Biology360:Evolution Spring,2016 Instructor: Office: OfficeHours: Email: Lecture: Text: LectureNotes: CourseWebsites: VanessaKoelling,Ph.D. Thompson257C F,10:00AM-1:00PM,orbyappointment vkoelling@pugetsound.edu MWF,9:00AM-9:50AM,TH197 Evolution(1stedition).CarlT.BergstromandLeeAlanDugatkin. W.W.NortonandCompany.2012. ProvidedonlineafterclassasaPDF. AMoodlesitewillbeusedtodistributelecturenotes,labsand othermaterials.http://moodle.pugetsound.edu/ COURSEDESCRIPTIONANDLEARNINGOBJECTIVES Evolutionisanelectivecoursecoveringthemajorprinciplesofmodernevolutionarytheory. Thiscoursewillexaminethemechanismsofevolutionandhowithasshaped,andcontinues toshape,thediversityoflifeonEarth.Whenyoufinishthecourseyoushouldhaveathorough groundingin: -howtheprocessofevolutionoccursinpopulations -majortransitionsinthehistoryoflifeonEarth -howevolutionisstudiedatboththemicroandmacroscale -importantconceptswithinmajorsubfieldsofevolutionarybiology Thiscoursewillalsogiveyoutheopportunitytofurtherdevelopyourabilitytoreadprimary scientificliteratureandcommunicateinwritingandotherforms. RESPONSIBILITY Iwillfacilitateaneffectivelearningenvironmentinwhichyoucanexplorethefundamentals ofevolution.Noquestionisunreasonable.However,ultimatelyyouareresponsibleforyour owneducation.Youmustdedicateampleenergyandtimetolearnthematerialonyourown. Youareresponsibleforobtaininginformationonanyschedulechangesaswellasonclass materialsyoumaymissduringabsencefromclass. TIMEREQUIREMENTS Pleasebeawarethatassessmentinthiscoursewillbebasedoncomprehensionand applicationofbiologicalknowledge.Assuch,youwillneedtoplanonSIGNIFICANTadditional timeoutsideoflecturesothatyoumaymovebeyondsimpleacquisitionofknowledge. EMAIL Iwillregularlyuseemailtosendoutannouncements,changesinthesyllabus,reminders abouttestsorduedates,etc.IwilluseyourPugetSoundemailaddress.Itisyour BIO360,Spring2016,V.A.Koelling 1 responsibilitytocheckthisemailaccountregularlytokeepup-to-datewiththese announcements. Iwillrespondtoemailwithin24-48hours. READINGS Thisisanupper-divisioncourseinwhichyouareexpectedtoacquirecontentknowledgeof greaterdepthanddetailthaninlower-divisioncourses;therefore,youareexpectedtodoa closereadingofthetextbook.Followingeachlecture,itisimportantthatyouintegratethe informationinthetextwiththatfromlecture.Irecommendthatyouusetheglossaryand practiceproblemsforeachchapter.Theseaidswillhelpyouidentifyimportantconceptsand termsassociatedwiththetopicsandhelpyouorganizeandapplyyourlearning. Youshouldalsoexpectsupplementalreadingassignments(e.g.,newsarticlesorscientific journalarticles).Assignmentsmaybeassociatedwiththisreadingmaterial.Inallcasesyou shouldconsiderthereadingmaterialrequiredunlessIstateotherwise. CLASSPARTICIPATION Iexpectyourregularattendanceandattentioninclass.Participationduringclassorlab discussionsofprimaryliteratureisrequired.Youwillearnpointsbasedonyourparticipation inpaperdiscussions. EVOBEAKERLABASSIGNMENTS YouwillbeassignedanumberofEvoBeakerLabsfromSimBio.Thesearesimulation experimentsbasedonrealworldexamplesofevolution.Theyaredesignedtohelpyou understandfundamentalevolutionaryprinciplesandwheretheycomefrom,aswellas practicesolvingproblemsinevolutionarybiologyusingthescientificmethod.Wewilluse somein-classtimefortheEvoBeakerLabs,butyouwillalsobeaskedtocompletesomeon yourownoutsideofclass.AttheendofeachEvoBeakerLab,youwillbeaskedtocomplete gradedquestions,awritingassignment,orboth. YoumustpurchaseasubscriptiontotheSimUTextforthecoursethatcontainstheEvoBeaker