Development and Evaluation of Conservation Tillage for Squash in Western Oregon by Brandon M. Reich A THESIS submitted to Oregon State University in partial fulfillment of the requirements for the degree of Bachelor of Science Presented June 10, 1996 Commencement December 1996 ABSTRACT Squash varieties of Golden Delicious and Squash Hybrid NK 530 were grown in disk-till, strip-till, and no-till conservation tillage systems following a cover crop of'Monida' oats (Avena sativa L.) and common vetch (Vicia sativa L.). Tillage systems were evaluated for effects on squash emergence and yield, weed density and biomass, and the time it took to hand hoe the growing area. Squash emergence was similar among treatments and squash yield was similar between strip-till and disk-till systems. However, propane flaming appeared to increase squash emergence in strip-till treatments and reduce emergence in no-till treatments. Weed abundance was lower in strip-till and no-till treatments where a cover crop mulch remained on the soil surface than in disk-till treatments where the soil was less covered. Weed biomass in no-till treatments, where a cover crop mulch lay on the surface, was reduced for up to six weeks after squash planting. However, once the mulch broke down, weeds became abundant and no-till plots were abandoned one month prior to harvest because of concerns over contamination of weed seeds into the soil. This one year study suggests that squash grown in a strip-till conservation tillage system with a cover crop has adequate weed control and equivalent yields with a disk-till system and that squash grown in a no-till system has similar emergence and inadequate weed control at this amount of mulch. TABLE OF CONTENTS Page INTRODUCTION ........................................................................................................1 Benefits and Challenges .................................................................................2 Vegetable Crop Establishment and Yields .....................................................6 Objectives of this Research ............................................................................7 MATERIALS AND METHODS ...................................................................................8 Cover Crop Management and Soil Preparation ..............................................9 Planting ........................................................................................................... 9 Weed Management .......................................................................................10 Squash Harvest ..............................................................................................10 Sampling ........................................................................................................10 Data Analysis .................................................................................................11 RESULTS AND DISCUSSION .................................................................................12 CONCLUSIONS ........................................................................................................14 BIBLIOGRAPHY .......................................................................................................16 LIST OF FIGURES Page Figure 1 Percent squash emergence by treatment ...................................................20 2 Weed abundance in-rows ...........................................................................21 3 Between-row weed abundance (by species) .............................................. 22 4 In-row weed abundance (by species) ......................................................... 23 LIST OF TABLES Table Page 1 Effects of tillage treatment on weed abundance and biomass ................ 24 2 Effects of tillage on the time it took to hoe the squash growing area ...... 25 INTRODUCTION World wide, 8.5 million ha of agricultural land are lost to erosion every year (Oregonian, 1996). A 1982 National Resources Inventory estimated that croplands in the United States lose to erosion an average of 16.4 Mg/ha at a cost, estimated by the Conservation Foundation, of $2.2 billion (Brady, 1990). Growers and researchers in Oregon have developed soil erosion management plans in the past and continue to develop management strategies, including whole-farm strategies and management practices for vegetables. One of the most common and widely developed practices developed to reduce soil erosion is conservation tillage. Conservation tillage has been defined as any set of soil management practices that leaves at least 30% of the available residue on the surface (Dickey et al., 1992; Pierce, 1985; Reeder, 1992). Originally, researchers thought that this was a minimum amount of residue that would protect the soil from erosion. Tillage systems that meet the 30% criteria include: disk-till, strip-till, ridge-till, and no-till systems. As researchers, growers, and others began evaluating other factors contributing to erosion (such as soil compaction) the