NIOZ-RAPPORT 2002 -1
E MACROBENTHIC FAUNA IN THE DUTCH SECTOR
rHE NORTH SEA IN 2001 AND A COMPARISON WITH
PREVIOUS DATA
R. Daan and M. M ulder
Nederlands Instituut voor Onderzoek der Zee
Monitoring Macrozoobenthos of the North Sea
©2002
This report is not to be cited without the
acknowledgement of the source:
Netherlands Institute for Sea Research (NIOZ)
P.O. Box 59, 1790 AB Den Burg, Texel
The Netherlands
ISSN 0923 - 3210
Cover design: H. Hobbelink
VLI Z (vzw)
VLAAMS INSTITUUT VOOR DE ZEE
FLANDERS MARINE INSTITUTE
O ostende - Belgium
THE MACROBENTHIC FAUNA IN THE DUTCH SECTOR OF THE NORTH
SEA IN 2001 AND A COMPARISON WITH PREVIOUS DATA
28782
R. DAAN AND M. MULDER
This report presents data of the monitoring program o f macrozoobenthos in the Dutch
Continental Shelf (DCS) of the North Sea, a cooperation between the National Institute for
Coastal and Marine Management/RIKZ (Rijkswaterstaat), the North Sea Directorate
(Rijkswaterstaat) and the Department o f Marine Ecology (NIOZ)
NETHERLANDS INSTITUTE FOR SEA RESEARCH
Monitoring Macrozoobenthos of the North Sea
NIOZ-RAPPORT 2002-1
CONTENTS
1. SUMMARY............................................................................................................................................................ 1
2. SAMENVATTING................................................................................................................................................. 3
3. INTRODUCTION.................................................................................................................................................. 5
4. MATERIAL AND METHODS.............................................................................................................................. 6
4.1. Sampling................................................................................................................................................. 6
4.2. Sample treatments................................................................................................................................ 6
4.3. Ashfree Dry weight.............................................................................................................................. 7
4.4. Statistics.................................................................................................................................................. 7
4.5. Sediment analysis................................................................................................................................. 8
5. RESULTS AND DISCUSSION........................................................................................................................... 9
5.1. Sediment composition......................................................................................................................... 9
5.2. Distribution of the macrobenthic fauna in 2001............................................................................... 10
5.2.1. Diversity, density and biomass .........................................................................................10
5.2.2. Temporal variation in density and biomass of individual species................................ 11
7. ACKNOWLEDGEMENTS................................................................................................................................. 15
8. REFERENCES...................................................................................................................................................... 16
Tables and Figures................................................................................................................................................... 18
Appendices............................................................................................................................................................... 46
THE MACROBENTHIC FAUNA IN THE DUTCH SECTOR OF THE NORTH
SEA IN 2001 AND A COMPARISON WITH PREVIOUS DATA
R. DAAN AND M. MULDER
This report presents data of the monitoring program o f macrozoobenthos in the Dutch
Continental Shelf (DCS) of the North Sea, a cooperation between the National Institute for
Coastal and Marine Management/RIKZ (Rijkswaterstaat), the North Sea Directorate
(Rijkswaterstaat) and the Department of Marine Ecology (NIOZ)
NETHERLANDS INSTITUTE FOR SEA RESEARCH
Monitoring Macrozoobenthos of the North Sea
NIOZ-RAPPORT 2002-1
Monitoring macrozoobenthos North Sea 2001
1. S u m m a r y
In this report the results are presented of a macrobenthos survey on the Dutch Continental
Shelf (DCS), carried out in spring 2001. The survey forms part of the 'Biological
monitoring programme of marine waters' (MON*BIOLOGIE, generally referred to as
‘BIOMON*) which was initiated by the National Institute for Coastal and Marine
Management (RIKZ). The purpose of the programme is to obtain insight into the year-toyear variations of the macrobenthic assemblages and to detect trend-like changes, that
possibly indicate anthropogenic influences on the marine environment {e.g.
eutrophication, pollution, beam-trawl fishery).
Within the framework of this project fieldwork is carried out every year in spring.
In 2001 the 100 BIOMON stations were sampled in the period between February 27 and
April 4. On the basis o f the results collected in 2001 and previous years an analysis is
made of the trends and fluctuations of some selected species and o f basic community
attributes over the period 1986-2001. The community attributes studied were the
diversity, abundance and biomass of the total macrofauna and of the 4 major taxonomic
groups. Temporal variation or trends were investigated separately for each of the four
subareas in the DCS i.e. the Coastal, Offshore areas, Dogger Bank and Oyster Ground.
The conclusions of this study can be summarized as follows:
1. The sediment composition in the four subareas was quite similar to that found in
previous years. Also at most of the individual stations the median grain size and silt
content of the sediment had hardly changed. There were only three stations with
strong year to year differences in sediment composition. At the stations OFF 2 and
OFF 6 the median grain size strongly fluctuated and at station OYS 8 the silt content.
It is suggested that these stations are situated at a sharp local gradient.
2. At the community level, there were only a few slight changes compared to preceding
years. In the offshore area and in the coastal area there seemed to be a tendency for
increased diversity. This was not due to increased species richness but to a more even
distribution in the abundance of the various species.
With respect to the share of the different taxonomic groups to the total biomass, a
decrease has been observed in previous years in molluscs, particularly at the Dogger
Bank and in the Oyster Ground. This decrease has come to an end in 2001 and turned
into an increase. Since molluscs also increased numerically, the increase in biomass is
caused by higher densities rather than by a larger size of the animals.
3. At the Dogger Bank a few species showed a remarkable decrease. The polychaete
Nephtys cirrosa shows a decrease from 2000 onwards, the sand star Acrocnida
brachiata and the bivalve Mysella bidentata from 1999 and the polychaete Aricidea
1
Monitoring macrozoobenthos North Sea 2001
minuta decreased in 1997 and has no longer been found since 1999. On the other hand
the population density of the gastropod Euspira nitida (formerly called Natica alderi)
had recovered from a dip in the period 1998 - 2000. The occurrence of Ensis
phaxoides is new for the Dogger Bank. Further, living specimens of the bivalve
Gouldia minima and the gastropod Turbonilla pusilla have not been found on the
DCS before.
4. In the Oyster Ground the decreasing trend that has been observed in previous years in
the brittle star Amphiura filiformis and the polychaete Nephtys hombergii had turned
into an increase in 2001. However, densities of A. filiform is were still low at the
Frisian Front. There was a number of species that have not been found before during
the BIOMON programme. The occurrence of the polychaete Nephtys assimilis is new
for the Oyster Ground. The records o f the polychaete Sabella penicillus, the bivalves
Montacuta tenella and Gari costulata and the gastropods Turbonilla pusilla and
Roxania utriculus are probably new to the DCS.
5. In the offshore area, a slightly increasing trend that has been observed in the sea
urchin Echinocardium cordatum in the preceding years has come to an end in 2001. In
contrast, the gastropod Euspira nitida showed a recovery of populations from 2000
onwards. Station OFF 33 that in 2000 had shown a rich fauna that was clearly
different from the other stations in the offshore area was still rich in 2001, but the
fauna composition did not show ‘exotic’ elements.
6. In the Coastal area there seemed to be a recovery of the populations of the gastropod
Euspira nitida. After an 8 year period o f very low densities the species returned at 5
stations, albeit in low numbers. The bivalve Tellina fabula also showed a steady
increase, after a dip in the second half of the nineties. At stations where banks of
Spisula subtruncata or Ensis americanus occurred, biomass values were very high.
2
Monitoring macrozoobenthos North Sea 2001
2. S a m e n v a t t in g
In dit rapport worden de resultaten gepresenteerd van een macrobenthos bemonstering die
in 2001 werd uitgevoerd op het Nederlands Continentale Plat (NCP). De bemonstering
vond plaats in het kader van het ‘Biologische Monitoring Programma Zoute Wateren’
(MON*BIOLOGIE, gewoonlijk aangeduid ais ‘BIOMON’), dat geïnitieerd is door het
Rijksinstituut voor Kust en Zee. Met het project wordt beoogd inzicht te krijgen in de
jaarlijkse fluctuaties van de macrobenthos gemeenschappen en vast te stellen of er op de
langere termijn trendmatige veranderingen optreden. Dergelijke veranderingen zouden
onder meer kunnen plaats vinden ais gevolg van anthropogene activiteiten (bijv.
eutrofiëring, verontreiniging, boomkorvisserij).
In het kader van dit project wordt jaarlijks veldonderzoek uitgevoerd in het
voorjaar. In 2001 zijn de 100 BIOMON stations tussen 27 februari en 4 april bemonsterd.
Aan de hand van de gegevens die in 2001 en voorgaande jaren zijn verzameld is een
overzicht verkregen van de trends en fluctuaties bij een aantal geselecteerde soorten en
een aantal kenmerken van de benthische gemeenschap ais geheel over de periode 1986 2001. Deze set kenmerken bestaat uit de diversiteit, de dichtheid en biomassa van de
totale fauna en de 4 belangrijkste taxa. Temporele variatie en trends zijn voor vier
subgebieden van het NCP, de Kustzone, het Offshore gebied, de Doggersbank en de
Oestergronden, afzonderlijk onderzocht. De conclusies van deze studie kunnen ais volgt
worden samengevat:
1. De doorsnee sedimentsamenstelling in de vier subgebieden vertoonde grote gelijkenis
met die welke in voorgaande jaren werd aangetroffen. Ook op de afzonderliojke
stations werden meestal geen grote veranderingen in mediane korrelgrootte of
slibgehalte gevonden. Er waren slechts drie stations waar sterke jaar-op-jaar
fluctuaties werden gevonden in de sedimentsamenstelling. Op de stations OFF 2 en
OFF 6 varieerde de mediane korrelgrootte sterk en op station OYS 8 het slibgehalte.
Verondersteld wordt dat deze stations gesitueerd zijn in gebiedjes waar scherpe locale
gradiënten voorkomen.
2. Op community niveau waren er slechts enkele kleine veranderingen ten opzichte van
de voorgaande jaren. In het offshore gebied en in de kustzone leek er sprake van een
toegenomen diversiteit, niet in de zin van een toegenomen aantal soorten, maar meer
wat betreft een meer evenwichtige verdeling in de dichtheden van de verschillende
soorten. Met betrekking tot het aandeel van de verschillende taxonomische groepen in
de totale biomassa is in voorgaande jaren een afname waargenomen bij de mollusken,
met name op de Doggersbank en in de Oestergronden. Deze afname is in 2001
3
Monitoring macrozoobenthos North Sea 2001
3.
4.
5.
6.
omgebogen in een toename. Aangezien mollusken ook numeriek toenamen moet de
biomassa toename eerder verklaard worden door hogere dichtheden dan door een
gemiddeld grotere afmeting van de organismen.