Labs.Thecostis$40.Ifyouwishtousefinancialaid,youmaypurchasearegistrationvoucher atthePugetSoundbookstore.Youwillthenneedtoregisterviaemailinstructionsfromme. PAPERS Youwillwrite1-2shortpapers.Furtherinstructionsforthesepaperswillbegiveninclass. Ifyouneedassistancewithresearchforthesepapersorotherassignments,pleaseconsultthe ScienceLibrarian.Heisavailableforone-on-onemeetingsandeagertohelpstudentsfindthe informationtheyneedintheCollinsLibrary.Hiscontactinformationisasfollows: BIO360,Spring2016,V.A.Koelling 2 EliGandour-Rood,ScienceLibrarian CollinsLibrary,Room1117 Phone:253-879-3678 Email:egandourrood@pugetsound.edu EVOLUTIONVIDEOPROJECT Yourfinalprojectinthiscoursewillbetomakeavideo(3minutesoralittleless)that explainsatopicinevolutionarybiology.ThisassignmentisbasedontheEvolutionFilm Festivalthattakesplaceeveryyearattheannualprofessionalconferenceforevolutionary biologists(theEvolutionmeeting).Examplevideosandtheguidelinesformakingthevideo arefoundhere:http://filmfestival.nescent.org/ Youhavetheoptionofworkingaloneorinpairsforthisassignment. EXAMS Therewillbetwomid-termexamsandafinalexam. Iwillnotdiscussanexamorexamgradewithin24hoursofreturningthegradedexamtoyou. Ifyouwishtodisputethegradeorfurtherdiscusshowtoimproveyourstudyhabits,Iwillbe happytomeetwithyouafterthe24-hourtimeperiod—onceyouhavethoroughlyread throughyourexamandthepostedanswerkey. RegradingPolicy:Ifyoufeeltherehasbeenamistakemadeingradinganexamproblem,you maysubmititforaregrade.Turninyourexamtomewithawrittenexplanationofwhatyou thinkiswrong.However,pleasebeawarethatwewillnotrevisithowpointsareassignedfor questionsthatreceivedpartialcredit.Regradesmustbesubmittedwithinoneweek(7calendar days)ofthequiz/exambeingreturned. GRADING(subjecttochangeatdiscretionofinstructoratanytime) GradingPolicies o NOMAKE-UPEXAMSWILLBEGIVEN.However,ifyouaresickorhavesomeother validreasonformissingtheexam,pleasecontacttheinstructor.Whilemake-up examsarenotgiven,wemaybeabletoworksomethingout. o Lateassignmentswillbepenalized10%perdaylate. o Noextracredit. GradingScale A≥94% A-=90-93.9% B+=87-89.9% B=84-86.9%B-=80-83.9% C+=77-79.9% C=74-76.9%C-=70-73.9% D+=67-69.9% D=64-66.9%D-=60-63.9% F<60% BIO360,Spring2016,V.A.Koelling 3 Componentsoftheclasswillbeworththefollowingpercentagesofyourtotalgrade: Exams 60% EvoBeakerLabs 20% Papers/Assignments 10% EvolutionVideoProject 10% APPROPRIATEBEHAVIOR Studentsareexpectedtobehaveinanappropriatemannerwhileattendingthisclass.During classtimeIexpectyoutobeworkingonmaterialsforthiscourseonly.Pleaseberespectfulto othersintheclass,especiallyconcerningtheuseofpersonaltechnology: • Switchyourphonestovibrateorno-ringbeforecomingtoclassandhavethemstowed inyourbagorpocket • Noemailortextmessaging • Youmaynothaveearbudsinatanytimewhileinclass • Digitaldevices(laptops,tablets,phones)canonlybeusedbypermissionofinstructor • Youmaynottouchoruseanyelectronicdevicesinanywayduringanexam • Youmaynotleavetheroomduringanexam STATEMENTCONCERNINGACADEMICMISCONDUCT AsastudentattheUniversityofPugetSound,youassumetheresponsibilitytoexhibitinyour academicperformancethequalitiesofhonestyandintegrity.Iwillnottolerateacademic misconductinanyform,especiallyplagiarism. Ifyouneedtorefreshyourmemoryaboutthenatureofacademicmisconductanduniversity policiesonthehandlingofacademicmisconduct,youmaygohere: http://www.pugetsound.edu/student-life/personal-safety/student-handbook/academichandbook/academic-integrity/ Plagiarismisdefinedaspresentingtheworkofanotherasone'sown.Morethanfour consecutivewordsfromasourceotherthanthewriterconstitutesplagiarismwhenthe sourceisnotclearlyidentifiedinappropriatedocumentationformat. Ifyouarefoundtohavecommittedplagiarisminthisclassyouwilllose100pointsor thevalueoftheassignmentthatwasplagiarized,whicheverisgreater.Inaddition,Iwill followthestepsforhandlingacademicmisconductoutlinedontheuniversitywebsiteabove. Ifyoudidnotreceiveinstructiononavoidingplagiarisminyourfreshmanseminarcourses, pleasetakesometimetocompletethecoursethatthelibraryhassetupforyou.Forthat course,gohere:http://research.pugetsound.edu/academicintegrity SPECIALACCOMMODATIONS Ifyouhaveaphysical,psychological,medicalorlearningdisabilitythatmayimpactyour coursework,pleasecontactPeggyPerno,DirectoroftheOfficeofAccessibilityand Accommodations,105Howarth,253.879.3395.Shewilldeterminewithyouwhat BIO360,Spring2016,V.A.Koelling 4 accommodationsarenecessaryandappropriate.Allinformationanddocumentationis confidential. CLASSROOMEMERGENCYRESPONSEGUIDANCE Pleasereviewuniversityemergencypreparednessandresponseprocedurespostedat www.pugetsound.edu/emergency/.Thereisalinkontheuniversityhomepage.Familiarize yourselfwithhallexitdoorsandthedesignatedgatheringareaforyourclassandlaboratory buildings. Ifbuildingevacuationbecomesnecessary(e.g.earthquake),meetyourinstructoratthe designatedgatheringareasoshe/hecanaccountforyourpresence.Thenwaitforfurther instructions.Donotreturntothebuildingorclassroomuntiladvisedbyauniversity emergencyresponserepresentative. Ifconfrontedbyanactofviolence,bepreparedtomakequickdecisionstoprotectyour safety.Fleetheareabyrunningawayfromthesourceofdangerifyoucansafelydoso.Ifthis isnotpossible,shelterinplacebysecuringclassroomorlabdoorsandwindows,closing blinds,andturningoffroomlights.Staylow,awayfromdoorsandwindows,andascloseto theinteriorhallwaywallsaspossible.Waitforfurtherinstructions. BIO360,Spring2016,V.A.Koelling 5 TentativeScheduleforBIO360,Spring2016 SubjecttochangeatanytimeatthediscretionofDr.VanessaKoelling Week/dates LectureTopics Readingfromthe Textbook Week1 Introduction/Evolutionary Ch.2,3 1/20-1/22 ThinkingBeforeandAfter Noclass9/18(MLKJr.Day) Darwin/NaturalSelection Week2 NaturalSelectionandEvolutionary Ch.3 1/25-1/29 Experiments Week3 PhylogenyandEvolutionary Ch.4,5 2/1-2/5 History/InferringPhylogeny Week4 InferringPhylogeny/OriginofLife Ch.5,11 2/8-2/12 Week5 MajorTransitions/Extinction Ch.12,15 2/15-2/19 Week6 TransmissionGenetics Ch.6,7 2/22-2/26 Review/PopulationGenetics Week7 PopulationGenetics Ch.7 2/29-3/4 Week8 EvolutioninFinitePopulations Ch.8 3/7-3/11 Week9 Noclass3/14-3/18 (SpringBreak) Week10 NeutralTheoryofMolecular Ch.8,10 3/21-3/25 Evolution Week11 EvolutionatMultipleLoci Ch.9 3/28-4/1 Week12 Species,Speciation,andHuman Ch.14 4/4-4/8 Evolution BIO360,Spring2016,V.A.Koelling EvoBeakerLabsand Exams DarwinianSnails& Experimentingwith Snails HowtheGuppyGotIts Spots DomesticatingDogs PaperDiscussion:2/12 Exam1:2/15 MendelianPigs/HardyWeinbergandKuru PaperDiscussion:3/11 GeneticDriftand BottleneckedFerrets HIVClock FinchesandEvolution Exam2:4/4 Week13 4/11-4/15 Week14 4/18-4/22 Week15 4/25-4/29 GenomeEvolution/Evo-Devo Ch.10,13 PaperDiscussion:4/15 EvolutionofSex/SexualSelection Ch.16,17 EvolutionofSociality Ch.18 Week16 5/2-5/4 Noclass5/6 Week17—FinalExam Wednesday,5/11,8:0010:00AM Examwillbecumulative Coevolution Ch.19 EvolutionVideo ProjectDue5/4 BIO360,Spring2016,V.A.Koelling