definition changed. Conservation tillage systems are defined now by the unique sets of benefits and challenges these systems share in common (Cook and Robertson, 1979; Dickey, 1992; Reeder, 1992). Conservation tillage systems are generally associated with increased water holding capacity of the soil, improved soil structure, reduced soil 2 erosion, increased amounts of organic matter, and greater densities of soil arthropods. However, a number of challenges toward adoption of conservation tillage systems have been identified. Cook and Robertson (1979) identified that conservation tillage systems must be created specific to an area's soil types, weather patterns, and equipment availability. A survey in Tillage for the Times identified growers' major concerns about the adoption of conservation tillage practices: 48% of growers were concerned about inadequate weed control, 21% identified yield reductions as a major concern, and 15% expressed concerns about managing excessive crop residue. Similarly, Nowak (1983) argues that growers' concerns are the major obstacle to adoption. Benefits and Challenges Conservation tillage systems usually include a cover crop that is planted the fall before spring planting of the main crop. Frequently used cover crops include cereal grains, legumes, and cereal/legume mixtures. Often it is not harvested and, instead, is allowed to remain on the surface as a mulch or partially incorporated into the soil. Growing a cover crop and allowing it to remain on the surface as a mulch increases the water available to the main crop. The mulch acts as a barrier to water evaporating from the soil (Blevins, 1983; Koskinen and McWhorten, 1986). The mulch also reduces the severity of runoff events. For example, Jones et al. (1969) measured the amount of water leaving areas of soil with and without a 3 mulch as a percent of the precipitation. He determined that 27% of the precipitation onto soil without a mulch became runoff water, while only 5% of the rainfall onto soil with a mulch became runoff water. Finally, improved soil structure increases the amount of water in the soil. The mulch influences soil compaction by protecting the soil from the impact of rainfall that can lead to compaction (Triplett et al., 1968). Under a mulch, soil is often found with a greater number of micropores, small airspaces that hold most of the soil's water (Froud-Williams et al., 1981). Blevins et al. (1971) grew corn in conventional tillage and no tillage systems and found soil moisture higher in the no tillage system, with effects most profound in the top 0-8 cm of soil. Jones et al. (1969) determined the soil moisture increased under a mulch to 30 cm compared to soil without a mulch. Conserving moisture in the soil for the main crop can be a major factor in increasing yields, especially in areas where crops are not irrigated (Triplett et al., 1968). Conservation tillage practices also reduce soil erosion. Koskinen and McWhorten (1986) determined that applying as little as one ton of wheat straw reduces soil losses to water runoff to nearly zero. They attributed this reduction in soil loss to (1) the mulch reducing the severity of rainfall impact, thus improving water infiltration; and (2) the mulch slowing water runoff and retaining soil particles from the water. They observed this benefits of a mulch on reduced soil erosion at rates as little as 0.7 Mg/ha and 1.4 Mg/ha (1/4 tons/ac and 1/2 tons/ac). Even at the lowest mulch rate of 0.7 Mg/ha, they observed a 75% reduction in soil loss compared to the soil loss on soil without a mulch. 4 Increased amounts of organic matter are also associated with conservation tillage systems. Blevins et al. (1983) determined that in corn grown in different tillage systems for ten years, there was twice as much organic matter in the first 5 cm of the no-till treatment compared with the conventional tillage treatment. Haines and Uren (1990) similarly found more organic matter near the surface of direct drilled systems than conventionally cultivated systems. Koskinen and McWhorten (1986) attribute increases of organic matter in reduced tillage systems to both the addition of plant biomass and to organic matter not being lost through erosion. Plant material can be introduced in conservation tillage systems through incorporating or disking the cover crop or crop stubble. Increased amounts organic matter, improved soil structure, and reduced soil disturbance creates beneficial habitat for microorganisms, bacteria, and earthworms. Haines and Uren (1990) found greater earthworm densities, more than twice as many in the top 10 cm, in direct drill systems compared to conventionally cultivated systems. However, the high moisture content of soils in conservation tillage systems, along with high amounts of organic matter levels, may create anaerobic conditions where bacteria can cause denitrification (Blevins, 1983; Doran, 1980). This effect may be, however, offset by nitrogen contribution from incorporated legume cover crops. When cover crops are