Op de Doggersbank vertoonden enkele soorten een opmerkelijke afname. De
polychaet Nephtys cirrosa neemt af sinds 2000, de slangster Acrocnida brachiata en
de bivalve Mysella bidentata vanaf 1999 en bij de polychaet Aricidea minuta zette de
afname in 1997 in. Deze laatste soort is sedert 1999 niet meer gevonden. Aan de
andere kant bleek de gastropode Euspira nitida (vroeger Natica alderi genoemd) zich
te hebben hersteld van een dal in de periode 1998 - 2000. Het voorkomen van de
bivalve Ensis phaxoides is nieuw voor de Doggersbank. Verder werden levende
exemplaren van de bivalve Gouldia minima en de gastropode Turbonilla pusilla
aangetroffen. Beide werden nog niet eerder op het NCP gevonden.
In de Oestergronden is aan de afnemende trend die in de voorgaande jaren is
waargenomen bij de slangster Amphiura filiform is en de polychaet Nephtys hombergii
een eind gekomen. Bij beide soorten was in 2001 in het algemeen sprake van een
toename. Niettemin waren de dichtheden van A. filiform is nog steeds laag op het
Friese Front. Er waren meerdere soorten die nog niet eerder gevonden zijn in het
kader van het BIOMON-programma. Zo werd de polychaet Nephtys assimilis nog niet
eerder in de Oestergronden aangetroffen. Het voorkomen van de polychaet Sabella
penicillus, de bivalven Montacuta tenella en Gari costulata en de gastropoden
Turbonilla pusilla en Roxania utriculus is waarschijnlijk nieuw voor het NCP.
Aan de licht toenemende trend die in de voorgaande jaren in het offshore gebied is
waargenomen bij de hartegel Echinocardium cordatum is in 2001 een eind gekomen.
Populaties van de gastropode Euspira nitida vertoonden daarentegen een herstel vanaf
2000. Station OFF 33 dat in 2000 een rijke fauna had laten zien, die duidelijk afweek
van de overige stations in het offshore gebied, bleek opnieuw rijk, maar de fauna
bevatte geen ‘vreemde' elementen.
In de kustzone leek er sprake van een herstel van de populaties van de gastropode
Euspira nitida. Na 8 jaar van zeer lage dichtheden bleek de soort op 5 stations weer
voor te komen, zij het in nog geringe aantallen. De bivalve Tellina fabula vertoonde
ook een gestage toename, na een dieptepunt in de tweede helft van de jaren negentig.
Op stations waar banken voorkwamen van Spisula subtruncata of Ensis americanus
waren de biomassagetallen zeer hoog.
4
Monitoring macrozoobenthos North Sea 2001
3. I n t r o d u c t io n
In 1989 the B iological M ONitoring programme of marine waters (project MON*
BIOLOGIE) was started with the goal to study the temporal variation of the marine
ecosystems on the Dutch Continental Shelf (DCS) including the Wadden Sea and the
Delta area. It is an initiative of the National Institute for Coastal and Marine Management
(RIKZ) of Rijkswaterstaat in association with several Dutch institutes (Yland, 1995). The
biological monitoring programme comprises besides the macrobenthos also plankton,
fish, seagrass, hard substrate populations, seabirds and mammals.
This report presents the data collected during the macrobenthos survey carried out
in spring 2001. Further the results of the 2001 survey are compared with the BIOMON
data collected in previous years (1991-2000) and those obtained during the ICES North
Sea Benthos Survey (ICES-NSBS, 1986) and the MILZON-BENTHOS programme
(1988-1993). In 1990 a pilot study of the BIOMON project was carried out at 7 locations
on the DCS and the results are also included in the data base.
The aim of the BIOMON programme is to obtain insight in the spatial and temporal
variation in the composition of the macrobenthos and to detect possible trendlike changes
on the DCS as a whole or in parts of it. During the first years (1991-1994) there were 25
stations located along 5 transects perpendicular to the Dutch coast. At these stations 5
replicate boxcore samples were collected each year. Athough in this way a rather detailed
picture was obtained of the fauna composition at each of these stations, it was argued that
(changes in) the macrobenthos composition of the DCS as a whole could better be studied
by spreading the sampling effort over a larger number of stations. Therefore, from 1995
onwards the sampling strategy changed and each year 100 stations were visited, that were
selected according to a stratified random sampling design in each of the 4 subareas of the
DCS, i.e. Dogger Bank, Oyster Ground, Offshore area and Coastal area (Fig. 1). The
number of stations within each subarea was proportional to its surface area. At each
station only one sample was taken. The 100 stations that were selected include the 25
original BIOMON stations. The selection procedure is described in more detail by Essink
(1995) and Holtmann et al. (1996)
The analysis of the results obtained in previous years (Daan & Mulder, 2001) has
shown that there were generally no clear trends at the community level (faunal density,
biomass, biodiversity parameters) in the 4 subareas. However, in most subareas there
seemed to be a slight decrease in the contribution of molluscs to the total benthic
biomass. At the species level there was a clear downward trend in the abundance of the
brittle star Amphiura filiform is in the Oyster Ground from 1993 onwards. Particularly at
5
Monitoring macrozoobenthos North Sea 2001
the Frisian Front a dramatic decrease was observed in the abundance of this species.
Further there was a decrease in the abundance of the polychaete Nephtys cirrosa and the
gastropod Natica alderi in the Offshore area and the Coastal area. The latter species also
decreased at the Dogger Bank. The new data will show to what extent the apparent trends
observed in previous years continued in 2001.
4 . M a t e r ia l
and m ethods
To ensure that any changes that are observed are not due to methodological differences,
the procedures for sampling and processing the fauna samples are standardized (Essink,
1991) and have remained unaltered since the beginning o f the monitoring project in 1991.
4.1. S a m p l i n g
In 2001 the BIOMON stations were sampled in the period February 27 to April 4. Most
stations have a water depth >5 m and were visited with the RV Mitra or the RV Arca
(North Sea Directorate, RWS). However, two stations in the Coastal subarea with a water
depth less than 10 m, viz. COA 13 & 14 were sampled with the RV. Delta (RWS).
Fig. 1 shows the positions of the stations . The exact geographical positions o f the
100 stations, together with the delta codes and selected abiotic characteristics
(depth/sediment) of the stations are summarized in Table la/b. More general information
about the cruise carried out with the RV. Mitra and the weather conditions during this
part of the survey in 2001 can be found in the cruise report of Rijkswaterstaat
(Anonymous, 2002).
4 .2 . S a m p l e
treatm ents
At each station two boxcore samples (0.068 m2, minimal depth 15 cm) were taken. One of
the samples was used for sediment analysis and the other sample was washed through a
sieve with round holes ( 1 mm) to collect the macrobenthic fauna. For sediment analysis 2
pooled subsamples (3.4 cm 0 , depth 10 cm) were immediately stored at -20°C. The
residue of the macrobenthos samples was preserved in a borax-buffered solution o f 4-6 %
formaldehyde in seawater and stored at room temperature.
In the laboratory the macrobenthos samples were stained with rose-bengal and
washed over a set of nested sieves with 0.7 mm as the smallest mesh size to facilitate
6
Monitoring macrozoobenthos North Sea 2001
sorting. The macrofauna was identified to species level, except for some notoriously
difficult taxa such as anthozoans, phoronids, priapulids and nemerteans, and subsequently
counted. Juvenile macrobenthic animals which because of their size could not be
identified to species level were recorded on higher taxonomic levels, usually the genus
level. Sizes (nearest 0.5 mm) were recorded for most molluscs and echinoderms.
4 3 . ASHFREE DRY WEIGHT
The ash-free dry weight (AFDW) of the different taxa was determined in one of the
following ways:
• Molluscs and echinoids:
By means of length-AFDW relationships of the form W =a*Lb (W=AFDW in g and
L=length in mm).
• Polychaetes. other worms. larger crustaceans and ovhiuroids:
Indirectly, by converting the (blotted) wet weight into AFDW by means of conversion
factors provided by Rumohr et al. (1987) and Ricciardi & Bourget (1998). W et
weights were measured with a M ettler PJ300 balance to the nearest mg.
• Remaining taxa:
Directly, by drying a sample at 60 °C for at least 60 hours and subsequently
incinerating at 520 °C for 2 hours (Duineveld & W itte, 1987).
Small molluscs, amphipods and cumaceans were assigned an average individual AFDW
of 0.2-0.5 mg. The same value is used by Holtmann & Groenewold (1992; 1994) in their
analysis of macrobenthos from the MTT ZO N -B F N T H O S project in the southern North
Sea between 1991 and 1993. This estimated individual weight is based on previous
determinations of the AFDW of the taxa in question (Duineveld; Holtmann, unpubl.).
4 .4. S t a t is t ic s
In addition to the density (ind./m2) and biomass (g AFDW /m2), the diversity of each
macrobenthos sample was calculated. In the literature a suit of biodiversity indices have
been used to identify possible changes of the benthic fauna (Hill, 1973; Peterson, 1977;
Pearson & Rosenberg, 1978; Harper & Hawksworth, 1994). In this report, we used three
indices each representing a different aspect of the faunal diversity. The species richness
7
Monitoring macrozoobenthos North Sea 2001
(Hillo) stands for the number of species per boxcore sample and is the simplest index. The
other two indices, the Shannon-Wiener index (H’) (Shannon & Weaver, 1949) and the
Simpson index (D) for dominance (Simpson, 1949), are based on the proportional
abundances of the individual species in the samples. The Simpson index is sensitive to
the abundance only of the more plentiful species and can therefore be regarded as a
measure of dominance (Hill, 1973). A high value for Simpsons index means low
diversity, whereas a high value for the Hillo or Shannon-Wiener index indicates high
diversity.
4 .5 . S e d i m e n t
a n a l y s is
At each station shown in Fig. 1, two subsamples were taken from an intact boxcore
sample and subsequently pooled for laboratory analysis o f the sediment composition (e.g.
grain size, content of calcium carbonate). The grain size was analysed with a Malvern
Particle Sizer by the laboratory of the National Institute for Coastal and Marine
Management (RUCZ, Middelburg). Two parameters were derived from the grain size data:
the median grain size (pm) and the percentage (by weight) of mud. We here define mud
as the total fraction mineral particles < 63 pm. However, for comparison with previous
years we also calculated the fraction 16-63 pm.
Sediment types were classified on the basis o f the median grain size as follows:
Characterisation of the sedimenttype according to
the median grain size (after Gullentops et al., 1977).
< 175 pm
Very fine sand
175 - 250 pm
Fine sand
250 - 300 pm
Medium-fine sand
300 - 350 pm
Medium-coarse sand
> 350 pm
Coarse sand
8
Monitoring macrozoobenthos North Sea 2001
5. Re s u l t s
a n d d is c u s s io n
5 .1 . S e d im e n t
c o m p o s it io n
The median grain size and silt content of the sediment at the stations sampled are listed in
Table 1. Spatial and temporal patterns are illustrated in Fig. 2, 3 and 4.
