2296 = 0.0 700 = G in SQU'N! CV in cubic Y G = basal area; CVTS = cubic volume of total stem; CV4 = cubic volume to a 4-inch (lO-cm) top d . i . b . ; IV5 • l nternationa 1 board- foot vol ume to a s-inch ( l3-on) top, 1/4-inch kerf; SV6 = Scribner board-foot volume to a 6-inch ( IS-em) top (Scribner volume is the fonnula version appl ied to 16- foot l ogs ) . CV4 , I V 5 . and SV6 excl ude vol ume in the stumP . Y For this comparison, a l l mercnantab l e roo rtality occurring within 10 years of final harvest is assumed to be salvable at final harvest. 1 10 80 70 50'��������-J� 30 40 50 60 70 80 80 1 00 1 1 0 1 2 0 STAND AGE (yea""j Fi gure 2 4 . --Mean ann ual incremen t of merchant ­ able cubic vol ume o f total stem (CVTS) a t ages before and after culmina tion, when minimum merchan table d . b . h . is 6 . 0 inches . 22 To p r o v i d e r e a l i s t i c e s t imat e s o f ga ins from t h i n n in g , w e h ave added e s t imat e s of sal v ab l e mor t a l i t y t o t h e l iv e vo l umes harve s t e d f rom un t h in n e d s t ands . Th i s is the e s t imat e d merchan t ab l e mort a l i t y wh i ch o c curred dur in g t h e 1 0 - year per i o d p r ior t o harves t . The amount t hat t h i s s a l vage adds to t o t al usab l e product ion d e c r e as e s with de c r e as i n g s i t e qual i t y an d wi th measures o f merchan t ab l e v o l ume wh i ch exclude progres s i ve l y mor e o f t h e t o t a l vo lum e . Th e correspon d i n g p e r c e nt a g e and ab s o l ut e gains i n CV4 y i e l d are i l l us t r at e d in f i gure 2 5 . Gains f rom o n l y comme r c i al th i n n i n g are great e s t on good s i t es , whe r eas t h e additio n a Z gains f rom p r e com­ m e r c i a l t h i n n i n g are greater on p o o re r - t h an - average s i t e s . With our as s umpt i o n s , t h e maximum i n c r e a s e in CV4 m . a . i . due t o p r e comme r c i a l t h i nn in g o c curs a t about s i t e i n d e x 1 00 . R e l at ive ga ins b y other me a s u r e s o f v o l ume p r o duct ion are shown in t ab l e 6 . so iab l e 6 --Gaine in mean annual. incremen t to Bpeaified potation ageB, oue m asUl'e of production; 'trees a. 0 by 81- te and !J inanss and larger in d.b. h.J ! Measure of producti onY Site, age, and trea tment CVTS ABSOLUTE GAIN Square S i te I , 85 years: Coomerc i a l thinning a l one Precoomercial thinning + comnere i a l thinning Oi fference 11 S i te I I , 90 years: Coomerc i a I thinning a lone Preccnmercial thinning + coornerc i a l thinning Oi fference 11 S i te I I I , 95 years: Conmercial thinning a l one Precoomerci a l thinning + conmerc i a l thinning Di fference 11 S i te IV, 100 years : Conmerc i a l thinning a l one Precoomerc ial thinning + coomerc i a l thinning Di fference 11 Site Y. lOS years: Conmerc i a l thinning a l one Precomnerci a l thinning + coomere i a l thinning Di fference 11 S i te I , 35 years: Conmercial thinning a l one Preccmnerci a l thinning + conmerci a l thinning Di fference 11 S i te l l , 90 years : Conmereial thinning a l one Precocrmerci a l thinning + conmerc i a l thinning Di Herence 11 Site I I ! , 95 years : Canmercia 1 thinning a lone Precoomerc i a l thinning + corrmerci a l thinning Di fference 11 Site IV, 100 years : COlmlereial thinning a l one Precoarnerci a l thinning + conmere i a I thinnino Di fference 11 S i te V, 105 years : Comnereial thinning a l one Precanmerci a I th i nni ng + conmercia I thinning Di fference 11 I SV6 CV4 - - Cubic - feet - - Board feet 1 . 6B 2 . 05 50 . 6 61.5 46. 1 56 . 7 236 321 170 262 . 37 10 . 9 10 . 6 B5 92 1 . 24 1 . 63 35.0 49 . 7 31.2 45.6 145 259 91 209 . 39 14 . 7 14.4 114 1 18 .81 1 . 27 20. 6 41.3 17.9 38 . 3 72 228 37 190 . 46 20 . 7 20 . 4 156 153 . 40 . 94 8.2 33. 1 6 .5 31 . 1 18 197 - 3 168 . 54 24 . 9 24 . 6 179 171 .57 22.8 21. 9 143 128 .57 22.8 21.9 143 128 PERCENTAGE GAIN - Percent 43 53 22 27 21 25 15 20 12 18 20 27 18 26 12 21 8 19 10 36 48 10 11 5 28 43 15 31 14 29 8 26 26 16 15 14 17 21 16 38 9 35 7 34 3 36 - 1 38 22 26 27 32 39 29 38 40 50 66 29 38 40 50 66 1/ Ga ins are relative to unthi nned plus s a l vabl e (from tab l e 5 ) . Y G = basal area; CVTS = cubic vo l ume of total stem; CV4 = cubic INITIAL SITE INDEX [100-yeBr-bBsis) vol ume to a 4-inch ( IO-em) top d . i . b . ; IV5 = lnterna tional board-foot vol une a s-inch ( I3-cm) top, 1/4-inch kerf; SV6 = Scribner board-foot volume to a 6-inch ( I S-em) top (Scribner vol tll1e is the fonnula vers ion appl ied to 16-foot 1 09S ) . CV4, IV5, and SVG exclude vol ume in til. stump. 11 Di fference between precOOTIl(!lx i a l thinning + co",, rcial thinni g and comnerc i a l alone is gai n due to precomnerc i a l thi nning. Figure 2 5 . --Gains in mercha n tab1.e cubic vol ume (CV4) due to thinning: B, percentage ga ins . A, Absol ute gains ; 23 o t e that r e l a t i ve gains from comm e r c i a l t h in n i n g decrease as l e s s of the t o t a l vol me in mer ­ cha n t ab l e t r ees i s deemed usab l e ; t h at is , gr e at e s t for CVTS and l e ast for SV6 . Th e t r end for add i t ional gains f rom p re comme r c i a l t h i n n i n g i s t h e reverse ; l e a s t f o r CVTS an d gre a t e s t f o r SV6 . R e l a t i ve gains on d i f f e r e n t s i t es a l so vary wi t h the measure o f produc t i on ; however , b y a l l measures , the , ' ab s o l u t e gain f r om p r ecomme r c i al t h i n n i n g is great e s t on s i t e I V . D i st ri bution o f Yi el d s Over Tim e Th e ac cumul a t ion o f t o t a l y i e l d o v e r t ime an d t h e d i s t r ibu­ t ion of this y i e l d amo n g thinn i n g s and f in a l harve s t , f o r t h e s e l e c t e d regime s , a r e i l l u s t rated i n f i gures 2 6 - 2 8 f o r s i t e s I I and I V in t e rm s t SOO 450 400 SITE I I A 22 r I SITE I I A 18 r 16 20 iJrlhll fO 14 "68 :t 10 W a: u « 0 a: w 6 a. 1 w :J :1 F 14 ..J 5+ 0IIC1'ln I 8 :0 I I 20 , I 30 , I 40 I I 50 I I 80 I 70 SITE