incorporated into the soil, they break down in the soil, can contribute nutrients, including nitrogen, to growing main crops. By comparing the yields of corn grown with fertilizer and cover crops, Mitchell and Teel (1977) estimated the nitrogen contribution of the incorporated cover crop to 5 the main crop. They estimated that a cover crop of oats or rye contributes 55 kg N/ha to corn in a no-till system; a cover crop of hairy vetch or crimson clover contributes 112 kg N/ha; and a combination of grain and legume contributes 180 kg N/ha. Similarly, Wagger (1989) estimated the nitrogen contribution to corn in a no-till system of a rye cover crop at 25 kg N/ha and a legume cover crop of crimson clover and hairy vetch at an average of 40-45 kg N/ha. He estimated that 36% of the nitrogen in the crimson clover and 30% of the nitrogen in the hairy vetch cover crop was returned to the corn crop. Nova (1995) estimated that cover crops of legume and mixtures of legumes and grains contributed 90 kg N/ha equivalent to broccoli in a disk-till system. Conservation tillage systems reduce weed abundance in a number of ways: through allelopathic cover crops, through shading weed seeds under a mulch, and through non-disturbance of weed seeds in soil. Research has identified allelopathic chemicals from cover crops that inhibit germination of seeds, thus reducing weed densities (Putnam et at., 1983). Mohler and Calloway (1992) identified reduced weed seed disturbance in the soil as the reason that lower densities of weeds were found in no-till treatments than in conventional tillage treatments. They also identified the presence of a mulch on the soil surface as a factor further reducing weed densities. Studies have shown that a mulch reduces weed germination by reducing sunlight infiltration through the mulch and by physically blocking seedling emergence (Teasdale et al., 1991; Teasdale, 1993). They determined that soil covered with a much on 42% of its surface will have lower weed density. 6 However, because seeds are not disturbed and stimulated to germinate, the seeds remain in the soil (Forcella and Lindstrom, 1988; Froud-Williams et al., 1983). On the other hand, high water content of soils in conservation tillage systems with a mulch, have been found to stimulate weed seeds (Buhler and Mester, 1991). A number of researchers have identified the weed species composition in conservation tillage systems, especially no-till systems and systems with a mulch, to shift from summer annuals and broadleaves to winter annuals and perennials (Blackshaw et al., 1994; Buhler et al., 1994; Cussan, 1975; Mohler and Calloway, 1992; Teasdale et al., 1991; Wrucke and Arnold, 1985). Shifts toward winter annuals may occur because winter annuals grow better than summer annuals in the cooler, moister soils often found in conservation tillage systems, especially under a mulch. Also, winter annuals may be able to better compete in low light conditions under a mulch. Shifts towards perennials may occur because of reduced cultivation that would control perennials. A number of researchers have identified the need for herbicides in conservation tillage systems, especially no tillage systems, to control weeds (Cussan, 1985; Koskinen and McWhorten, 1986; Triplett and VanDoren, 1974). Vegetable Crop Establishment and Yield The main factors affecting crop establishment in conservation tillage systems, especially no-till systems, are increased water content and reduced temperatures in the soil under a mulch (Knavel et al., 1977; Weston, 1990). Large seed size and rapid germination are two factors that may predict success of crop 7 establishment (Weston, 1990). Lima beans and squash have been established successfully in no-till systems (Beste, 1973; Mullins et al., 1980; NeSmith et al, 1994). Some studies have found that cucumber (Knavel et al., 1977) and snap beans (Mullens et al., 1988) establish poorly. However, most studies have found that tomatoes, peppers, cucumber and snap beans establish as well or better in no-till systems than in conventional tillage systems (Beste, 1973; Knavel et al., 1977; Morse and Seward, 1986; Mullins et al., 1980; NeSmith et al, 1994; Weston et al., 1990). Vegetable crops with good stands grown in a no-till system that includes a pre-plant or a pre-emergence herbicide, generally have as good or better yields than conventional tillage systems (Abdul-Baki, 1993; Bellinder et al., 1984; Bennett, 1975; Beste et al, 1973; Loy et al., 1987; Masiunas et al., 1995; Morse and Seward, 1986; Mullins et al., 1980; NeSmith et al., 1994; Skarphol et al., 1987). Wilhoit et al. (1990) determined that cabbage grown in a strip-till system had greater yields than no-till or conventionally tilled systems. Some researchers identified poor weed control as the main factor contributing to reduced yields in no- till systems, even when an herbicide was used (Knavel et al., 1977; Knavel and Herron, 1981; Loy et al., 1987; Mullins et al., 1988). Objectives of This Research The following research project was conducted to assess effects, if any, of disk-, strip-, and no-till systems on squash production in western Oregon. The 8 effects of tillage systems on squash stand and yield, weed density and biomass, and hoeing times were evaluated. MATERIALS AND METHODS A cover crop of 'Monida' oats (Avena sativa L.) at 56 kg/ha and common vetch (Vicia sativa L.) at 50 kg/ha was seeded on 24 October 1994 in a 0.4 ha plot at the Oregon State University Vegetable Research Farm, near Corvallis, Oregon. Seed was drilled in 15 cm row spacing using a Nordsten® grain drill. 