The spatial pattern in the sediment composition in 2001 was quite similar to that
in the preceding years. A look at the mean median grain size in the 4 subareas between
1995 and 2001 (Fig. 4) shows that the grain size is not only very stable, but that the
variation around the mean is extremely small. Also at most of the individual stations the
median grain size did not substantially change compared to preceding years. A
comparison between the values measured in 2001 (Fig. 2) and those found in previous
years shows that in more than 80% the size class did not change and that in most of the
other cases the difference was not more than 1 size class. There are, however, two
exceptions. At station OFF 2, north of Schiermonnikoog, the median grain size was about
340 pm in 1997 and 1999, but substantially lower, about 210 pm, in the other years. In
our previous report we already suggested that OFF 2 is situated at a sharp local gradient
where the sediment changes from fine sand into medium-coarse sand within a short
distance. As a consequence, small year to year differences in sampling position could
result in rather strong differences in sediment composition. At station OFF 6, west of
Texel, a different situation was met. Here the median grain size was 375 pm in 1995 and
dropped to values of about 310 pm in the period 1996 - 2000. In 2001 an even lower
value of 200 pm was found. It is not clear how this should be explained. Maybe the
situation is similar to that at OFF 2, i.e. a sharp local gradient. However, the possibility
that there has been a change in sediment composition over time should also be
considered. This would imply that there has been either a deposition of fine material at
OFF 6 or a disappearance of the larger size fractions. If a deposition of fine material had
occurred one might expect that silt concentrations had increased compared to previous
years. However the silt fraction was at the same low level (0.5%) as in previous years. On
the other hand a selective disappearance of the larger size fractions can only be explained
when there has been a local physical disturbance, probably by human activities. It is not
known whether such activities have taken place at OFF6.
The distribution of silt in the sediment also showed the same spatial pattern as in
the preceding years (Fig. 3). There was only one station where a substantial difference
was found in the silt content compared to 1999 and 2000. This was station OYS 8, at the
southern edge of the Frisian Front. In the preceding years the silt concentrations at OYS 8
were between 22 and 28 % but in 2002 it was less than 9 %. In this case we know that the
9
Monitoring macrozoobenthos North Sea 2001
station is located in an area with a sharp north-south gradient in silt concentrations which
is linked up with a relatively steep depth gradient. This means that a small difference in
positioning possibly could have a strong effect on the silt content measured. Another
explanation could be that the local depth is critical with respect to sedimentation and
resuspension of silt. This could be a cause of strong temporal variability of silt contents.
5.2. D is t r ib u t io n
o f t h e m a c r o b e n t h ic f a u n a in
2001
5.2.1 Diversity, density and biomass
A total number of 208 species/taxa were identified in the 100 boxcore samples in 2001,
including 16 that were identified to genus level only (most juveniles) and 12 higher taxa
(identified to family level or higher). The total number o f taxa is within the range of
previous years (181 - 231). The distribution of the species over the stations
(presence/absence) and the scientific names are given in A ppendix-1. The basic data on
macrobenthic abundance, biomass and diversity are listed in Appendix-2.
The mean number of species per sample (Hill 0) was, like in previous years, the
highest on the Dogger Bank and the lowest in the coastal and offshore area (Table 2, Fig.
5,8). There is an overall patterm of high species richness in the North (below the 30 m
depth contour) and low species richness in the south. In neither of the subareas a clear
long term trend could be observed in species richness. In the offshore area there were four
stations with a relatively rich fauna. The samples collected at OFF 1, 4, 5 and 33 all
contained more than 30 species. OFF 33 had been identified already in 2000 as a
remarkably rich station in an area, that is poor in fauna. The other three stations are
situated just south of the 30 m isobath, so close to the rich Oyster Ground. Therefore, the
relatively high number of species at these stations is not surprising.
The highest Shannon W iener diversity was also observed at the Dogger Bank,
whereas Simpson’s dominance was the lowest in this area (Table 2, Fig. 9,10). Numbers
o f individuals are more or less equally distributed over the different species and there are
no species which strongly dominate the fauna community by number. The lowest
Shannon Wiener diversity was found in the offshore and coastal area. However, there
seems to be a slight tendency for increasing diversity in these areas since the middle of
the nineties. On the other hand Sim pson's dominance seems to decline in this period. This
might indicate that dominance by one or a few species is less pronounced than it has been
and that the fauna is more equally distributed over the different species. Indeed, it seems
that some polychaete species (viz. Lanice conchilega, Magelona mirabilis and
10
Monitoring macrozoobenthos North Sea 2001
Spiophanes bombyx) which often have dominated the fauna in the two areas, were not so
abundant in 2001.
The mean fauna density was as usual the lowest in the offshore area (Table 2, Fig.
6). The fauna in this area was dominated by crustaceans and polychaetes O f the other
three areas the density was the highest in the Oyster Ground, but there were no large
differences. The Oyster Ground had particularly high numbers of echinoderms and
molluscs. At the Dogger Bank Crustaceans and polychaetes were the dominant groups, in
the coastal area molluscs and polychaetes. There were no trendlike changes in fauna
abundance (Fig. 11).
Biomass values were generally hardly different from those in 2000 (Fig. 12).
There does not seem to be a trendlike change. The highest mean biomass, but also the
strongest variation was found again in the coastal area (Fig. 7). As in previous years the
peak values found at some stations in the coastal area were generally caused by dense
populations of the mollusc Spisula subtruncata or by Ensis americanus.
W ith respect to the share of the different taxonomic groups to the total biomass a
shift has been noticed in some areas in 1999. Particularly at the Dogger Bank and in the
Oyster Ground the contribution of molluscs had decreased. Since this decrease was
compensated for by other taxa, there was no overall change in biomass. Mollusc biomass
in 2000 was still at the low level of 1999. However in 2001 we can see an increase of
mollusc biomass, both in an absolute sense and as share in the total biomass. Since
molluscs increased also numerically in three of the areas, the increase in biomass is
caused mainly by higher densities, rather than by a larger size of the animals.
5 .2 .2 . T e m p o r a l v a r i a t i o n in d e n s i t y a n d b io m a s s o f i n d i v i d u a l s p e c i e s
Figs. 13-16 illustrate the temporal variation in density or biomass of a number of
individual species in the 4 subareas during the period 1986-2001.
D o sser Bank (Fig. 13a-c)
On the Dogger Bank there were a few species that showed a remarkable decrease in 2001.
The sand star Acrocnida brachiata, the bivalve Mysella bidentata and the polychaetes
Nephtys cirrosa and Arcidea minuta occurred in lower densities than ever in the nineties.
A. brachiata and M. bidentata show a decrease since 1999, but of these species there were
still several tens of individuals per m 2 on average in 2001. N. cirrosa occurred already in
11
Monitoring macrozoobenthos North Sea 2001
low abundance in 2000, but that was supposed to be only a temporal dip. However, in
2001 there was a further decrease and the species was found at only two stations. In A.
minuta the decrease had started already in 1997 and from 1999 onwards the species has
no longer been found. The gastropod Euspira nitida (formerly called Natica alderi)
showed an increase compared to previous years. The species was found in low numbers
by the end of the nineties, but in 2001 population densities have recovered to a level
similar to that before 1998. The densities of other species that in previous years have been
found to be relatively abundant were more or less stable in 2001.
The brittle star Amphiura chiajei that colonised the Dogger Bank area in 1999
(Daan & Mulder, 2001) was abundant again in 2001. On average it was found in densities
of 150 in d .m '.
The occurrence of Ensis phaxoides is new for the Dogger Bank. The species was
found at two stations. We could find only a very few mentions of living E. phaxoides on
the DCS. The species has been found in beam trawl hauls at two stations in the offshore
area in the seventies by van Noort et al. (1979, 1984). Further the species was found in
two van Veen grab samples in the south-western Oyster Ground (Creutzberg, unpubl.
data).
Further we found two species that probably are new to the Dutch malacofauna.
Living specimens of the bivalve Gouldia minima and the gastropod Turbonilla pusilla.
have, to our knowledge, not been found on the DCS before. We found living T. pusilla
also at one station in the Oyster Ground and at one station in the offshore area. In
previous years we sometimes found empty shells of the species.
The occurrence of some new species on the Dogger Bank did not lead to increased
diversity in this area. Both, the number of species per sample and the Shannon-Wiener
index were in the same order as found in previous years. Apparently, the number of
samples in which the species were found was too low to have a substantial effect on the
diversity. However, the total number of species (87) on the Dogger Bank in 2001 was
higher than found in any of the previous years (66 to 80) and the new species have
certainly contributed to this overall high species richness.
Oyster Ground (Fig. 14a-c)
In previous years a declining trend has been observed in the brittle star Amphiura
filiformis and the polychaete Nephtys hombergii. In 2001 this decrease seems to have
come to an end. Both species showed an overall increase in their abundance.
Nevertheless, the densities of A. filiformis at the Frisian Front were still low compared to
12
Monitoring macrozoobenthos North Sea 2001
the numbers that were found here in the early nineties. Further, relatively high numbers
were found of the bivalves Corbula gibba and Nucula nitidosa (formerly called N.
turgida). The other more or less abundant species showed stable densities.
There was a number of species that have not been found before during the
BIOMON programme, or that are even completely new to the Dutch fauna. The
polychaete Nephtys assimilis that was found at OYS 36 has previously been found only in
the ‘Loswal' area west of Scheveningen (Aquasense, 1996,1997; Kluijver & van
Nieuwenhuizen, 1998). For another polychaete, Sabella penicillus, we could not find any
reference for its occurrence on the DCS. The other new species are all molluscs. The
bivalve Montacuta tenella was found at four stations in the Oyster Ground. This species
is known to live as a commensal of the echinoid Brissopsis lyrifera (Ockelman, 1965).
Indeed, B. lyrifera was found in two of the four samples too. We have inspected some
older data and found that B. lyrifera was found in 1997 in combination with Montacuta
spec. (Holtmann et al., 1998). A reidentification revealed that this was M. tenella. B.
lyrifera was found at four stations in 2001. This indicates that the species occurs in
increasing numbers in the Oyster Ground, which implies that M. tenella may also increase
in abundance in this area.
The gastropod Turbonilla pusilla was found at station OYS 38. The species also
occurred at the Dogger Bank and in the offshore area. In previous years only empty shells
of this species have been found. Completely new are the bivalve Gari costulata (found at
OYS 17) and the gastropod Roxania utriculus (found at OYS 3). Both have, to our
knowledge never been found on the DCS before.