IV I 8 I 0 I I I I 80 1 00 80 lOC 0 12 > 8 6 10 4 2 L L 350 24 0 0 300 10 20 30 TOTAL AGE {years) 40 50 8e 70 8 0 Figure 27 . --Trends of cubic vol ume (CV4) A , Site II ; production over time : B , site IV. 250 200 1 50 100 50 0 0 400 LO 20 30 40 50 60 70 80 80 1 0C SITE IV 8 350 300 UM 'Mn!O 250 200 150 1 00 50 0 a 10 20 30 AGE ( years) 40 50 60 70 80 80 1 00 Figure 26 . --Trends of basal area production A , Si te II; B , Si te I V . over time : 24 o f b asal a r e a , CV4 , and IV5 . The t o t a l - y i e l d curves i n these f i gur e s suppl ement t h e f o r e go in g d i s c ussion an d further i n d i c a t e r e l at i ve g a i ns i n y i e l d w i t h s ho r t e r rot a t i on s . Th e fol lowing d i s cu s s ion wi l l focus on the r e s i dual s t an d- -b e f o r e a n d a f t e r each cut t i n g . A t t ent ion is dir ect e d f i rst t o the vo l ume r ema i n i n g f o r t h e f in a l harve s t an d then to vo l ume r emo ved in t hi nn i n gs . �--- r : -�---. SITE I I A l :J C l 2Q L : L , , 7 F 0 SO SC 4C :J C O" O D t ! I I r- L L l f • i : ! I 00 80 3C 7 0 60 2 1 Fi �-�-----��-- f 10 , ' 20 , I 30 40 I ! SO SITE IV , I SO , I 70 , I 80 , ! J 90 I 1 00 B Vo l um e Remo ved in Th i n n i n g s :- I I L-,''--.J1--'-l'--.JI-----1--:-1:':-'-' ' :-'-1-:::1:-'""' -:,-::1-"-,;;,;:::' ' ;::,;:' .,.J Q ure c omme r c i a Z t h i n n i n g , t h i s vo l u me at f i n a l h a r v e s t i s about 8 5 p e r c e n t o f t h e vo l ume o f l i v e me r chan t ab l e t r e e s i n n a t ur a l s t a n ds o f t h e s ame s i t e and age . Th i s r a t i o i n c r eas e s s l i gh t l y w i t h i n c r e a s i n g s i t e i n d e x an d is s l i gh t l y g r e a t e r f o r b o a r d ­ f o o t vo l ume t h an f o r cub i c - f o o t v o l um e . F o r s t ands t h a t w e r e p e c o mm e r c i a Z Z y t h in n e d , t h i s r a t i o o f vo l ume a t f in a l h ar ve s t t o f i n a l vo l ume o f comp a r ab l e n at u r a l s t an ds i n c r e ases wi t h d e c r e a s i n g s i t e qual i t y and , e s pe c i a l l y o n poor qu a l i t y s i t e s , i s s ub s t an t i a l l y g r e a t e r for b o ard- f o o t vol ume t h an f o r cub i c - f o o t vo l ume , On poo r s i t e s , b o ar d - f o o t vo l ume at f i n a l harve s t ( as we l l as t o t a l produ ct ion ) is gre at e r t h an t h at in comp a r ab l e un t h i nn e d s t an d s . 10 2C 30 2 8 . - - Trends prod uction 4C SC TCTAL AGE SO ( years 1 70 of board-foot over t i me : A, Site 80 9C 1 0C volume (IVS) II ; B, site IV. Vo l um e at F i n a l H a r v e s t The cu b i c vo l ume remo v e d a t f i n a l h a r v e s t a v e r ag e s about 7 0 p e r c e n t o f t h e t o t al vol ume harve s t e d from t h e s t and ( t ab l e 7 ) ; t h e r ema i n ­ d e r i s harv e s t e d i n t h i n n i n gs . Th e p ro po r t ion o f vo l ume at f i n a l h a r v e s t var i e s somewh a t w i t h t imespan a n d growt h r a t e a n d w i t h t h e me asure of m e r c h an t ab l e v o l ume . Th us , it is a l i t t l e gr e at e r on poor s i t es t h an on good s i t e s a n d is g r e a t e r i n t e rms o f bo a r d - f o o t vo l ume t han in t e rms o f t o t a l cub i c - f o o t vo l um e . ( I f b e l o w MAX I MUM , vo l ume at f i n al h a r v e s t wo u l d b e p r o po rt i o n a t e l y less . ) The vo l ume a v a i l ab l e a t f in a l h a r ve s t o f t h i nn e d s t an d s is g e n e r ­ a l l y l e s s t h an t h a t o f c o r r e s po n d i n g un t h i n n e d s t an d s ( f i gs . 2 6 - 2 8 , t ab l e 8 ) . For s t an d s r e c e i v i n g o n l y Vo l ume remo ved i n t h i n n i n g s ave r a g e s a b o u t 30 p e r c e n t o f t h e t o t a l me r c h an t ab l e cub i c vo l ume p roduc ed : a l i t t l e mo re on goo d s i t e s , wh i ch r e q u i r e mor e f r equen t t h i nn i n g , t h an on p o o r s i t e s . Propo r t i o n a t e l y l e s s o f t h e t o t a l produ c t ion o f board f o o t vo l ume is removed i n t h inn i n gs b e c a u s e much o f t h e vo l ume in t h i n n i n gs is in r e l at i ve l y sma l l t r ees w h i c h con t ribu t e propo r t i o n at e l y mo re to t o t a l vo l ume t h an to b o a r d - f o o t vo l ume , W i t h our s e l e c t e d r e g imes , t h e f i r s t comme r c i al t h in n i n g removes an ave r age of about 1 , 36 0 cub i c f e e t p e r acre ( 95 m 3 p e r h a ) f rom s t an d s t h a t w e r e n o t p r e c omme r c i a l l y t h in n e d and about 1 , 2 1 0 cub i c f e e t p e r a c r e ( 85 m 3 p e r h a ) f rom t h o s e t h a t were . Th e I I - pe r c e n t l ow e r vo l ume i n p r e c omme r c i a l l y t h i n n e d s t an ds r e s u l t s f r om t r e e s b e i n g youn ge r - - and t h us s ho r t e r -- wh en t h e y re ach the s pe c i f i e d d . b . h . S imi l ar di f f e r e n c e s b e t we e n s t ands that we re pre comme r c i a l l y t h i n n e d an d t h o s e t h a t w e r e n o t p e r s i s t t h ro u gh out a l l s ub s eq ue n t t h i n n i n g s . The amo u n t o f vo l ume remo ved t ends t o i n c r e a s e w i t h e ac h s u c c e s s i ve t h i n n i n g , at a f a i r l y s lo w rat e fo r cub i c vo l ume but m o r e r ap i d l y f o r b o ar d - f o o t vo l ume ( f i gs . 2 7 a n d 28 ) . 25 Tab l e 7- F':"ncZ c::T"'.)est i n :::h irmed stands a n percenr:. ?f' =:nei'l' ;o:a:Z ?l'cduc::ion o spt?a{; :'ed rotatiol1 ages, by site} ;;'l'ea::mer.!.", e su:t'es of production,' trees 6. 