'Monida' oats was chosen because of apparent weed suppression qualities reported by Nova (1995). Common vetch was selected for the mixture because it is a nitrogen-fixing legume and was expected to contribute nitrogen to the squash. A split-plot, randomized complete block experiment was established in the spring of 1995, with three tillage systems, disk-till, strip-till, and no-till. Treatments plot size was 12 m by 15 m and four rows of squash were planted in each plot on 1.5 m centers. Each main treatment tillage plot of four rows of squash was divided into: two squash rows with weed control (flaming, cultivating, and hoeing) and two squash rows with no weed control. Each two row weed control subplot measured 6 m by 15 m. Between blocks, in 7 m strips, buckwheat was seeded at 56 kg/ha to maintain soil cover and to suppress weeds during the growing season. 9 Cover Crop Management and Soil Preparation In the disk-till treatments, the cover crop was flailed on 17 May 1995. On 18 May 1995 plots were disked twice and field harrowed twice with a Lely Roterra® power harrow. Five days before planting (1 June 1995) plots were again field harrowed once. In the strip-till treatments, 56 cm wide strips in the cover crop were mowed using a rotary lawn mower on 28 April 1995. Two weeks before planting (21 May 1995) mowed strips were tilled three times at an 8 cm depth with a rototiller. Plots were tilled again one week before planting (1 June 1995) using a tractor-mounted rotovator. Three days after planting (9 June 1995) cover crop regrowth was killed with glyphosate at 3.6 kg a.i./ha. In the no-till treatments, the cover crop was killed with glyphosate at 3.6 kg a.i./ha one month before planting (3 May 1995). No additional soil preparation occurred. Planting A mixture of Golden Delicious and Hybrid NK 530 Squash (50:50 by volume) was planted 6 June 1995 at 6 seeds/row m on 1.5 m centers. A modified no-till "slot planter" was used to plant all plots and liquid fertilizer was banded at the time of planting at the rates of 56 kg/ha (N) and 191 kg/ha (P). Additional nitrogen fertilizer (70 kg/ha) was surface applied five weeks after planting (14 July 1995). 10 Weed Management The squash rows in the weed control plots were flamed ten days after squash planting (but prior to squash emergence) with a propane hand torch delivering 2 kg/cm2 (30 p.s.i.) of propane and the operator walking approximately 5 km/hr. Disk-till and strip-till treatments were cultivated five weeks after planting with a tractor mounted sweep cultivator. Strip-till treatments were only cultivated in the mowed strips. Disk-till and strip-till treatments were hand-hoed on 13 July, 23 July and 8 August 1995. The time it took to hand hoe each of the 29 m2 squash growing areas was timed with a stop watch and averaged over treatments Squash Harvest A harvest area of 29 m2 in the weed control subplots, were hand harvested on 10 and 11 October 1995. Entire plots were weighed on truck scales accurate to 9 kg. No-till treatments were flailed on 14 August 1995 to minimize weed seed production and contamination of the research plots and, therefore, were not harvested. Sampling Cover crop biomass was estimated by collecting and sorting all above ground material in four 0.25 m2 quadrats in each plot (3 May 1995). Oats, vetch, and weeds were sorted and placed in paper bags. Samples were dried at 49 °C for 83 hrs and dry weights determined. The percent cover of the soil by cover crop was estimated by recording the presence of residue at 30 cm intervals along a 11 diagonal across the middle of each plot. Percent soil cover was estimated in the disk-till plots on 19 June 1995 and in the strip-till and no-till plots on 12 July 1995. Squash seedlings was estimated for each tillage treatment plot one month (4 July 1995) after planting and compared to the number of seeds planted to estimate percent seedling emergence. Rows that had been flamed for weed control were compared with rows that were not flamed to estimate any effects of flaming on squash emergence Four weeks after planting (5 July 1995) weed density was estimated in all treatment plots. Within each treatment, two 0.25 m2 quadrat samples were randomly taken between rows of squash, from inside the rows of squash in weed control subplots, and from inside the rows of squash in the no-weed control subplots. Weeds were identified and counted. Rows that had been flamed for weed control were compared with rows that were not flamed to evaluate any effects of flaming on weed density. One week later (12 July 1995), samples from four 0.25 m2 quadrats were taken from the rows of squash in weed control subplots were clipped and dried at 67 °C until dry and dry weight determined. Data Analysis All weed abundance data were square root transformed for analysis and means LSD separated. Other data not transformed and means LSD separated. I considered p-values up to p = .21 significant enough to report because I was willing to risk a Type II error in order to separate differences between means that are large enough to be significant to growers, but not scientifically significant. 