In spite of the relatively high number of new species in the Oyster Ground, there
was no increased diversity in the area. The mean number of species per sample and the
Shannon W iener index were within the range of values found in previous years. Also the
total number of species (158) in the Oyster Ground was not particularly high compared to
previous years (135 to 178). This is explained by the fact that there was a number of
species that have been found now and then in previous years, but were not found in 2001.
The absence of the latter species apparently neutralized the presence of the new species,
so that the diversity did not change.
Offshore area (Fig. 15a-c)
The plots of the population densities o f 11 of the commonest species do not indicate that
there were substantial changes in 2001. From 1995 onwards, there has been a slightly
increasing trend in the sea urchin Echinocardium cordatum, but this increase has come to
an end. The opposite holds for the gastropod Euspira nitida (formerly called Natica
13
Monitoring macrozoobenthos North Sea 2001
alderi): a decrease during the second half of the nineties and a recovery of population
densities from 2000 onwards.
The Offshore area had, as usual, the poorest benthic fauna. Total fauna densities,
biomass and diversity were, like in previous years, the lowest in this area. In 2000,
however, there was one station with a remarkably rich fauna, that was clearly different
from all the other stations. This station (OFF 33), situated west of Umuiden, showed high
densities of the crustacean Callianassa subterranea and the brittle star Amphiura chiajei
and there were four mollusc species that were not or only very sparsely found alive
before. It was suggested that the sample could have been taken close to a ship's wreck. In
2001 the station appeared to be very rich again. However, in spite of the high species
richness (the sample contained 35 species), the species composition was not particularly
different from that at the other stations.
The gastropod Turbonilla pusilla was found at station OFF 11. It is remarkable
that this species of which in 2001 for the first time living specimens were found, did
occur at the Dogger Bank, the Oyster Ground and in the offshore area. Apparently, the
species does not seem to have a specific preference for a particular sediment texture.
Coastal area (Fig. 16a-c)
In the coastal area there seemed to be a recovery of the population densities o f the
gastropod Euspira nitida (= Natica alderi). This species had strongly decreased in 1992
and 1993 and occurred in very low numbers up to 2000. However, in 2001 the species
returned at 5 stations. There was also an increase in the bivalve Tellina fabula. After a dip
in 1996 and 1997 the species shows a steady increase up to 2001. The apparent increase
in the densities of the bivalve Mysella bidentata is caused by the extremely high density
at one station. At this station, COA 14, about 2000 individuals occurred per m".
The other more or less abundant species did not show substantial differences with
the preceding years. The densities of the polychaete Nephtys cirrosa were still at a low
level. After a steep decline in 1996 this species has never reached its former densities. N.
cirrosa is a species of sandy sediments and also occurs at the Dogger Bank and the
offshore area. Since the species showed low densities also at the Dogger Bank and, to a
lesser extent, in the offshore area, there seems to be a trend that covers the whole DCS.
Like in previous years, high biomass values were found at stations where banks of
bivalves occurred. Particularly high values were found at the stations COA 4, 9 and 12.
The banks at these stations were formed by Spisula subtruncata and/or Ensis americanus.
At stations where such banks were absent, the biomass values were generally low. For
example, an extremely low value of 0.2 g AFDW per m2 was found at station COA 13, in
front of the coast of Zeeland.
14
Monitoring macrozoobenthos North Sea 2001
6. Acknowledgements
The monitoring programme is initiated by the National Institute for Coastal and Marine
Management (RIKZ), with J. de Vlas and M. Latuhihin as project leaders, and is carried
out in cooperation with the North Sea Nirectorate (DNZ) and the department of Marine
Ecology of the NIOZ. We want to thank the captain and crew on board of the RV Mitra,
the RV Arca and RV Delta for their assistance during the fieldwork, W. Schreurs and G.
den Hartog (RIKZ Middelburg) for the analysis of the sediment samples, J. de Vlas for
critically reading the original manuscript, M. van Arkel for his contribution in the
organisation and H. Hobbelink for the cover design.
15
Monitoring macrozoobenthos North Sea 2001
7.
References
A n o n y m o u s , 2002. Meetverslag ms. Mitra, Arca, Rotterdam, jaargangen 2000, 2001 en
2002. Mon*Biologie Benthos. -Rijkswaterstaat, Directie Noordzee: 176 pp.
A q u a S e n s e , 1996. Effecten van de verplaatsing van Loswal Noord op het macrobenthos.
In opdracht van: Rijksinstituut voor Kust en Zee/RIKZ. Rapportnummer 96.0929.
A q u a S e n s e , 1997. Effecten van de verplaatsing van Loswal Noord op het macrobenthos de Tl-situatie. In opdracht van: Rijksinstituut voor Kust en Zee/RIKZ. Rapportnummer
97.1090.
D a a n , R. & M . M u l d e r , 2001. The macrobenthic fauna in the Dutch sector of the North
Sea in 2000 and a comparison with previous data. NIOZ-report 2001-2, NIOZ, Texel,
The Netherlands: 1-93.
D u in e v e l d , G.C.A., H.J. W itte , 1987. Report on an intercalibration exercise on methods
for determining ashfree dry weight of macrozoobenthos. - ICES CM 1987/L:39: 6pp.
E ssin k , K., 1995. Change of strategy for monitoring macrozoobenthos in the Dutch
sector of the North Sea. -National Institute for Coastal and Marine Management/RIKZ/
Working-document OS-95.606x: 5pp.
G u ll en to ps , F., M. M o e n s , A . R in g e le & R , S eng ier , 1977. Geologische kenmerken
van de suspensies en de sedimenten. -In: J. Nihoul & F. Gullentops (eds):
Mathematisch Model Noordzee. Vol 4. Sedimentologie.
H a r pe r , I.E. & D .L . H a w k s w o r t h , 1994. Biodiversity: measurement and estimation.
Phil. Trans. R. Soc., Ser. B, 345: 5-12.
H ill , M.O., 1973. Diversity and evenness: A unifying notation and its consequences.
Ecology 54(2): 427-432.
H o l t m a n n , S.E. & A . G r o e n e w o l d , 1992. Distribution of the zoobenthos on the Dutch
Continental Shelf: the Oyster Ground, Frisian Front, Vlieland Ground and Terschelling
Bank (1991). -NIOZ-report 1992-8, NIOZ, Texel, The Netherlands, NIOO-CEMO
rapporten en verslagen 1992-6: 129pp.
H o l t m a n n , S.E. & A . G r o e n e w o l d , 1994. Distribution of the zoobenthos on the Dutch
Continental Shelf: The western Frisian Front, Brown Bank and Broad Fourteens
(1992/1993). -NIOZ-report 1994-1, NIOZ, Texel, The Netherlands, NIOO-CEMO
rapporten en verslagen 1994-1: 136pp.
H o l t m a n n , S.E., J.J.M. B e l g e r s , B. K r a c h t & R. D a a n , 1996. The macrobenthic
fauna in the Dutch sector of the North Sea in 1995 and a comparison with previous
data. -NIOZ-report 1996-8, NIOZ, Texel, The Netherlands: 102 pp.
16
Monitoring macrozoobenthos North Sea 2001
H o l t m a n n , S.E., G.C.A. D u in e v e l d , M. M u l d e r & P.A.W.J.
d e W il de , 1998. The
macrobenthic fauna in the Dutch sector of the North Sea in 1997 and a comparison
with previous data. -NIOZ-report 1998-5, NIOZ, Texel, The Netherlands: 103 pp.
K l u u v e r , M.J. de & A. van Nieuwenhuizen, 1998. Effecten van de verplaatsing van
Loswal Noord op het macrobenthos - de T2 situatie van 1998. In opdracht van:
Rijksinstituut voor Kust en Zee / RIKZ. Aquasenserapport: 98.1280.
N o o r t , G.J. v a n , F. v a n L e e u w e n & F. C r e u t z b e r g , 1979. Aurelia cruise reports on
the benthic fauna of the southern North Sea; report 3 (1972). NIOZ-report 1979-6,
NIOZ, Texel, The Netherlands.
N o o r t , G.J. v a n , F. C r e u t z b e r g & R. D a p p e r , 1984. Aurelia cruise reports on the
benthic fauna of the southern North Sea; report 8 (1976). NIOZ-report 1984-1, NIOZ,
Texel, The Netherlands.
O c k e l m a n n , K.W., 1965. Redescription, distribution, biology and dimorphous sperm of
Montacuta tenella LovÉN (Mollusca, Leptonacea). Ophelia, 2: 211-221.
P e a r s o n , T.H. & R. R o s e n b e r g , 1978. Macrobenthic succession in relation to organic
enrichment and pollution of the marine environment. -Oceanogr. Mar. Biol. Ann. Rev.
16: 229-311.
Pe t e r s o n , C.H., 1977. Species diversity and perturbations: predictions of a non­
interactive model. -Oikos 29: 239-244.
R ic c ia r d i , A. & E. B o u r g e t , 1998. Weight-to-weight conversion factors for marine
benthic macroinvertebrates. -Mar. Ecol. Prog. Ser. 163: 245-251.
R u m o h r , H., T. B r e y & S. A n k a r , 1987. A compilation of biometric conversion factors
for benthic invertebrates in the Baltic Sea. -Baltic Marine Biology Publ. 9: 1-56.
S h a n n o n , C.E. & W. W e a v e r , 1949. The mathematical theory of communication. Univ. of Dlinois Press, Urbana.
S im p s o n , E.H., 1949. Measurements of diversity. -Nature, 163: 688-688.
Y l a n d , E., 1995. Biologisch monitoringprogramma zoute wateren, stand van zaken
1995. -Werkdocument RIKZ IT-95.170X: 39pp.
17
Monitoring macrozoobenthos North Sea 2001
Tables and Figures
18
ÍHETM)
(men* 'o
/(fe)
Stations Biomonitoring
North Sea
'* .3 6
(war.
< Dogger B ank {OOG)
♦ O yster G ro u n d '^ Îy ï^
C Offshore (O FF)
• Coasf {C O A )
\
Fig. 1. Locations of the sampling stations
* BIOMON 1991-1994
snn Dutch C ontinental S helf
i
r
median grain size (i^m)
<175
1 7 5 -2 5 0
•
250 - 300
300 - 350
#
>350
Fig. 2: Median grain size (|im) of the sediment in 2001
20
55 -
54 -
53 -
52 silt content (%)
•
#
2
3
4
5
<2
> 15
6
7
Fig. 3: Silt content (fraction <63 |iim) of the sediment in 2001.
21
3 40 3 20 300 2 80 -
median grain size
2 60 2 40 -
220
-
200
-
-A-
180 160
140 -
120
-
100
—
1995
1996
1997
1998
\~
1999
I
2000
2001
year
Fig. 4: T e m p oral trends in the m ean m edian grain size in the fo u r subareas.