0 i nches Z"'..i ::na' !.la:J>':OUG, Arr'g2Y' :.n a . o. fl . Measure o f producti on.1i S i t e , age , and treatment CVTS CV4 IV5 SV6 Percent S i te I, 85 years : Commerc i a l th n i ng a l one Precomerci a 1 thinning + conmercia 1 thinning 15 13 56 55 66 66 67 67 71 71 74 75 Conmerc i a l thinning a l one Precormlerc i a 1 thinning + conmer c i a l thinning 19 16 58 56 66 67 67 68 72 72 76 76 27 21 61 58 68 67 69 68 73 73 76 77 42 29 66 62 72 69 72 70 77 74 81 80 45 57 72 73 77 82 62 69 70 74 78 S i te l l , 90 years : S i te I I I , 95 years : Conmer c i a 1 thinning a l one Precorrmerc i a 1 thinning + corrrne r c i a l thinning S i te I V , 100 years : Corrmerci a 1 thinning a l one PrecofTlTler c i a 1 thinning + corrrne r c i a 1 thinning S i te 'I, 105 years : Precorrmercia 1 th i nn i n g corrme rc i a 1 thinning + verage N number o f trees; G ;: ba s a l area; CVTS ;: cub i c vo l ume of total stem: CV4 = e uo i e '(o l ume to a 4 - i nch ( l O-cm) top d . i . b . ; IV5 = International board-foot vol ume to a 5-i nen ( l 3-em) top, 1/4- inch kerf; SV6 = Scribner board-foot vol ume to a 6 - i n c h ( l 5-cm) top ( S c r i bn e r vol ume is the fonnula version appl ied to !6- foot l ogs ) . CV4 , I V5 , and SV6 exclude vol ume i n the s tump. 11 ;: Tab l e 8--P::r..:z t h(Zr"Jest i n hinned s'tands relative to harvest of !ive ';]'ees n unthinned stwtds at specified l"otation ages, by site, !:Teatmel1t, znd Val""': ous measta'es of' " production; trees 6 . 0 inches and la:l'gel' in d. D. h.l/ S i te I a g e , and Measure of producti onY reatment S i te 1 , 8 5 yea rs : Conmerci a 1 thinning a l one ?recolmlerci a 1 th i n n i ng 01cOlTTle l r c i a l thinning Precorrmerci a 1 thinning g a i nl! S i te I I , 90 yea rs : Og CV4 I 1V5 SV6 Percent 56 50 122 134 85 89 85 89 85 89 86 90 87 93 83 88 83 90 83 90 84 92 86 94 10 Conmerc i a l thinning a l one Precomne rcia 1 thinning oj. corrme rc i a l t h i nni n g Preconrne r c i a l t h i n n i n g ga i n 60 50 Conme rc i a l t h i n n i n g a l one ?recornne rcia 1 t h i n n i n g 040cornne r c i a l thinning Precornne r c i a 1 thinning 9a i n 64 50 Conmerci a l t h i n n i n g . l one Precorrmerci a 1 thinning -+corrmerc i a 1 thinning Precornne rc i a 1 thinning ga i n 74 51 S i te I l l , 9 5 y e a r s : CVTS lI8 133 10 12 1 14 132 82 88 16 82 92 82 92 83 96 82 100 12 12 16 22 Si te I V , 1 0 0 years : 105 132 81 89 81 97 81 98 82 105 83 113 26 10 20 21 28 37 130 90 103 105 118 138 S i te V. 105 yea r s : Preconmerc i a 1 thinning ... cornne rci a 1 thinning 54 1/ Expressed as percent of v o l ume i n l i ve trees i n a n unthi nned stand o f the same i n i t i a l s i te i ndex. S a l vage o f morta l i ty i n unthinned stands would reduce these . l i t tl e . -percentages Y N = number of tree s ; og = average d . b . h . ; G . basal area ; CVTS = cubic volume of total s tem; CV4 = cub i c volume to a 4 - i nch ( IO-cm) top d . i . b . ; IV5 = Interna tiona l board-foot vol ume to a s - i nch ( l 3-cm) top , 1/4- i nch kerf; SV6 = Scribner board-foot vol ume to a 6 - i nch ( 1 S-em) top (Scribner vol ume is the fonnu l a version a pp l i ed to 16- foot l o g s ) . CV4 , I V 5 . and SV6 excl ude vol urne i n the stump. Y P recornne rci a i thinn i ng ga i n i s r ( precoomerc i a l thinning ... conmerc i a l thi nn i ng) - conmer e i a l thinni ng) t conme rc i a l thinni n g . 26 E ff e cts of Al t e r n at e Thj n n i n g Re g i m es M i n imum Merchant ab l e D . b . h . The e f f e ct o f min imum me r ch an t ­ ab l e d . b . h . ( dm ) on t o t a l merchan t ab l e p r o du c t i o n i s gr e a t e r for st ands r e c e i v i n g o n l y commerc ial t h i n n i n g t h an f o r p r e comme r c i a l l y t h i n n e d s t a n d s ( t ab l e 9 ) b e cause t h e l at t er were t h i n n e d t o s t an d dens i t i e s comme n s u r at e w i t h t h i s min imum d.b.h. For s t an ds r e c e i v i n g on Z y c ommer a i a Z t h i nn i n g , merchan t ab l e y i e l ds wi t h a dm o f 7 . 5 in ch e s ( 1 9 em ) a r e l e s s t h an t h o s e wi t h a dm o f 6 . 0 i n c h e s ( 1 5 em ) b y a l l Re l a t ive meas u r e s o f vo l ume . r e du c t i o n s i n y i e l d are great e s t o n s i t e I V and p r o gr e s s ive l y l e s s w i t h i n cr e as i n g s i t e i n d e x . Yield r e duc t i o n s a r e l e s s wh en i n c r e a s i n g p r o p o r t io n s o f t o t a l vo l ume are nonmerchant ab l e ; t h u s , l es s for board- foo t vo l ume t han for cub i c vo l um e . F o r pre a ommer c i a Z Z y t h i nned s t and s , d i f f er e n c e s a s s o c i ­ at e d w i t h dm- - an d , t h us , w i t h s p a c i n g o f t r e e s - - ar e mino r . As Tab l e 9--Effecr; :;j 'ninimwn meraham;ab le ':. CJ . h . (am) o n merchantable yield to specif ed ro ation ages) Db $: e) trea entJ uarious measures of :JoZwne; when am ·is - , S i.nohes Versus 0 . 0 inches Meas ure o f vol umel! Treatme n t , s i te , and age CVTS CV4 IV5 SV6 ABSOLUTE D I F cc R E NC E - - Cub i c feet - - - - Boa rd feet - - Commerc i a l t h i n n i ng a l one : S i te I , 85 years S i te I I , 90 years S i te l I I , 95 years S i te I V , 100 years 994 990 951 879 858 850 813 744 4 , 499 4 , 1 32 3 , 818 3 , 19 5 2 , 958 2 , 450 1 , 98 1 1 , 390 S i te S i te S i te S i te S i te 299 323 324 305 259 198 220 218 199 155 1I - 19 81 -13 - 1 60 - 29 7 -1 ,416 - 1 , 45 3 - 1 , 56 3 - 1 , 72 4 - 1 , 80 1 Precommerc i a 1 thi nni n g + corrrn e rc i a 1 t h i nn i ng : I , 85 years I I , 90 years I I I , 9 5 years I V , 1 0 0 years . V , 1 0 5 years PERCENTAGE D I Fc:RENCES Percent Commerci a 1 thi nni ng a l one : S i te I , 85 years S i te I I , 90 years S i te I I I , 9 5 years S i te I V , 100 years 4.2 5.1 6.4 8.5 3.7 4.6 5.7 7.6 2.8 3.3 4.3 5.6 S i te S i te S i te S i te S i te 1.2 1.6 1.9 2.4 3.0 .8 1.1 1.4 1.6 1.9 0 Precommerc i a 1 th i nn i ng + commerc i a l th i nn i ng : 11 ( IO-cm) ( 1 3-cm ) ( I S-cm) l og s ) . I , 85 years I I , 90 years I I I , 9 5 years I V , 100 years V , 105 years .1 0 - . 2 -.7 2.1 2.3 2.7 3.2 -1.0 -1.2 -1.8 -2.8 -5.3 CVTS = cub i c vo l ume o f tota l s te m ; CV4 = c ub i c vo l ume to a 4 - i nch top d . i . b . ; IVS = I n terna t i ona l board- foot vo l ume to a 5 - i nc h top , 1 / 4- i nch kerf ; SV6 = Scri bner board-foot vo l ume to a 6 - i nch top ( Scri bner v o l ume is the form u l a vers i o n a p p l i e d to 1 6-foot CV4 , IV5 , and SV6 e x c l ude vo l ume in the s tump . Di fference i s y i e l d when dm (mi n i mum mercha n ta b l e d . b . h . ) = 6 . 