12 RESULTS AND DISCUSSION Cover crop biomass was, on 3 May 1995, was: 'Monida' oats, 2,355 kg/ha; common vetch, 2,341 kg/ha; and weeds, 283 kg/ha. Weeds were less than 6% of the total biomass. The cover crop was approximately 60 cm high at this time. Prior to planting, percent soil covered by cover crop residue was: disk-till, 63%; strip-till, 95%; no-till, 89%. Cover crop growth occurs rapidly in the spring and because the no-till treatments were killed early, they had less biomass covering the soil at planting time. This difference was evident at approximately six weeks after planting when the cover crop cover began to decay, whereas the cover crop on the strip-till treatments still remained intact. Pre-emergence flaming caused an apparent increase in squash emergence in strip-till treatments (p = .04) and an apparent decrease in no-till treatments (p = .15) (Figure 1). This was an unanticipated effect and further research is needed to determine whether flaming squash rows affects emergence differently among tillage treatments. There was no significant difference in wet weight harvest yields between treatments: disk-till, 28 tons/ac; strip-till, 31 tons/ac. Both yields are comparable to typical commercial squash yields in western Oregon. Both no-till and strip-till treatments, with the mulch residue on the soil surface, had fewer weeds between squash rows than the disk-till treatment (p = .11) (Table 1). Weed density between rows was greater in the disk-till treatments than in strip-till and no-till treatments for the four most abundant weed species 13 sampled: common groundsel (Senecio vulgaris L.), common purslane (Portulaca oleracea L.), hairy nightshade (Solanum sarrachoides Sendtner), and redroot pigweed (Amaranthus retroflexus L.) (Figure 3). No hairy nightshade was found in either strip-till or no-till treatments; no common purslane was found in strip-till treatments; and no redroot pigweed was found in no-till treatments. Within squash rows, there was a significant interaction between treatment and weed control effects (p = .01). Strip-till plots had fewer weeds (p = .06) when flamed than when non-flamed (Figure 2). When strip-till treatments were tilled, the soil was wet and tillage was difficult; the soil formed large clods that were not consistent with other treatments. Perhaps, because the soil was open and exposed, weed seedlings were exposed to the flaming and weed densities were lower in flamed strip-till than in non-flamed strip-till treatments. Further research may determine whether this interaction is a result of the disk-till treatment responding differently to flaming than other treatments. Within rows, weed density by species responded differently depending on whether the rows were flamed (Figure 4). Common groundsel and common purslane responded similarly to tillage and flaming treatments. Hairy nightshade was generally unaffected by tillage or flaming treatments, expect in no-till treatments where flaming increased seeding abundance. Redroot pigweed had low abundance in all tillage and flaming treatments. Dry weight biomass of weeds in-row at six weeks after planting differed among treatments (p = .09). Less biomass was found in the no-till treatment, which had a mulch residue around the rows of squash (Table 1). This result is 14 consistent with Mohler (1991) who found a reduction of weed biomass with a rye cover crop. Strip-till treatments required less time to hoe (140 person hours/ha) than the disk-till treatments (199 person hours/ha) (p = .21). Because different teams hoed different treatments at different dates, it is difficult to accurately attribute differences in hoeing times to tillage treatments. However, a difference of nearly 50 people hours/ha is an important economic difference and it is demonstrated throughout the season (Table 2). CONCLUSIONS Tillage did not have an effect on emergence of squash seedlings; however, flaming did effect emergence in strip-till and no-till treatments. The tillage system did not affect crop yields; yields of both disk-till and strip-till were similar to and consistent with growers' yields of squash in western Oregon. The mulch residue in the strip-till and no-till treatments reduced weed density for approximately four weeks compared with the disk-till. The mulch residue in the no-till treatment reduced weed biomass, compared to disk-till and strip-till treatments for approximately six weeks. This time period for weed control is consistent with the research of Putnam et al (1983) who recorded adequate weed control under a mulch for four to six weeks. Further research is needed to evaluate the effects of flaming within different tillage treatments on weed density. 