22
diversity (Hill-0)
•
< 16
•
16-20
•
21-25
•
26-30
•
>30
Fig. 5: The number of species per sample (Hill-0) in 2001
d ensity (ind./m2)
•
<1000
1000-20 00
•
•
2000 - 3000
•
3000 - 4000
•
> 4000
Fig. 6: The total fauna density in 2001.
24
55 -
54
• •
53 -
••
••
52 biom ass (g A FD W /m 2)
•
2
3
4
5
•
< 10
10-20
•
#
4 0 -8 0
> 80
6
7
Fig. 7: The total biomass (g AFDW/m2) of the macrobenthos in 2001.
25
60
-,
Dogger Bank
504030-
2010-
nr. of species per sample
1985
1990
1995
60-,
2000
Oyster Ground
504030-
20101985
1990
1995
60-,
2000
Offshore area
504030-
20101985
1990
1995
2000
60-,
Coastal area
504030-
20101985
1990
1995
2000
year
Fig. 8: Temporal patterns in species richness (Hill-0) between 1986 and 2001
26
Dogger Bank
1985
1990
1995
2000
Oyster Ground
1985
1990
1995
2000
Offshore area
1985
1990
1995
2000
Coastal area
1985
1990
1995
2000
year
Fig. 9: Temporal patterns in Shannon-Wiener diversity between 1986 and 2001.
Dogger Bank
1.00.80.60.4-
0.20.0
1985
1990
1995
2000
Oyster Ground
1.0-
0.60.4-
0.2-
Simpson index
0.0
1985
1990
1995
2000
Offshore area
1.00.8-
0.60.4-
0.20.0
1985
1990
1.0-
1995
2000
Coastal area
0.60.4-
0.20.0
1985
1990
1995
2000
year
Fig. 10: Temporal patterns Simpson's dominance between 1986 and 2001.
10 0 0 0 n
Dogger Bank
8000600040002000-
1985
1990
1995
2000
Oyster Ground
10000-
8000-
(gLu-pu!) Áüsuap
600040002000-
1985
1990
1995
30000-,
2000
Offshore area
20000-
10 0 0 0 -
1985
1990
1995
2000
40000-,
Coastal area
30000-
2 00 0 0 -
100 00-
1985
1990
1995
2000
year
Fig. 11: Temporal patterns in macrobenthos density between 1986 and 2001.
29
□
40-1
Dogger Bank
30-
2010-
biomass (g AFDW per m )
1985
1990
1995
75-,
2000
Oyster Ground
60453015-
1985
200
1990
1995
-,
2000
Offshore area
160120 -
8040-
1985
1990
2000
1995
500-,
Coastal area
400300200100 -
1985
1990
2000
1995
year
Fig. 12: Temporal patterns in biomass between 1986 and 2001.
400
-,
Acrocnida brachiata
300-
2001001985
1990
1995
2000
1990
1995
2000
1990
1995
2000
1600-,
Bathyporeia elegans
1200 -
800-
400-
1985
7501
Mysella bidentata
600-
ind. per m'
450300150-
1985
120-,
100-
Euspira nitida (=Natica alderi)
806040-
201986
1988
1990
1992
1994
1996
1998
2000
year
Fig. 13a: Densities of 4 species at the Dogger Bank (1986-2001)
31
2002
1000
-,
Tellina fabula
800600400-
2001990
1985
1995
2000
1990
1995
2000
1990
1995
ind. per m
750-,
Magelona mirabilis (=M. papillicornis)
600450300150-
1985
250-,
200-
Nephtys cirrosa
150-
10050-
1985
2000
year
Fig. 13b: D ensities of 3 sp e c ie s at th e D ogger Bank (1986-2001)
32
120
-,
100 -
Aricidea minuta
6040-
201985
1990
1995
2000
1995
2000
1995
2000
250-,
Chaetozone setosa
200150-
10050-
number, per m‘
1985
1990
2500-,
Spiophanes bombyx
20 0 0 -
15001000 -
500-
1985
1990
750-,
Urothoe poseidonis
600450300150-
1986
1988
1990
1992
1994
1996
1998
2000
year
Fig. 13c: Densities of 4 species at the Dogger Bank (1986-2001)
33
2002
4000-1
Amphiura filiformis
320024001600800-
1985
1990
1995
2000
1995
2000
1995
2000
1995
2000
200Echinocardium cordatum
O
160
Q.
lí 120
80-I
40
1985
1990
50-,
Echinocardium cordatum
40CM
E
<
u 30a.
§Q 20LL
1985
1990
300-,
Callianassa subterranea
240-
E
O)
Q.
■cO
180-
12060-
1985
1990
year
Fig. 14a: Densities (and biomass for E. cordatum) of 3 species in the Oyster Ground
(1986-2001).
34
300
N atica
alderi
240180-
12060-
ind. per m'
1985
1990
1995
2000
1990
1995
2000
1995
2000
250-,
200-
Nucula turgida
150-
10050-
1985
200-,
Nephtys hombergii
160-
1208040-
1985
1990
year
Fig. 14b: Densities of 3 species in the Oyster Ground (1986-2001)
35
300-
Chaetozone setosa
250-
200150100 -
50-
01985
200
1990
1995
2000
-,
Chaetopterus variopedatus
150-
100
50-
1985
1000
,
—»—
1990
1995
I
J
«
—1—
2000
Corbula gibba
100,
10,
1985
1990
1995
1990
1995
2000
300-,
number per m
Harpinia antennaria
240180-
12060-
1985
2000
year
Fig. 14c: Densities of 4 species in the Oyster Ground (1986-2001).
36
Echinocardium cordatum
density
300
250
200
150
100
1985
1990
1995
2000
1995
2000
1995
2000
75-,
g a f DW per m'
60-
Echinocardium cordatum
biomass
45-
1985
2000
1990
-,
Bathyporeia elegans
ind. per m
16001200 -
800400-
1985
1990
150-,
120-
Euspira nitida (=Natica alderi)
E
<5
Q.
■co
6030-
1985
1990
1995
2000
year
Fig. 15a: Densities (and biomass of E. cordatum) of 3 species in the offshore area
(1986-2001).
37
Tellina fabula
600-
400-
200-
1985
1990
1995
2000
1990
1995
2000
1990
1995
2000
1990
1995
8000-,
Magelona mirabilis (=M. papillicornis)
6000-
4000-
2000-
1985
400-,
Nephtys cirrosa
300-
200-
100-
1985
800-,
Scoloplos armiger
600c\i
E
0-Q.)
400-
200 -
1985
2000
year
Fig. 15b: Densities of 4 species in the offshore area (1986-2001)
38
250
Aricidea minuta
200
150
100 -
50-
1985
1000
Chaetozone setosa
100 ,
number per m'
10
1985
Lanice conchilega
1985
10000,
1990
1995
2000
1995
2000
Spiophanes bombyx
1000 ,
100
10,
■
1985
1990
year
Fig. 15c: Densities of 4 species in the offshore area (1986-2001).
39
150-1
Echinocardium cordatum
density
120-
906030-
1985
1990
1995
2000
1995
2000
1990
1995
2000
1990
1995
2000
60-,
50-
Echinocardium cordatum
biomass
40302010-
1985
1990
2500-,
Urothoe poseidonis
2000 -
g AFDW per m'
15001000-
500-
1985
2000
E
(Q.1)
-,
Mysella bidentata
1500-
10 0 0 -
500-
1985
year
Fig. 16a: Densities (and biomass of E. cordatum) of 3 species in the coastal area
(1986-2001).
40
100
-,
Euspira nitida (=Natica alderi)
80604020-
1985
1990
1995
2000
1990
1995
2000
1990
1995
2000
1990
1995
6000-,
Spisula subtruncata
50004000300020001000 -
1985
1200 -,
Tellina fabula
1000 -
ind. per m'
800600400200-
1985
500-,
Nephtys cirrosa
400300200100 -
1985
2000
year
Fig. 16b: Densities of 4 species in the coastal area (1986-2001)
41
1000
-,
Capitella capitata
800600400200-
¡
1985
100004
number per m
1985
i
' r M I
1990
1995
2000
1990
1995
2000
1995
2000
1995
2000
Lanice conchilega
Magelona mirabilis (=M. papillicornis)
1985
1990
Spiophanes bombyx
1985
1990
year
Fig. 16c: Densities of 3 species in the coastal area (1986-2001).
42
Table 1a. Station number, position, date, depth and sediment composition of the survey 2001.
Station (name)
NIOZ
DONAR
Code
Code
DOG 1 DOGGBK07
DOG
DOG
DOG
DOG
DOG
DOG
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
OYS
2 DOGGBK02
3 DOGGBK03
4 TERSLG235
5 DOGGBK04
6 DOGGBK05
7 DOGGBK08
1 OESTGDN43
2 FRIESFT16
3 OESTGDN02
4 OESTGDN03
5 FRIESFT02
6 OESTGDN04
7 OESTGDN05
8 FRIESFT03
9 FRIESFT04
10 OESTGDN06
11 FRIESFT05
12 OESTGDN07
13 OESTGDN08
14 OESTGDN09
15 OESTGDN10
16 OESTGDN11
17 OESTGDN12
18 FRIESFT06
19 OESTGDN13
20 OESTGDN14
21 TERSLG50
22 OESTGDN15
23 OESTGDN16
24 BREEVTN34
25 OESTGDN17
26 FRIESFT07
27 OESTGDN18
28 FRIESFT08
29 OESTGDN19
30 BREEVTN02
31 FRIESFT09
32 FRIESFT10
33 OESTGDN20
34 FRIESFT11
35 FRIESFT12
36 FRIESFT17
37 TERSLG100
38 BREEVTN26
39 OESTGDN22
40 OESTGDN21
41 OESTGDN23
42 ROTTMPT70
Geographical position
E
N
04°03'00'
03°38'30'
03°30'00'
03°09'26"
03°14'00'
03°05'0003°00'00'
03°25'30'
05°32‘30'
04°00'00'
02°56'00‘
04°55'00'
04°22’48"
04°18'00"
04°54'00‘
03°37‘50’
03°42'30‘
05°10'00"
04°26'00'
03°30'00'
04°44'30"
04°21 '20'
05°03'00"
03°25'08'
05°54'00'
03°19'00'
02°51'51"
04°46'03'
03°38'30‘
03°22'00'
03°29'46"
04°32'00'
04°4730‘
05°00’00'
03°30'00"
03°00'00'
o s - is ^ r
04°09'06'
05°05'00"
04°03'00"
04°16'37*
03°52'24"
04°30'00"
04°20'27*
03°00'00'
04°00'00'
05°00'00'
03°1736"
06°12'5T
55°28'18"
55°10'00"
55°15'00"
55°10'14'
54°54'42"
54°57'06"
SS'O O W
54°23'0054°11'30"
55°00'00"
54°33'00'
54°01'10‘
55°18'24”
54°53'00‘
53°44'40’
53°45'20'
54°39'00"
53°55'30'
54°10'OO"
54°45'00"
54°20'00’
54°28'30’
54°38’30"
54°00‘2 r
54°11'20"
54°20'00'
54°05'00"
53°46'04'
54°18'30"
54°49'24"
ÔS^'OO54°39'00‘
53o55'2054°30'0053°45'00'
54°30'00'
53°31 '30"
53°50'42"
54°15'30'
54'>16'00"
53°3740"
53°51 '31 “
53°42‘05'
54<>09'04"
53°30'00'
54°30'00"
öö^OO'OO'
54°51'42"
54°0r03’
Date
Depth (m)
14/03/2001
14/03/2001
14/03/2001
13/03/2001
14/03/2001
14/03/2001
14/03/2001
15/03/2001
06/03/2001
15/03/2001
15/03/2001
08/03/2001
14/03/2001
15/03/2001
08/03/2001
03/04/2001
13/03/2001
08/03/2001
13/03/2001
13/03/2001
06/03/2001
13/03/2001
06/03/2001
04/04/2001
06/03/2001
15/03/2001
04/04/2001
08/03/2001
15/03/2001
14/03/2001
03/04/2001
15/03/2001
08/03/2001
06/03/2001
03/04/2001
15/03/2001
03/04/2001
08/03/2001
06/03/2001
15/03/2001
08/03/2001
08/03/2001
08/03/2001
13/03/2001
03/04/2001
13/03/2001
06/03/2001
14/03/2001
06/03/2001
43
30.0
36.2
28.1
30.1
35.7
23.0
25.0
45.5
39.3
47.6
34.0
43.0
46.0
50.3
36.7
37.5
44.3
39.5
47.0
44.5
46.8
50.1
46.7
44.0
37.1
48.2
51.5
38.3
43.7
41.5
33.5
49.6
41.5
43.7
35.0
36.1
35.0
43.8
43.8
47.8
37.5
40.2
38.8
49.3
33.5
44.7
41.4
39.3
32.7
Sediment composition
Med.Gr.