0 i nches mi nus yi e l d when d m = 7 . 5 i nches . Percent i s th i s d i fference d i v i ded by the forme r . 1I Nega t i ve d i fference means y i e l ds a re greater when dm = 7 . 5 i nc h e s ( i . e . , wi th wi der spac i ng ) . 27 wi t h s t ands rece i v i n g o n l y comme r c i a l t h i nn i n g , reduc t i o n s 7.5 c ub i c v o l ume y i e l d when dm i s i n c h e s ( 1 9 cm ) i n s t e a d o f 6 . 0 ( 15 site cm ) increase with index . In t e rms vo l ume i n mer c han t ab l e t r e e s wh i c h die prior t o thinn ing . Th i s l o s s is part l y o f f s e t b y s a l v a g e o f in inches decreas i n g some o f t h e s e d e a d t r e e s when t h e f i r s t t h i nn i n g i s made . An addi­ of t ional , S c r ib n e r t re e s ; the howeve r , p r o du c t i on t h e s t and w i t h f ew e r r e l at ive improveme n t increases with b ut may The del a y e d Th i s Th i n n i n g maj o r Tabl e thinning is l o ss is the of a sma l l , is i l l us t ra t e d for a site t ab l e 10 . examp l e , gains i n m e r c h an t ab l e CVTS due to In comme r c i a l th is thinning 10 - - Effeat on usab le production (CVTS) of de laying aOlTTTleraial thinning in a site II n a t ura l s tand w h e n m i n imum merahantab le d. b . h . i8 6. 0 inohes Stand age (yea rs ) Stand and i tem 30 Unthi nned s tand : Cumu l a ted mo rt a l i ty Cumu l a ted growth 0 0 " Captured" mo r t a I i ty 0 T h i nn d at age Net ga i n Adj usted net ga i Th i nned at age 0 0 ! 51 I - - - - C u b i c feet - 439 3 . 09 3 1 . 534 6 . 82 2 I 67 3 . 032 1 1 . 09 3 90 Percent of poten t i a l ga i n - 4 . 953 1 6 . 088 792 439 193 1 . 534 426 3 . 032 693 4 . 953 1 . 006 3 . 94 7 3 . 155 0 0 n1! 100 39 : Y 390 Cumu l a ted " ca ptured" morta l i ty Cumu l a ted " l os t " growth Net ga i n Adjusted n e t ga i Th i nned at age I 39 30 : Cumu l ated " c a ptured" mo r ta l i ty Cumu l a ted " l os t " growth 0 1 . 485 317 2 . 983 680 4 . 904 1 . 105 3 . 79 9 3 . 007 nY 95 51: Cumv l a ted "ca ptured" morta l i ty Cumu l a ted " l os t " growth Net ga i n Adjusted net g a i 890 o 2 . 388 491 4 . 309 1 . 066 3 . 24 3 2 . 451 nY 78 11 Adj us ted net ga i n = d i fference between captured morta l f ty and " l os t " potent i a l growth mi nus sa l va b l e morta l i ty at fi na l harvest in unthi nned s tands . Y Assumes that s a l vage at fi r s t commer c i a l th i n n i ng · 80 perce t of tota l lO-year mercha n ta b l e morta l i !l t diD ra t i o at f i rs t commerc i a l th i nn i n g . diD i s the Thus , ra t i o o f o f cut trees to ug of a l l merchantab l e trees present before. thi n n i n g . diD 1 .0 .9 .8 28 due progre s s i ve l y l on ge r de l ay r e s u l t in s ub s t an t i a l l o s s es . s t an d i n loss caused by comme r c i a l smal l e r , i s made . The net e ffect s h o r t de l a y may b e q u i t e d e c r e as i n g s i t e qual i t y . De l a ye d F i r s t but t o a gre a t e r r e du c t i o n in growth pe r acre o n ce t h e f i r s t t h i n n i n g v o l ume ( SV6 ) , is g e a t e r in Percent s a I vab I e 80 89 100 II - - - ----. .. - . .- . - - - - - . - - - . --- ��------ .------ are 9 5 percen t as gr e at whe n th i n n i n g i s d e l a y e d 9 y e a r s ( t o age 3 9 ) as i f it wer e done at t h e f i r s t poss ib l e a g e ( age 3 0 ) . If the f i r s t t h i n n i n g i s d e l a y e d 2 1 y e ars ( t o a g e 5 1 ) , g a i n s a r e o n l y 7 8 p e r ce n t a s g r e at a s i f i t w e r e d o n e at a g e 3 0 . Frequency and S e v e r i t y o f T h i n n i n g With o u r a s s ump t i o n s , d i f f er e n c e s in r e s i dual s t o ck i n g l e v e l ( wi t h in s u g ge s t e d l i mit s ) an d / or f requen cy of t hinn ing h ave no e f f e c t o n t o t al cub i c volume p r o duct ion . Thus , o t he r r e g im e s a l t er o n l y t h e d i s t r ib ut ion o f h ar v e s t e d vo lume o v e r t im e and For amo n g var ious sizes of t re e s . the a l t e rnate r e g imes d i s c us s e d p r e v i o us l y under numb e r an d s iz e o f t r e e s ( f ig . 2 2A ) , d i f f e r e n c e s i n t o t a l produ c t ion b y a l l me a s u r e s o f D i f f er e n c e s vo l um e are n egl i g ib l e . i n vo l ume at f i n a l harvest in t h e s t an d ma i n t a i n e d at t h e l ow · average stocking l e ve l is 4 t o 7 p e r c e n t great e r t han in t h e s t an d main t a i n e d at t h e h i gh average s t o ck i n g l ev e l . C o r r e s pondin g l y , s t an ds w i t h t h e great est vo l um e a t t ime o f f in a l harve s t have a l i t t l e l es s vo l ume r emo v e d in t h in n i n g s . Type o f Thi n n i n g L ikewi s e , t y p e o f t h i n n i n g i s assumed to h ave no e f f e ct on t o t a l cub i c vo lume product ion , an d i t h a s o n l y a s l i ght e f f ect on t o t al p r o ­ duct i o n b y o t h er me asures o f vo l um e . I f t h e d/D r a t i o p ro gr e s s e s f rom 0 . 9 to 0 . 8 r at h e r t h a n f rom 1 . 0 t o 0 . 8 ( f i g . 2 2B ) , t o t a l p r o du c t i on by al l me asures o f vo l ume r ema i n s The amo un t cut vir t u a l l y t h e s ame . at each t h i n n i n g i s a l i t t l e l e s s - ­ e s p e c i a l l y a t t h e f ir s t two t h i n n i n gs - - s o vo l ume rema i n i n g at f in a l harve s t i s a l i t t l e g r e at e r . Th i s d i f f e r e n ce in f i n al harves t vo l ume is , howeve r , l es s t h an 3 p e r c ent . Extrapolation to Other Stand Conditions S t ructure and devel o pment o f mos t n a t ural s t ands d i f f e r f r om t h a t o f t h e average n a t u r a l s t an d wh i c h w e have d e s c r ib e d . Some s t an d s a r e u n d e r s t o cked and others o v e r ­ s t o c ke d , r e l a t ive t o t h e aver age . S ome a r e very uni f orm and o t h e rs e xt reme l y var i ab l e in compo s i t ion . I n some s t an ds , the p at t ern o f h e i gh t growt h r e l a t ive to s t and a g e i s qui t e d i f f e ren t f rom what we have as s um e d - - s o s i t e index does not r emain const ant . We do not at t emp t to s p e c i f i c a l l y est imate the e f f e c t s o f t h e s e var i a t i on s , b u t i n t h i s s e c t i o n w e b r i e f l y discuss t h e i r p o s s ib l e e f f e ct s . U N I F O R M LY U N D E R ST O C K E D STA N D S A n un d e r s t o cked s t and i s one i n wh i ch aver age d . b . h . ( D g ) is sma l l er t h an NORMAL ( f i g . 