15 Results of this one year study indicate that a grower could expect weed control from a mulch residue for four to six weeks and expect yields from both the strip-till and disk-till system to be consistent with commercial squash yields in western Oregon. 16 BIBLIOGRAPHY Abdul-Baki, A.A. and J.R. Teasdale. 1993. A no-tillage tomato production system using hairy vetch and subterranean clover mulches. HortSci. 28(2):106-108. 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A systems approach to conservation tillage. Chelsea, Michigan:Lewis Publishers. Putnam, A.R., J. DeFrank, and J.P. Barnes. 1983. Exploitation of allelopathy for weed control in annual and perennial cropping systems. J. of Chem. Ecol. 9(8):1001-1010. Reeder, R. 1992. Making the transition to conservation tillage. 3-7. In: Midwest Plan Service. Conservation tillage systems and management. Ames, lowa:lowa State Univ. Skarpol, B.J., K.A. Corey, and J.J. Meisinger. 1987. Response of snap beans to tillage and cover crop combinations. J. Amer. Soc. Hort. Sci. 112(6):936941. Teasdale, J.R. 1993. Interaction of light, soil moisture, and temperature with weed suppression by hairy vetch residue. Weed Sci. 41:46-51. 19 Teasdale, J.R., C.E. Beste, and W.E. Potts. 1991. Response of weeds to tillage and cover crop residue. Weed Sci. 39:195-199. Tillage for the times. Agrichem. Age. June, 1983. 30. Triplett, G.B., Jr. and D.M. VanDoren, Jr. 1974. Agriculture without tillage. Sci. Amer. 236:28. Triplett, G.B., Jr., D.M. Van Doren, Jr., B.L. Schmidt. 1968. Effect of com (Zea mays L.) stover mulch on no-tillage corn yield and water infiltration. Agron. J. 60:236-239. Wagger, M.G. 1989. Cover crop management and nitrogen rate in relation to growth and yield of no-till com. Agron. J. 81:533-538. Weston, L.A. 1990. Cover crop and herbicide influence on row crop seedling establishment in no-tillage culture. Weed Sci. 38:166-171. Wilhoit, J.H., R.D. Morse, D.H. Vaughn. 1990. Strip-tillage production for summer cabbage using high residue levels. Applied Agricultural Research. 5(4):338342. Wrucke, M.A. and W.E. Arnold. 1985. Weed species distribution as influenced by tillage and herbicides. Weed Sci. 33:853-856. Pumpkin emergence flame no-flame a 70 b 40 disk-till strip-till no-till tillage treatment Figure 1. Percent squash emergence by treatment, sampled 4 July 1995. Data analyzed as percent of squash seeds planted and means LSD seperated. Letters above bars indicate significant differences between means (within a tillage treatment): disk-till flame v no flame, p =.55; strip-till flame v no flame, p =.04; no-till flame v no flame, p =.15. Weed abundance in-rows 300 a flame 250 ® no-flame 200 150 b b a 100 50 0 disk-till strip-till no-till tillage treatment Figure 2. Weed abundance in-rows, two samples, four replicates per mean, sampled 5 July 1995. Data square root transformed and means LSD seperated. Letters above bars indicate significant differences between means (within a tillage treatment): disk-till flame v no flame, p =.01; strip till flame v no flame, p =.01; no-till flame v no flame, p =.03. 22 Total weed densities between rows 40 purslane 35 ® niteshade o pigweed 30 ® groundsel 25 zero counts 20 15 10 5 0 disk-till strip-till no-till tillage treatment Figure 3. Between-row weed abundance (by species), two samples, four replicates, sampled 5 July 1995. 2 Pigweed Pursiane 20 20 flame b 18 flame ® no-flame D = .16 N 16 E ® no-flame E15 014 m CL CL 0> 12 c 010 m 0 I- 8 d .0 E6 c 4 b 2 0 0 disk-till strip-till no-till 20 no-till strip-till disk-till tillage treatment tillage treatment Groundsel Nightshade 60 flame flame ® no-flame 50 a 40 no-flame 15 p=.14 10 p=.02 30 20 5 10 0 disk-till strip-till tillage treatment no-till disk-till strip-till tillage treatment no-till Figure 4. In-row weed abundance (by species), two samples, four replicates, sampled 5 July 1995. Data square root transformed and means LSD seperated. Letters above bars indicate significant differences between means (within a tillage treatment). 24 Table 1. Effects of tillage and weed control treatments on weed abundance (two samples, four replicates, sampled 5 July 1995) and biomass (four samples, four replicates, sampled 12 July 1995). Data square root transformed and means LSD seperated. weed abundance weed dry between rows biomass in-rows (number/m2) (g/m2) disk-till 232 a 104 a strip-till 19 b 128 a no-till 30 b 19 b p-value 0.11 0.09 Tillage treatment 25 Table 2. Effect of tillage on the time it took to hoe the 37 m2 squash growing area (sampling dates listed). Means LSD separated. time to hoe (people hours/ha) July 13 July 23 August 8 Season average disk-till 31 a 100 a 67 199 a strip-till 14 b 54 b 72 140 b p-value 0.15 0.06 0.90 0.21 Tillage treatment