Mud (%)
Mud (%)
Size (pm) Fr.<63 pm Fr.16-63 pm
218
186
205
201
173
217
209
112
210
114
138
105
151
87
202
182
114
83
92
114
128
88
156
194
220
121
166
113
159
133
80
107
134
172
200
125
122
143
158
105
111
160
103
95
139
112
153
149
228
1.3
1.6
0.4
0.5
1.3
1.2
1.1
10.9
9.4
8.2
1.9
20.9
4.1
19.4
8.7
2.8
6.7
38.2
19.7
5.8
16.4
26.4
7.3
3.0
2.1
6.4
18.0
23.8
5.8
4.7
33.7
16.7
18.6
3.7
1.9
3.3
17.3
8.6
12.6
13.9
22.6
3.7
29.9
17.0
6.6
11.6
3.8
2.4
1.1
0.9
0.0
0.0
0.0
0.0
0.9
0.8
5.3
3.6
2.7
0.0
10.1
0.0
12.0
2.5
0.0
1.8
27.0
10.6
1.8
7.6
16.2
0.9
0.0
0.0
1.8
9.2
11.0
0.9
0.9
26.7
10.7
6.9
0.0
0.0
0.9
9.8
2.6
4.3
6.9
12.8
0.0
15.1
9.2
1.7
6.3
0.0
0.0
0.0
Table 1a. Station number, position, date, depth and sediment composition of the survey 2001.
Station (name)
NIOZ
DONAR
Code
Code
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
OFF
COA
COA
COA
COA
COA
COA
COA
COA
COA
COA
COA
COA
COA
COA
COA
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
FRIESFT13
WADDKT07
WADDKT02
FRIESFT14
FRIESFT15
BREEVTN03
BREEVTN04
BREEVTN05
BREEVTN06
BREEVTN07
BREEVTN08
BREEVTN09
BREEVTN10
BREEVTN11
BREEVTN12
BREEVTN13
BREEVTN14
BREEVTN15
BREEVTN16
BREEVTN17
BREEVTN18
BREEVTN19
BREEVTN20
BREEVTN21
BREEVTN22
BREEVTN23
BREEVTN24
BREEVTN25
ROTTMPT50
TERSLG30
BREEVTN27
NOORDWK30
NOORDWK50
NOORDWK70
WALCRN30
WALCRN70
WADDKT03
WADDKT04
HOLLSKT03
HOLLSKT02
WADDKT05
WADDKT06
ROTTMPT3
TERSLG4
HOLLSKT04
NOORDWK2
NOORDWK10
VOORDTA2
VOOR DTA3
VOORDTA4
VOORDTA5
Geograph cai position
E
N
05°59‘00'
06°06'25"
05°49'37"
04°57'30"
04°22'30"
04°26'32"
04°18'22*
04°00'30"
04°13'50'
03°50'30°
03°31'1803°23'30°
03° 11'36°
03°17'20"
03°1718'
03°30'00‘
03°12'12"
03°11'25’
03°24'42'
03°30'00‘
03°00'00'
03°59'15'
04°09'5003°42'58"
03°24'26‘
03°11'34"
03°14'28‘
02°52'48"
06°18'36'
04°56'17’
03°55'0r
04°02'53‘
03°4707°
03°31 '53"
03°06'49"
02°40'45"
05°59'53"
05°374804°31'50"
04°40'00"
04°41 '20"
06°11'0306°32'46"
05°09'02"
04°30'00'
04°24'20‘
04°18'0103°23'15"
03°36'02'
03°48'48"
03°55'09'
53°51 '30'
53°3729"
53°36'40'
53°40'00"
53°29'00'
53°11'1653°05'5953°01'30"
52°49'20‘
52°45'40"
53°1700'
53°03'55‘
53°02'58‘
52°53'53'
52°50'12"
52°45'00"
52°27'43’
52°20'25'
52°15'10‘
52°15'00"
52°00'00'
52°16'30‘
52°23'08‘
52°00'00'
52°06'12’
51 °56'07"
51°41 ‘40°
51 °52'40‘
53°5714'
53°36'56'
52°59'53'
52°23'15'
52°28'30"
52°34'10"
51 °43'06"
51 “5725“
53°32'34’
53°30'19"
52°32'50‘
52°50'00'
53°03'23'
53°32'0953°34'57"
53°24’54"
52°45‘00‘
52°15'3652°1741‘
51 °3704"
51 °42'23"
51 °47'26"
51 °55'20‘
Depth (m)
Date
06/03/2001
27/02/2001
27/02/2001
08/03/2001
08/03/2001
09/03/2001
09/03/2001
05/03/2001
05/03/2001
28/02/2001
03/04/2001
16/03/2001
16/03/2001
16/03/2001
16/03/2001
28/02/2001
16/03/2001
03/04/2001
02/04/2001
02/04/2001
02/04/2001
28/02/2001
28/02/2001
02/03/2001
02/04/2001
02/04/2001
01/03/2001
01/03/2001
05/03/2001
08/03/2001
05/03/2001
28/02/2001
28/02/2001
28/02/2001
01/03/2001
02/04/2001
27/02/2001
27/02/2001
28/02/2001
27/02/2001
27/02/2001
27/02/2001
27/02/2001
27/02/2001
26/02/2001
28/02/2001
28/02/2001
01/03/2001
22/03/2001
22/03/2001
01/03/2001
44
31.1
23.3
26.2
31.4
28.6
30.8
35.7
29.2
26.4
30.0
26.8
28.0
29.4
32.8
33.3
26.5
26.8
29.1
28.8
31.4
37.0
23.3
22.5
24.0
31.1
29.0
26.7
34.0
29.7
25.0
26.2
23.3
30.0
31.0
28.4
44.0
18.1
9.1
18.0
11.3
11.4
9.4
7.2
12.0
21.4
13.0
18.5
11.5
5.1
3.6
14.5
Sediment composition
Med.Gr.
Mud (%)
Mud (%)
Size (pm ) Fr.<63 pm Fr.16-63 pm
211
215
188
200
216
200
249
143
198
291
152
242
268
272
301
268
300
337
363
388
462
376
344
429
435
516
358
549
381
214
254
354
289
308
357
493
229
182
228
188
212
169
183
234
228
265
342
281
276
280
201
1.7
1.2
1.8
2.7
2.1
0.5
1.1
4.4
0.3
0.8
5.8
2.6
0.6
1.0
0.6
0.4
2.3
0.3
0.3
0.4
1.3
0.8
0.6
0.6
0.2
0.2
0.6
0.3
0.3
0.5
0.3
0.5
0.8
0.5
0.6
0.3
0.8
0.9
1.3
4.4
1.0
2.8
0.7
0.6
1.6
1.1
0.6
0.8
0.1
0.6
1.3
0.0
0.0
0.0
0.0
0.0
0.0
0.0
3.6
0.0
0.0
4.6
1.9
0.0
0.0
0.0
0.0
1.9
0.0
0.0
0.0
0.9
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
Table 2. Mean values of abiotic and biotic parameters in the 4 areas in 2001.
AREA
Dogger Bank
No. of stations
Median Grain Size (pm)
Silt content (fr.< 63um,%)
silt (fr. 16- 63um, %)
Depth (m)
7
Oyster Ground
42
Offshore area
36
Coastal area
15
201
1.1
0.4
30
138
11.9
5.7
42
309
1.1
0.4
29
233
1.2
0.0
12
Diversity:
Total num ber of species
Number of specier per core
Shannon- W iener diversity
Simpson's dominance
87
35.7
2.76
0.12
158
29.8
2.35
0.20
103
17.0
2.23
0.14
51
16.9
1.92
0.24
No. individuals (ind./m2):
Crustaceans
Echinoderms
Molluscs
Polychaetes
Miscellaneous
887
251
288
596
108
209
607
797
460
167
326
45
130
430
52
398
21
769
642
29
2130
2240
983
1859
0.6
7.0
2.3
3.7
0.8
3.4
5.3
3.2
3.7
1.6
0.3
6.0
5.5
2.7
0.4
1.5
7.6
40.7
5.0
0.2
14.4
17.2
14.9
55.0
TOTAL D E N S IT Y
Biomass (g AFDW /m2):
Crustaceans
Echinoderms
Molluscs
Polychaetes
Miscellaneous
TOTAL B IO M A SS
m
E E E E E E s
S
«
S
4
1
2
1
2
3
5
6
O '/
s te r G ro u n d
in
in
in
in
m
<n
S
<A
S'
S' S ' S
S
S
S
S
4
5
6
7
8
9
10
11
12
13
14
+
+
+
+
+
+
+
+
k
7
vi
$
M
<0
S
3
+
in
I
Oys
O
Û
gger B an
Oys
Appendix-1 Biomonitoring 2001 (+=presence)
S
in
in
S
S
15
16
17
18
C ode
Species nam e
ABRA ALBA
+
+
ABRA PRISMATICA
ABRAALBA
A B R A PR IS
+
ACANECHI
ACANTHOCARDIA ECHINATA
A C ID O B E S
ACIDOSTOMA O BESUM
ACROCNIDA BRACHIATA
+
+
+
+
+
ACROBRAC
+
ALTENAEUM DAW SONI
A LTED A W S
+
AMPELISCA BREVICORNIS
+
+
+
+
+
+
AMPELISCA TEN U IC O R N IS
+
+
+
+
+
+
+
+
+
+
+
AMPHFINM
A M PH ILSP
+
AMPHILOCHUS S P E C .