4 ) for the numb e r o f trees present . Th e d e g r e e o f under s t o cking decreases as t h e t r e e s grow . A u n i form Z y un d e r s t ocked st and has adequat e crop t r ee s - - t r e e s t hat w i l l re ach merch a n t ab l e s i z e - -but f ewer e x c e s s t re e s t h an h as a n aver age s t an d . Th i s may b e due t o e i ther nume rous s m a Z Z o p e n i n gs w i t h in the s t and or a un i formly mis s in g s i ze compon ent of sma l l ( general l y younger ) t r e e s . I f , f or examp l e , a s t and has 1 , 0 0 0 t r ee s per acre ( 2 , 4 7 0 per ha ) w i t h we l l - d i st r ib ut e d crop t r e e s , it w i l l Zik e Z y develop in a manner s im i l ar t o that shown in f i gure 2 9 . The s t a n d w i l l reach MAXI MUM at a Dg o f about 4 . 3 inches ( 1 0 . 9 cm ) . At about t h i s t ime , the smal l e r t r e e s w i l l b ecome supp r e s s e d and some w i l l die ; t hus , t he st and wi l l s l owly approach NORMAL through a comb in a t ion o f growth and mo n : a l i t y . Wh en t h e s t an d with 1 , 000 t rees p e r a c r e ( 2 , 4 7 0 p e r h a ) reaches ;\OlAX I MUM , t h e l ar gest 4 0 0 t r e e s per acre ( 99 0 per ha ) w i l l have a Dg of abo ut 5 . 1 i n c h e s ( 1 3 . 0 cm ) ; the l at t er mus t t h en grow to 8 . 0 inch e s ( 2 0 . 3 c m ) at the age o f " f i r s t po s s ib l e " comme r c i a l t h innin g . The s e 4 0 0 t r e e s per acre ( 99 0 p e r h a ) wi l l , have grown w i t h l e s s t h an average compe t i t i o n dur i n g part o f t h e i r l i f e and , thus , reach t h i s s iz e at a youn ger age . By the t im e t h e s t a n d is ready i N' comme r c i a l t h i n n in g , it wi l l f o r a l l pract i c a l purp o s e s h ave the char ct e r i s t ics of a f u l l y s t o cked s t an d . 29 300 200 " " ;: 1l. :1 w [ U " [ W Q " W U) <1 ill 100 80 50 50 40 <i 30 20 2.000 1 .000 800 NUMBER 600 OF 400 TREES PER 300 200 100 ACRE Fi gure 29 . --Typical devel opment of a uni forml y understocked stand . With our assumpt ions , the e f f e c t s o f this p a s t under s t o c k i n g on merchant ab l e y i e l d shoul d b e r e f l e c t e d en c ir e l y i n t h e age and s i t e index at t ime o f the f i r s t commer c i a l t h inn i n g . Therefor e , once t h i s age and s i t e index are known or e s t imat e d , one can p r e d i c t f u t u r e de ve l opment o f the m e r c h a n tab Z e port ion o f the s t an d . Each year gained i n age at f i r s t p o s s ib l e com­ mercial t h i nn ing w i l l add about 1 t o 2 percent t o merch an t ab l e vo l ume y i e l d . The maj o r gain i n age wi l l o ccur prior t o the t ime t he s t and reaches MAXI M11M- -wh e n crop t r e e s h ave l e s s than normal compe t i t ion ; there wi l l b e a s l i ght add i t ional gain t h e r e af t e r , s i n c e t r e e s w i l l be youn ger th an normal for t h e s i t e . The gain , r e l a­ 30 t i ve t o t h e normal t ime r e q u i re d , wi l l de c r e a s e as numb e r of t r e e s increases . W i t h 1 , 00 0 t re e s p e r acre ( 2 , 4 7 0 p e r h a ) , we e x p e c t a g a i n o f about 6 . 5 p e r c en t . Thi s means a gain o f about 1 ye ar o n t h e b e s t s i t e s an d u p t o about 2 . 5 years on poorest s i t e s , for s t ands havin g t h e s ame current s it e i n d e x . No t a l l uni f o rml y un d e r s t o ck e d s t an ds wi l l , however , b e h ave in t h i s I f t r ee s ize is very un i ­ mann e r . f orm , due t o min imal var iat ion in f a c t o r s cont r ib u t i n g t o var i at ion i n s ize--age , genotype , an d m i c ro s i t e--the s t an d may not t h i n i t s e l f adequat e l y a f t er r e a c h i n g MAXI MUM . I t wi l l then b ecome o v e r s t o c k e d , w i t h s ub s e quent det r i ­ men t al e f f e c t s o n s t and d e v e l opmen t . ------ -------- - -------"---- O V E R ST O C K E D STA N D S An o ve r s t o ck e d s t an d i s o n e in wh i ch Dg i s l arger t h an NORMAL f o r the number o f t r e e s p r e s e n t . One po s s i b l e cause o f t h i s h a s a l r e ady been not e d . I f a s t a n d is ve r y u n i f o rm , t h er e w i l l b e l i t t l e c rown c l a s s d i f f er e nt i at i o n ; t h u s , f ew t r e e s w i l l b e l o s t t o mort a l i t y and most t r e e s wi l l cont inue t o incre ase i n s i ze-- at a r e du c e d rat e ( f ig . 1 5 ) . When s u b s t ant i a l mor t a l i t y f i n a l l y d o e s o c cur in s u c h s t ands , i t t e n ds to remove f a i r l y l ar g e groups of t r e e s r at h e r t han s c a t t e r e d i n d i v i du a l s ; t h i s c r e a t e s l arge ope n i n gs and r e s u l t s i n a l os s o f Thus , some o f t h e crop t r e e s . growt h p e r acre is s u b s e q u en t l y Z e s s than t h a t o f t h e average na t u r a l s t an d o f the s ame s i t e and age . Exces s i ve o v e r s t o c k i n g may a l s o c aus e a r e du c t ion in h e i gh t . growt h . Gains f rom p r e comme r c i a l t h i n n i n g in s u c h a s t an d woul d , thus , b e g r e a t e r t h an g a i n s f rom t h i n n i n g an average n a t u r a l s t and o f the s ame s i t e an d age . Anot h er po s s i b l e c a u s e o f overst o c k i n g h a s much l e s s impact on s ub s equent s t an d d e v e lopment . Even in s t an ds whe r e s t and s t r u c t u r e c l o s e