AMPHIOXUS LANCEOLATUS
+
AMPHIURA CHIAJEI
+
AMPHLANC
+
+
+
AMPHCHIA
+
+
AMPHIURA FILIFORMIS
+
+
+
+
+
+
+
+
+
ANTHOZOA
+
+
+
+
+
+
+
+
AO N IPA U C
APHEMARI
+
+
APHELOCHAETA MARIONI
A PH ER B IS
APHERUSA B ISPIN O SA
APH EOVAL
APHERUSA O V A LIPES
+
APHERUSA S P E C .
+
APHRODITE ACULEATA
+
+
+
+
APH ESPEC
APH RA CU L
+
+
APLACOPHORA
A PLA C O PH
+
+
+
+
+
ASTERIAS R U B E N S
A ST R IR R E
+
A STRO PEC TEN IRREGULARIS
+
+
+
ATYLFALC
+
ATYLSW AM
+
ATYLUS SWAMMERDAMI
BATHYPOREIA ELEGANS
+
BATHYPOREIA GUILLIAMSONIANA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ BATHELEG
+ BATHGUIL
+
+
+
+
BIVAINDE
BIVALVE INDET.
+
B RISSO PSIS LYRIFERA
+
CALLIANASSA SU BTERR A N EA
BRISLYRI
+
+
CALLIANASSA S P E C . JUV.
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ CALLJUVE
CA LLSUBT
CALLTYRR
CALLIANASSA TYRRHENA
CAPICA PI
CAPITELLA CAPITATA
+
CAPRELLIDAE
+
CAPRELLI
+
+
+
CH A ETO PTER U S VARIOPEDATUS
CHAETOZONE S E T O S A
+
CHAMELEA STRIATULA
+
+
+
+
+
CERILLOY
+
+
CERIANTHUS LLOYDII
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
CHAEVARI
+
+
+
C H A M SP E C
CH O N D U N E
CHONE DUNERI
+
CORBULA GIBBA
+
+
CORY STES CA SSIVELAUNUS
CRANGON C RA N G O N
+ C H A E SE T O
CHAM STRI
+
CHAMELEA S P E C . JUV.
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ C O R B G IB B
+
C O R O IN SI
+
+
CO RY CA SS
+
CRA N CRAN
CUCUMARIA FR O N D O SA
C U C U FR O N
+
CULTELLUS PELLUCIDUS
+
CYLICHNA CYLINDRACEA
DIASTYLIS BRADYI
ARCTISLA
ARICMINU
A ST E R U B E
ARICIDEA MINUTA
COROPHIUM INSIDIOSUM
+ AMPHFILI
A N TH O ZO A
+
+
+
AONIDES PAUCIBRANCHIATA
ATYLUS FALCATUS
A M PEB R EV
A M PETEN U
+
AMPHARETE FINMARCHICA
ARCTICA ISLANDICA JUV.
in
+
+
+
+
+
+
+
+
+
+
+
+
+
+
46
+
+
+
+
+
+
+
+
+ CU LTPELL
+ CYLICYLI
+
+
DIASBRAD
Appendix-1 Biomonitoring 2001 (+=presence)
O
a
i
?
1 2
99
er Ban k
SS
3
4
I
5
Oyster Ground
Ss
6
7
w «
ô* S'
1 2
+
è
<0
U
)
S' S'
S
3
4
5
6
+
+
CO CO <0 co co CO co CO CO CO
CO
S S’ S' o S' S' s S' S'
7 8 9 10 11 12 13 14 15 16 17 18
S' S' S'
Species nam e
C ode
DIPLOCIRRUS GLAUCUS
+
+
+
+
+
DONAX VITTATUS
DONAVITT
DOSINIA EXOLETA
+
DOSINIA LUPINUS
+
+
+
+
+
+
+
ECHINOCYAMUS PU SILLUS
EDWARDSIA CLAPAREDII
+
+
EBALIA CRANCHII
ECHINOCARDIUM CORDATUM
+
+
+
+
+
DOSIEXOL
+
DOSILUPI
+
+
+
+
+
+
+
+
EBALCRAN
+
+
+
+
+
+
+
+
ECHICORD
+
ECHIPUSI
+
EDW ACLAP
EN SIS AMERICANUS
ENSIAMER
EN SIS ARCUATUS
ENSIARCU
EN SIS ENSIS
EN SIS PHAXOIDES
+
+
+
+
ENSIENSI
+
ENSIPHAX
ETEONE BARBATA
+ ETEO BA RB
ETEONE LONGA
+
EUDORELLA TRUNCATULA
+
EU D O RELLOPSIS D EFO R M IS
+
+
+
+
+
+
+
EUMIDA SANGUINEA
+
+
+
ETEO LON G
+ EUDOTRUN
+
EU D O D EFO
+
EUMISANG
EUSPIRA CATENA
EUSPIRA NITIDA
EU SPC A T E
+
+
+
+
+
+
+
+
+ EUSPNITI
EUZONUS FLABELLIGERUS
EXOGONE HEBES
EUZOFLAB
+
GOULDIA MINIMA
EX O G H EBE
+
GOULMINI
GARI COSTULATA
GARI FERVENSIS
+ DIPLGLAU
+
+
+
+
GATTYANA C IR RO SA
+
+
+
+
G A RICO ST
+
GARIFERV
+
G A TTCIRR
GLYCERA LAPIDUM
GLYCLAPI
GLYCERA ROUXI
GLYCROUX
GLYCERA S P E C . JUV.
G LY C SPE C
GLYCINDE NORDMANNI
+
GOLFINGIA ELONGATA
GOLFINGIA PR O C ER A
+
GOLFINGIA VULGARIS
+
GONIADA MACULATA
GLYCNORD
+
+
+
+
+
+
+
+
+
+
+
+
+
GOLFELO N
GO LFPRO C
+
+
+
+
+
+
+
+
+
+
+
+
GOLFVULG
+ GONIMACU
GONIADELLA BOBRETZKII
G Y PTIS CAPEN SIS
G O NIBOBR
+
+
+
+
+
G Y PTC A PE
+ HARMGLAB
HARMOTHOE GLABRA
HARMOTHOE IMBRICATA
HARMIMBR
HARMOTHOE LJUNGMANI
HARMLJUN
HARMOTHOE LUNULATA
HARMOTHOE S P E C . JUV.
+
+
+
HARPINIA ANTENNARIA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
HETEROM ASTUS FILIFORM IS
HETEFILI
HIATELLA ARCTICA
+
HIPPOMEDON DENTICULATUS
HYALA VITREA
HIATARCT
HIPPD EN T
+
+
HYPERIIDAE S P E C .
+
+
+
IPHIMEDIA OBESA
+
+
+
+
+
+
+
HYALVITR
HYPERIID
+
+
+
+
+
+
IONE THORACICA
IPHINOE TR ISPIN O SA
+
HARMLUNU
+ H A RM SPEC
+ HARPANTE
IONETHOR
IPHIOBES
+
KELLIA SUBORBICULARIS
+
+
47
+
IPHITRIS
+
KELLSUBO
Appendix-1 Biomonitoring 2001 (+=presence)
D o<g g e r B a n
g
G
E E
1I2
0 )/s te r G ro u n d
k
S
E
S
9
o
3
4
5
6
7
+
+
<0
S
v>
&
a'
a aa
1
2
3
4
<0
V)
V)
5
<0
6
IO
co
IO
co
a* §
&
&
a* a-
7
ê
IO
&
8
10
11
12
13
9
14
CO CO
a'
a
a
15
16
17
18
Code
Species nam e
LANICE CONCHILEGA
LANICONC
+
LANIJUVE
LANICE S P E C . JU V .
LEM BLONG
LEMBOS LO N G IPES
+
+
+
+
+
+
+ LEUCINCI
+
LEVIGRAC
+
LEVINSENIA G RACILIS
LIOCARCINUS S P E C . JUV .
LE PT SQ U A
+
LEPTON SQUAM OSUM
LEUCOTHOE INCISA
L IO C S P E C
+
LUMBRINERIS FRAGILIS
+
LUMBRINERIS LATREILLI
+
+
LUM BFRAG
+
LUMBLATR
+
+
LYSILOVE
+
LYSILLA LOVENI
MACOBALT
MACOMA BALTHICA
MACTRA CORALLINA
+
MAGELONA ALLENI
+
+
MAGELONA MIRABILIS
+
+
+
+
+
+
+
+
+
+
+
+
+
MAGEALLE
+
+
+
+
+
+
M ACTCORA
+
+
+
MEDIOMASTUS FRAGILIS
M EGALUROPUS AGILIS
+
+
+
+
+
+
+
+
+
M EGAAGIL
MELITOBTU
+
MICRMACU
M IC RO PR O TO PU S MACULATUS
M O D ISPE C
+
MODIOLUS S P E C . JUV.
+
MONTACUTA TENELLA
M O N TTENE
+
+
+
M YATRUNC
+
MYATRUNCATA
MYRIOCHELE DANIELSSENI
MYSELLA BIDENTATA
+
MYSIA UNDATA
+
+
+
+
+
+
+
+
+
+
+
+
M YRIHEER
+
+
+
+
+
+
+
+
+
+
+
+
+
+ M YSEBIDE
MYSIUNDA
+
+
+
NEM ATODA
NEMATODA
+
+
+
+
+
+
+
+
+
+
+
+
+
NEPHTYS CAECA
+
NEPHTYS C IR R O SA
NEPHTYS HOMBERGII
+
+
+
+
+
+
+
+
+
NEPHCAEC
+
+ N E PH C IR R
+
+
+
+
+
+
+
+
+
+
+
+
NEPHTYS S P E C . JU V .
N EREIS LONGISSIMA
+
+
NEPHSPEC
+
+
+
+
+
N O TO LA TE
+
+
+
NUCUNITI
+
+
N ER EL O N G
+
NOTOM ASTUS LATERICEU S
+
+
NUCULA NITIDOSA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
N UCU TEN U
+
NUCULA TENUIS
O D O SSPEC
+
ODOSTOMIA S P E C .