l y appr o x imat e s t h a t o f ave r a ge s t an ds , mor t a l i t y i s e r r at i c i n s t e a d o f f o l l owin g a smo o t h t r en d as i n average s t an ds . Commo n l y , a f ew years may p a s s in wh i c h t h e r e i s l i t t l e mort a l i t y ; supp r e s s e d t r e e s r emain a l i v e ( a l t h o u gh t h e y d o not grow ) , caus i n g t h e s t an d t o b e come overst ocke d . Then a drought o r other s u c h c l imat i c var i ab l e w i l l t r i gg e r h e avy mort a l i t y i n o n e year . Thi s remo v e s t h e accumul at ion o f supp r e s s e d t r e e s wh i c h " norma l l y " might h ave d i e d p r e v i ou s l y and , l ik el y , a l s o some wh i ch normal l y might n o t d i e unt i l l at er . Th i s e r r at i c l o s s o f suppr e s s ed t re e s wi l l , g e n e r a l l y , h ave l i t t l e e f f e c t on devel opment o f t h e r e s t o f t h e s t an d . N O N C O N STA N T S I T E I N D E X I f h e i gh t growt h d o e s n o t f o l l ow t h e a v e r a g e pat t ern r e l a t i ve to s t a n d age , s i t e i n de x wi l l not b e a const ant . Thus , i t is not uncommon f o r s i t e i n d e x t o i n c r e a s e or d e c r e a s e as t h e s t an d b e comes o l der . If s i t e index i s i n c r e a s i n g ( r e l at ive t o our s ) , vo l ume growth p e r a c r e wi l l b e greater t h an w i t h a const ant s i t e i n de x ; if s i t e in dex i s d e c r e as ing , vo l ume growt h w i l l b e l es s . Th e approxima t e magn i t ude o f t h e e f f e c t c a n b e e s t im at e d . F o r e xamp l e , i f s i t e index i n c r e a s e s f rom 1 70 t o 1 8 0 a s t h e st and grows f rom ages 3 0 t o 6 0 , we e s t ima t e that t o t al cub i c vo l ume growt h d ur i n g t h i s p e r i o d wi l l be a.bout 1 3 p e r c ent g r e a t e r t h an i f s i t e i n d e x r ema i n e d a t 1 70 , a n d about 5 p e r c e n t g r e a t e r t h an i f i t h a d always b e e n 1 8 0 . I R R E G UL A R S TA N D S O t h e r t h an t h e s e r a t h e r s y s t em­ a t i c d e v iat i o n s f rom the ave r a g e natural s t a n d , t h e r e a r e i r r e g u l a r i ­ O n e such t i es i n s t an d compo s i t i on . irregu l a r i t y i s the p r e s enc e o f Z arg e open i n gs w i t h i n t h e s t an d which are nonproduct ive . Th e r e i s , howeve r , a w i de gray ar e a b e t w e e n th e s e l ar g e o p e n i n g s and t h e sma l l open i n gs d i s c us s e d p r e v i o us l y . Trees b o r de r i n g on open i n gs wi l l grow mo re rap i d l y than i f t h e y we r e comp l e t e l y s urrounded b y comp e t i t o rs , but not eno ugh to o f f s et the m i s s i n g crop t r e e s . Each s u c h s i tuat i o n i s un i que and must b e s o evaluat e d . Anoth er i r r e gu l a r i t y i s t h e presence of other species . All the f o r e g o i n g a s s umes p u r e Dougl a s - f i r . I f t h e t r e e s w i t h i n a s t an d i n c l ud e o t h er s pe c i e s , b o t h y i e l ds an d re comme nded s p a c i n g o f t re e s m i gh t b e qui t e d i f f erent . The e f f ec t s o f i n c l us io n s o f o t h e r s p e c i e s an d other i r r e gu l ar i t i es in s t a n d com­ pos it ion are beyond the s c o p e o f t h i s p ap e r . Literatu re Cited Bruce , David , Don a l d J . DeMars , an d Don a l d L . Reuk ema . 1977 . Dou g l as - f i r man ag e d y i e l d s imul a t o r --DF I T - - us er ' s g u i d e . USDA For . S e r v o Gen . Tech . Rep . PNW- 5 7 , 2 6 p . P a c . No r t hwe s t Fbr . a n d Range Exp . St n . , Port l an d , Or e g . 31 McAr d l e , Richard E . W a l t e r H . M e y e r , and Dona l d Bruce . 1961 . The y i e l d o f Doug l as - f i r in t h e Pac i f i c orchwe s t . [ . S . Dep . Agric . Te ch . BUl l . o . 2 0 1 , 74 p . Wash i ng t on , D . C . . Re inek e , L . H . 1933. Perfect ing a s t an d- de n s i t y index for e ven- aged f o r e s t s . J . Agric . Res . 4 6 : 6 2 7 -6 3 8 . Reukema , Don a l d L . 1 9 70 . Fo rty-year deve l opmen t o f Doug l a s - f i r st ands p l ant e d a t various spaCings . USDA For . Serv o Res . Pap . PNW- I O O , 2 1 p . Pac . No rthwest :or . and Ran g e Zxp . St n . , Po r c l and , O re g . Reukema , Don a l d L . 1 9 72 . Twen c y-one y e ar deve l o pment o f Dougl aS- f i r s c ands r e p e a t e d l y 32 t h inned a t vary i n g int erval s . USDA For . Serv o Res . Pap . PNW- 1 4 l , 23 p . Pac . Nor t hWest For . and Range Exp . St n . , Por t l an d , Or e g . Reuk ema , D o n a l d L . 1 9 75 . Guide l in e s for pre com­ merc i a l t h i n n i n g of Dougl as- f i r . USDA Fo r . Serv o Gen . Tech . Rep . PNW- 3 0 , 1 0 p . Pac . NorthWe s t For . and Range Exp . St n . , P o r t l an d , Ore g . Reukema . D o n a l d L . , an d Leon V . P i enaar . 1973 . Y i e l d s with and without r e p e at e d commer c i a l t h in n ings i n a h i gh - s i t e - qua l i t y Dougl as ­ USDA For . S e r v o f ir s t an d . Res . Pap . PNW- 1 5 5 , 1 5 p . , i l lus . Pac . Northwest For . and Range Exp . S t n . , Port l an d , O r e g . A p pendix GLO S SARY Ag e : tot a l a g e f r o m s e e d . Ave r age d . b . h . ( D g ) : D i ame t e r at b r e a s c he i ght o f t r ee o f average b a s a l area . May r e f e r t o e i t h e r t o t al s t an d or a s p e c i f i e d com­ ponenc of the st an d . Average n a t u r a l s c a n d ( n atural s t an d ) : A stand h av i n g t h e c h a r a cc e r i s t i c s d e s cribed in t h e s e c t ion enc i t l e d " n a1: ural s t an ds . " The s e s t an d s may a l so b e c al l ed " n o rmal " s 1: ands . Crop t r ee s ( merch an 1: ab l e compon e n t ) : Trees wh i c h have at t ained , o r are e xp e c t e d 1: 0 a t t a i n , a spe c i f i e d d.b.h. A curve Cumu l a t e d- s c and curve : s howing t h e cumu l a t e d cont r ibu­ t ion o f l ar ge s 1: t r e e s t o t o t al n umber an d b a s a l a r e a per acre as s u c ce s s i ve l y sma l l e r t r e e s are added . Cub i c vo lume o f t o t a l s t ems , CVTS : i n c l ud i n g S 1: ump and t i p . CV4 : Cub i c v o l ume o f s t ems e x c l u­ s i v e o f ( 1 ) t i p above a 4 - i n ch d . i . b . an d ( 2 ) s t ump be low 1 foot o r o n e - t e n t h o f t o t al h e ight , wh i ch e v e r is l ower . d(D r at io : The r a t i o of average d . b . h . of cut t r e e s c O average d . b . h . of all mer ch an t a b l e t r e e s b e f o r e t h i n n in g . DF I T : A s t an d s imul ator con s t r u c t e d t o provide de t ai l e d output r e ­ g ar d i n g man aged- s t an d y i e l d . Dg : S e e " av e r age d . b . h . " ( mi n imum merch an t ab l e d . b . h . ) : The smal l e s t d . b . h . at wh i ch a t r e e is c o n s i de r e d t o be o f merchan t ab l e ( mark e t ab l e ) s i z e . dm G : B a s a l a r e a per a c r e ; may r e f e r t o e i th e r t o t al s t an d or spe c i f i e d c omponent o f t h e s t and . Gro s s growt h ( growt h p e r a c re ) : A c t u a l growt h o f a l l t re e s i n c l ud e d at s t art o f p e r i o d , n o t r e d u c e d b y mor t a l i t y . May r e f e r to the t o t a l s t an d o r t o sp e c i f i ed com­ pon e n t o f t h e s t an d . Gross p r o d u ct ion ( gr o s s y i e l d ) : Tot al vo l ume , o r b a s a l a r e a , produced by t h e s t an d s in c e i t s origin , including that l o s t to May r e f e r mor t a l i t y o r c ut t i n g . t o e i t he r t o t a l s t an d o r t o spe c i f i e d me r ch a n t ab l e compon en t . Gross y i e l d : S e e " gro s s p roduc t ion . " IV5 : I n t e r n at i o n a l b o a r d - f o o t vo l ume of st ems e x c l us i ve of ( 1 ) t ip above 5 - i n ch d . i . b . and ( 2 ) s t ump a s f o r " CV4 " ; 1 ( 4 - i n ch k e r f . M.a.i. : Me an ann u a l i n c remen t . MAXIMUM : De f in e d b y f i gu r e 5 a n d cor r e s p o n d i n g t ex t . ( 1 ) Dg , N , and G at wh i ch t h e maximum numb e r o f t r ee s l ar g e r t h an a s pe c i f i e d d . b . h . i s r e a c h e d i n aver age nat u r a l s t an d s ; ( 2 ) the approxi ­ mat e max imum s t o ck i n g l e v e l t o whi ch a given number o f merchant­ ab l e t r e e s s ho u l d b e grown i n a man a g e d s t an d . Merch an t ab l e comp o n en t : t re e s . " S e e " cr o p Merchan t ab l e . . . : P e r t a in s t o mer ­ chan t ab l e comp o n e n t o f s t an d ; dep e n d e n t on measure o f v o l ume . Min imum merch an t ab le d . b . h . : " dm . " See N : Numb e r o f t r e e s p e r a c r e ; may r e f e r to e i t h e r t o t al s t an d o r spe c i f i e d compon en t o f t h e s t an d . Natural s t an d : s t an d . S e e ave r a g e n a t u r a l 33 Net y i e l d : Excludes vo l ume l o s t t o mor c al i t y ; may r e f e r t o e i t h e r t o t al s t and or s p e c i f i e d merchan t ­ ab l e component . NORMAL : D e f i n e d b y f i gure 4 and corresponding t ex t . A s t and­ den s i t y i n de x ; D g , N , and G of t o t al average n at ur a l s t an d . RECOMMENDED RES I DUAL ( or RECOMMENDED ) : The min imum s t o c k i n g l ev e l t o whi ch t h e merchan t ab l e p o r t ion o f a s t an d wi l l b e reduced a t e ach t h i n n in g . S i t e index : Ave r a g e h e i gh t o f domi ­ nant and codom i n an t t re e s at age 1 0 0 , i f h e i gh t g r owth f o l l o ws the normal p a t t ern . S ubmer ch an t ab l e t r ee s : Tre e s wh i ch are exp e c t e d t o d i e without at t a i n ­ i n g the min imum merchan t a b l e d . b . h . SV6 : S c r ibner b o a r d- f oot vo l ume o f s t ems e x c l u s i v e o f ( 1 ) t ip above 6 - i n ch d . i . b . a n d ( 2 ) s t ump a s f o r " CV4 . " VA R I A B LE S A N D R E LATI O NSH I PS E XA M I N E D P l an t a t i on P r e comme r c i a l t h inn i n g Comme r c i a l t h i n n i n g Cub i c vo l ume grow t h ( 6 V ) Poten t i al 6 V f ( s i t e , age ) Reduced 6 V w i t h t h i n n i n g Produc t i on b y t o t a l s t an d Product ion b y s t an d compone n t s V f ( h e i gh t ) Height growt h ( 6H ) Impro ved 6H w j e ar l y s pa c i n g 6 o f t o t a l s c an d 6 D g o f sc an d compone n t s f ( H ) ; over t im e Dg f ( H ) at g i ven t imes D.b.h. Dg , N , G of t o t a l s t an d Dg , N , G o f s t an d component s Cumul at ed- s t and curve Mort al i t y , t o t al s t an d Mor t a l i t y , s t an d compone n t s S t a gn at i on = = = 34 SAMPLE JFIT P,OG C O NT R O L DrA THE CARD ETER I hI S P E C I FI E S THA T OF UH E Q C H A NT A6 L " E RC N A " , A 8 l E 0 8 H THE AT AGE H A P VE S 1 = H A R V E S T S T U_ F A G E = l T H l h N I h G STUHP AG = I D I SC O UNT + OO T O T AL CODOM GE YEARS 35.9 H E I G HT 36 R A T E: · f N 1) 8 1I NC Jo E S FE E T 73.5 I '! R S T PaSS 73.7 TOT A L 8 E F CRE: STANO CO P G hHIT S UB I1 E R C f< M ER C H A N T A B L E " E R C H t >: A V E C UT T OT A L OF 4 . ,J g '; . 0 1 B • J I 5.C2 6 .73 'Y PEP THOUSAND T HO U S A N D "''= A t-. SO FT . 26 73 VCL CU q 7 . ;; 2 VCL /G RATIO . 1 303 . I ' 2 3 . l3 2 G . 2 6 87 .35 02 34 97 • 3 5 04 . 24 7 1 • 7 . 58 1 0 . 20 1 0 . 1 9 1 0 . 2 1 .ee 2S. I? "QRT G . O Wl' H ,E R C H A N T A e L E CUT CH L E A V, TOT AL L E AVE S U I1 G"OSS SU SUM "ERCH OF CUT VOL GRO 'H SUBHfRCH AT t l FT CU 28 . 2 0 ' 484 1 29. 1 4 ' . 1 03 " 29. 1 3 ' ?7 . 3 5 " .. 4 8 26 1 3 67 2 7 J6 1474 • •d2 9 . 27 " . J' 1 0 7 9. I 9 • I e , a 9.3 • . I 2 E6 6 d7 . 1 3S3 . 5535 .4602 . ;; 9 g .47 17 3. CO 15.71 , 5 f: 18. ! t 'G . I ;; • 04 J2 o" e" ' 2322 504 3 .51 · 529 1 2 3 . 72 25 . ;4 " 34 . ; 33. J0 30 .. • , ;8 ... 34 3 I -1' 2 4 15 5046 1 34 3 37 0 3 39 4 8 7 1 63 27 1 I OPT 4.9 1 2 . 9 5.21 • I 3 54 . 5C3 . 1 4 80 2. .3 25.34 " 02 27 . 1 6 " 32.S4 • • ' . 74 G R A I. O P O S S I SL E FT 4 - I NCH CU ' " - I N(,H FFF VOL U H E CO ERC I A L ACRf TNTL 114V 286 1 163 46 86 E - I NC H T H I N N I NG SC R n V TRHS • 8 <) 1 6 3E 14 1 20 " 2409 285 484 1 4 1 01 4 8 56 1 57 "699 1 23 " 3 4 f' , 3 E 22 SqO 6 3 5"3 1 66 1 2 89 1 6 <; 5 3 3 2 9 67 I I U79 I I J 74 50 4 a9 16 8916 5 ·;:T 1 29 7 252 4 222 25042 6 ;t 8 1 87 0 3 I en6 3 1 3 2'\ Q <; 7 5 1 1 8 72 1 0? 1 2 1 &349 5 HARVEST , 94 9 ' 5 9 49 1 6E I 2 2438 6 290 - E R C H A N T A 8L E I 1 0 . I F IR S T 718 64 1 4 T O T AL 9 E F O R F. SU8HERC H 00 T HE IS 8 . 00 INCHE S . a OA P O F E F T SCR I e N E R . B O A RD E El S C R I E N ER . ' 2 9 . I '" l A dlES 1 3 f>7 T O T A L B E F O R F. SU8 HFOCH HE 7.02 MEAN YIE D N I NG . GUT S U q hf R C H VOL 9 1 .6 L FA V E 7 . 10 1 20 . 0 0 EA' S . PfF T"t GROSS fRCH SUM SUM SU 46 60 0 . • 60 1 50 . 0 ' DFI T P R T OF T F S TANQ IS 6 . 0 0 :NCH S. OT M A C A P ECC"HERCI AL THE STANO HtS T H E T H I N N I NG I T E R V A L I S 1 0 A 'S . N U M 8 E R OF C O " E R C I t T H I N N I N G S = 2. T O T AL P R I N T O UT A N A GfD O O U G . A S- F 1 R THE AM 3 7 7 5'< 177'3 4 1 77 0 5 1 77 0 5 4 22e 1 3477 64 1 . 5 A C R£ SG FT 639. 0 17 1 . 7 402. 1 1 34 . 2 140.8 46 . 9 236.8 • G 'Y FER PE" A C R£ 2 6 7 . <: S 04 . 7 1 4. 2 3 gJ.9 1 24.7 9. E 22 1 . 9 I a .. "3' "0 . 3 108 . 0 1 17 .6 1 539 37 356 2656 507 .. I Jq5 . 2 226 87 . 3 17 1 . 6 3 0 349 2 0 1 .8 303 9 1 80.3 168.4 180.3 I E6 . 4 3 0 349 1 46.9 69.0 2 0 n7 2 2 0 672 7 1 95 1 48 3 283 1 1 40 . 8 2 1 . 3 249 . 2 CU PRICE 7 17 I .7 1 57 . 9 1 3 4 77 P R IC E D I SC .9 167.9 87. E 180.3 pe I C E 17 I .4 33 6 . 9 267 . 9 CURR£NT 2 1 • S 3.2 I 1 24 1 7 64 T Ol' A l S Y ?.I SUM SUH GROSS "PCH SUH O' Basal B o a rd CUT a re a . • I 00 2 927 0 9270 9 I 8I e 1 29 8 7 2 ., 5 0345 49 26 4 96 2 6 J7544 37 544 7 5"4 402. 1 2e6 . 5 2 5 .6 2 55 .6 fe e t . f: I I 35 w: o T JT A L C 0 1 0 t< ; GE YEARS 36 €A N STAND ET COMPONENT 74 gTAL 92 g EF O R E , , 0 cu I N C HE S ' 7.02 " 8. C I · · 2 e F C R F. 9 . 27 A N T A 9L E , 0 . 0 7 · 8 E F O R :: 1 2.49 13.09 · T O T AL HfRC · 03H ER CHHTA9LE 46 T O T AL ERCHA N 1 A 8 L E · VOLUHE C U FT 4 6 '< 1 4 1 0J 529 I 5046 6646 6560 " " - H C >! CU FT , - I NC H 6 - I HCH • If" ' 2 89 ' 6 · 6 39 . 0 4 2. I 31 R AC R E I N TL b14 40S4 ! l 4Y 1 7 4 84 25264 , 7705 · 25042 1 77 0 5 · E349 6 2 S ry 3H2 37 1 54 " 0 l49 10349 · SIn r O T AL ..\ G E YEARS 36 • MEAN OeH COD0H . :: n HE I G H T 73.7 . Z6 d 7 • 1 3 03 .3502 7 . 58 4 .82 . 1 2 E6 6.79 . 25 1 3 S U8 '1 E R C H 9. 1 3 5.05 • H E R C H A N T A aL 9.7 1 . 5 1 42 1 7 . 44 33. 92 S US H E R C H 4.9 7.63 . 1 3 54 . 3 1 76 3 . 44 2 5 . 39 ' 1 0 . 6 1 33. 4 1 ' TOTAL HERCHA NT A elf S U O H E R C H > O DT HERe H TOTAL HERC M 60 I 1 0 . I .U Y B o a rd Basal O RT TCT A L OqT H O Q, S U B "ERCM HERCHA n A R E I 1 . 69 5 . 2 1 1 2 . 0 5 SO • 4 54 2 I 393 . 7458 . 1 480 . 79 2 5 3. 1 2 1 0. 2 0 .oo VCl / G Y. RATIO • 2 , q2 .. FT 718 2 S .. 2 0 • 48 4 1 29. 1 4 • " 103 .9 1 57 . 9 • B 0.3 171 .6 1 68 .4 267 . 9 14e 20 I I · • T A ?L E ( lOO - YEAR BAS I S ) 4 - I N CH CU GY P '.: J? ACH 17 1 .7 1 40.a A C P {' VOL U H E 7.02 4 . 09 8 . 0 1 I NC H E S C O HPO N E N T G r ST A I>I O " X = 1 4 0 . 0 HA VCl CU F T SU3 HfQCH 9I .6 "ON 2/ FT STANO S CRI R Y ' · 89 1 4E 48S6 N A TUP A L DOH T Q F' f S P I' '' • I GHT F Conti nued SAMP L E O F I T P R I N TOUT S U 'i A P ¥ ., E L · CU FT 4 0 5" 440 3E 14 P"F I GY ACRE ; N i 1 7 4 84 72 1 66 1 2 4 y. Y FER CU "E N T C RE "RICE $ S C FT n6.B 89 1 6 402. 1 1 40 . 8 21 .3 23.72 1If, 504 292 5 25 1 67 . 9 29. 25 ' 295 245 1 0 " 40.2 I .. 2 2 , 3.5E 33. 5 1 5 . 5<; 246 • • 1 63 917 a1 2 0 967 430 . 9 69.0 195.7 209 7 36 1 . 9 186. 1 1 39 3E97 ' " d56 1 f l S9 · 84q4 59 8 100 1 0 . I 30740 1 02 3 8 . 74 27 . J3 17 1 . 7 3 0492 B08 3d.90 .9 09. 0 7.35 • 3 8916 47.5 6.4 e6.4 27 . 5 37072 2 97 . C 12 1 . 5 3 . 2 37072 27 5 . 5 1 16.3 1 3 37 3 14 5 1 756 556 1 1 37 3 fee t . a r ea . GPO 36 1 337 9.6 2 0 64 2 1 . 5 O I SC DRICE 999-537 I· I T h e mission of the P A C I F I C N O R TH W E S T F O R E ST A N D R A N G E E X P E R I M E NT STATI O N is to provide the knowl. edge , technol ogy, and a l te rn atives fo r present and futu re p rotection , m anagem ent, a n d use of fo rest, range , and rel ated envi ron m e nts . W i th i n th is o v e r a l l m i s s i o n , the Statio n condu cts a n d sti m u la tes rese a rch to fac i l i tate and t o accelerate progress toward the fol l ow i n g goals: 1. P rov id i n g safe a n d efficient tech n ol ogy for inventory, protecti o n , and use of resou rces. 2 . Developing and evalu ating alte rnative methods and levels of resou rce management . 3 . Ach ieving opti m u m sustai n e d resou rce pro d u ctiv ity consistent w i th m a i n ta i n i n g a h i gh q u a l ity forest environme nt. The area of resea rc h encompasses O rego n, Wash i n gton, A l aska, and, i n some cases, Californi a, H aw a i i , the Western States , and the N ation. R es u lts of the resea rch are made available p romptly . Proj e ct headqu arters a re at: La Grande, O regon Portl and, O regon O l y m p i a , Wash i n gton Seattle, Washi ngton W enatche e , W ashi ngton A nchorage, A l as k a F a i rbanks, A l as k a J u neau , A l aska B e n d , O regon Corval l i s , Oregon Mailing address : Pa cific Northwest Forest and Range Experiment Station P. O. Box 3 1 4 1 Portland, Orego n 9 7208 The F O R EST S E R V I CE of the U.S. D e pa rtment of Agricultu re is dedicated to the principle of multiple use manage m e nt of the N ation's forest resou rces for sustained yields of wood, 'watei< 'forage'>i.liidlife, and recreation. Through forestry research;, ' cooperation with ' the States and private forest owners, and ma nagement of the N ational F orests a nd' N ational G rasslands, it strives - as d i rected by Congress: - to ' p rovide increasingly greater service to a growi ng N ation. ; :: ' " " -" ;-�", " , The U.S. D e partment oLAgriculture is .an" E q ua l ;Opportunity E m pl oyer. A p p l i ca nts for all D epartrTlent progra ms ,will be given equal considerati on without regard to age, race,' col r;,sex, religior ,or" 'national ori g i n . \ . '" /..I: .i:./>f ::. · " ' ----... -