O LIGO CH A
OLIGOCHAETA
OPHELIA LIMACINA
+
+
+
+
+
OPHELINA ACUMINATA
OPHIURA TEXTURATA
+
OPHIURA S P E C . JU V .
+
+
+
+
+
OPHELIM A
O PH EA C U M
+ O PH IFLEX
+ OPHIALBI
+
O PH IT EX T
+
+
+
O P H IS P E C
O R B ISE R T
+
ORCHHUM I
+
+
+
+
O R CHNANA
+
ORCH SPEC
+
ORCHOM ENE S P E C . JUV .
OWENIA FU SIFO RM IS
+
+
+
+
+
OPHIURA ALBIDA
+
+
+ +
+
OPHIODROM US FL EX U O SU S
ORCHOM ENE NANA
N EPH H O M B
NEPHINCI
NEPHTYS INCISA
ORCHOM ENE HUMILIS
+ NEM ERTIN
N EPH A SSI
NEPHTYS ASSIMILIS
ORBINIA SERTULATA
+ MAGEMIRA
M EDIFRAG
+
+
MELITA OBTUSATA
NEMERTINI
co
O
+
+
+
O W E N FU SI
+
+
PARAONIS FU LG EN S
48
+
+
PARAFULG
Appendix-1 Biomonitoring 2001 (+=presence)
Do gg er Ban
E
S E
1
2
3
E E
E
5
6
4
0 /ster Ground
C
E
7
M
os
$ § £ $
1
2
3
4
5
<0
co
co
co
co
os
co
S $ £ S S s E ê s S $ $ $
6 7 8 9 10 11 12 13 14 15 16 17 18
Species name
Code
PECTINARIA AURICOMA
+
PECTINARIA KORENI
PERIO CU LO D ES LONGIM ANUS
+
+
+
+
+
+
+
+
PH O LO E MINUTA
+
+
+
+
PHORONIDA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
PE C T K O R E
+
+
+
+
PECTAURI
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
PERILO N G
PHOLMINU
+
PHYLLODOCE GROENLANDICA
PH O RO N ID
PH Y LG RO E
PHYLLODOCE MACULATA
PHYLLODOCE RO SEA
PHYLMACU
PH Y LR O SE
+
PHYLLODOCIDAE
+
+
+
+
PHYLLODO
PO D A R K EO PSIS HELGOLANDICA
PO DAHELG
POECILO CHA ETU S S E R P E N S
POLYDORA SP E C .
+
+
+
+
+
PO LYN O E KINBERGI
+
+
+
+
+
+
PO ECSERP
POLYDORA
+
POLYKINB
+
PO N TO C R A TES ALTAMARINUS
PO N T ALTA
PO N TO PH ILU S B ISPIN O SU S
PO N T B IS P
PR IO N O SP IO CIRRIFERA
PR IO C IR R
P R O C E S S A EDULIS C R A S S IP E S
PRO CED CR
P R O C E S S A NOUVELI HOLTHUISI
P R O C E S S A PARVA
PSEU D O CU M A LONGICORNIS
PR O C N O H O
PR O C PA R V
+
+
+
+
+
ROXANIA UTRICULUS
+
ROXAUTRI
SABELLA PENICILLUS
SCALIBREGMA INFLATUM
SC O LE LE PIS BONNIERI
+
SA B EPEN I
SCALINFL
+
+
SC O L O PL O S ARMIGER
+
+
+
+ SC O LBO N N
+
+
+
+
+
+
SEMIERYCINA NITIDA
+
+
+ SCOLARMI
+
SIGALION MATHILDAE
+
+
S P IO FILICORNIS
SP IO PH A N E S BOMBYX
PSEU LO N G
+
+
+
+
+
+
+
+
+
SEMINITI
+
+
+
+
+
+
+
+
+
+
+ +
SPIO PH A N E S KROEYERI
+
+
+
+
SPISU LA ELLIPTICA
+
+
+
+
+
+
SIGAMATH
SPIOFILI
+
+ SPIO BO M B
SP IO K R O E
+
SPISELLI
SPISU LA SOLIDA
SP ISSO L I
SPISU LA SUBTRUNCATA
S P IS S U B T
SPISU LA SP E C . JUV.
S P IS S P E C
STHENELAIS LIMICOLA
SYLLIDAE
+
+
+
+
+
+
+
+
+
+
+
+
+
SYLLIDAE
SYNCHELIDIUM H A PLO C H ELES
SYNCHELIDIUM MACULATUM
SYNCHAPL
+
+
+
SYNELMIS KLATTI
+
TELLIMYA FERUGINOSA
+
TELLINA FABULA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ SYNCMACU
SYNEKLAT
TELLFER U
TELLFABU
TELLPYGM
+
TELLINA TENUIS
THARYX KILLARIENSIS
THIA SCUTELLATA
+
TELLTENU
THARKILL
TH IASCU T
THRACIA CONVEXA
THYASIRA FLEXUOSA
+
+
TELLINA PYGMAEA
THRACIA PHASEOLINA
STHELIMI
+
+
+
+
+
+
+
+
+
+
+
+
+
+
TH RACONV
+ TH R A PH A S
THYAFLEX
TRACHYTHYONE ELONGATA
TR A CELON
TRAVISIA FORBESII
TR A V FO R B
49
Appendix-1 Biomonitoring 2001 (+=presence)
Do ggerBan k
O /ster Ground
E E E E EE E
1
2
3
4
5
6
7
u>
CO
(S' ê (S' ê «1 <§ ê $
1
2
3
4
5
6
7
8
co
co
co
co
ë
ê <1
ë
ë
9
10
12
13 14 15 16 17 18
11
d $ (S' (S' Ç
S p ecies name
C ode
TRID O N TA TRIANGULARIS
TURBELLARIA
TRIDTRIA
+
+
TURBONILLA PUSILLA
+
+
+
U PO G EB IA STELLATA
+
+
+
+
+
+
+
+
+
+
+
+
50
U PO G ST E L
+
UROTBREV
+
+
+ 1
U PO G D ELT
+
+
U R O T H O E BREVICORNIS
VITREOLINA ANTIFLEXA
TURBELLA
+
TU R B PU SI
U PO G EB IA DELTAURA
U R O T H O E POSEIDONIS
+
+
+
+
+
U R O TPO SE
VITRANTI
Appendix-1 Biomonitoring 2001 (+=presence)
0 /s te r Ground
CO
&
<1
(S'
co
CO
CO
CO
co
co
S
(S'
S
S
§ S S
(S'
§ ê (S' 1
27
19 20
21
22
23
24
25
26
+
+
+
+
+
+
+
28
29
30
31
32
+
+
+
+
+
33
34
co
co
CO
39
40
CO
(S
1 £ S S S d (S'
35
36
37
38
41
42
+
+
S p ecies nam e
ABRA ALBA
C ode
+
+
+
+
ABRA PRISM ATICA
+
ACANTHOCARDIA ECHINATA
ABRAPRIS
ACANECHI
+
ACIDOSTOM A OBESUM
ACIDOBES
ACROCNIDA BRACHIATA
+
ALTENAEUM DAWSONI
+
+
AMPEBREV
+
+
ACROBRAC
ALTEDAW S
+
AMPELISCA BREVICORNIS
AMPELISCA TENUICORNIS
AM PH A RETE FINMARCHICA
ABRAALBA
+
+
+
+
AMPETENU
+
AMPHFINM
AM PHILOCHUS SP E C .
AMPHILSP
AMPHIOXUS LANCEOLATUS
AMPHIURA CHIAJEI
AMPHLANC
AMPHIURA FILIFORMIS
AMPHCHIA
+
+
+
ANTHOZOA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
AONIDES PAUCIBRANCHIATA
APH ELOCHAETA MARIONI
AMPHFILI
ANTHOZOA
+
+
AONIPAUC
APHEMARI
+
A PH ER U SA BISPINOSA
+
A PH ERBIS
A PH ER U SA OVALIPES
+
APHEOVAL
A PH ER U SA S P E C .
+
A PH E S P E C
A PH R O D ITE ACULEATA
APHRACUL
APLACO PH O RA
+
ARCTICA ISLANDICA JUV.
+
APLACOPH
+
+
ARICIDEA MINUTA
ARICMINU
A STERIA S R U BEN S
A S T R O P E C T E N IRREGULARIS
ARCTISLA
+
A STER U BE
+
+
+
A STRIRRE
ATYLUS FALCATUS
ATYLFALC
ATYLUS SWAMMERDAMI
ATYLSWAM
BATHYPOREIA ELEGANS
+
BATHYPOREIA GUILLIAMSONIANA
BIVALVE INDET.
+
+
+
+
+
+
+
+
+
+
+
+ BATHELEG
+
BATHGUIL
+
B R IS S O P S IS LYRIFERA
+
CALLIANASSA S P E C . JUV.
+
+
+
CALLIANASSA SUBTERRANEA
+
+
+
BIVAINDE
BRISLYRI
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
C A LU U V E
+
+
+ CALLSUBT
CALLIANASSA TYRRHENA
CALLTYRR
CAPITELLA CAPITATA
CAPICAPI
CAPRELLIDAE
CAPRELLI
CERIA N TH U S LLOYDII
CERILLOY
C H A E T O PT E R U S VARIOPEDATUS
C H A ETO ZO N E SETO SA
+
+
CHAMELEA STRIATULA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
CHAEVARI
+
+
+
+
+
CHAMELEA S P E C . JUV.
+
+ + + +
+ + + +
+
+
+
+
+
+
+
+
+
+
+
+
CO RO PH IU M INSIDIOSUM
C O R Y S T E S CASSIVELAUNUS
CRANGON CRANGON
CUCUMARIA FRONDOSA
+
CHAMSTRI
C H AM SPEC
CHONE DUNERI
CORBULA GIBBA
+ C H A E SE TO
+
+
+
+
CHONDUNE
+ CORBGIBB
+ COROINSI
C O R Y C A SS
CRANCRAN
CU CU FR O N
+
CULTELLUS PELLUCIDUS
CYLICHNA CYLINDRACEA
+ +
DIASTYLIS BRADYI
+
+
+
+
+
+
+
+
+
+
+
+
+
+
51
+
+
+
+
+
+
CULTPELL
+
CYLICYLI
DIASBRAD
Appendix-1 Biomonitoring 2001 (+=presence)
Oyster Ground
d
d
d
d
d
d
d
d
d
d
m
d
d
d
d
d
d
d
d
d
d
d
d
d
d
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
C ode
S p ecies nam e
+
+
D IPLO CIRR U S GLAUCUS
+
+ +
+
+
DIPLGLAU
DONAVITT
DONAX VITTATUS
DOSINIA EXOLETA
DOSIEXOL
+
+
DOSINIA LUPINUS
+
+
ECHINOCARDIUM CORDATUM
+
DOSILUPI
+
+
EBALIA CRANCHII
ECHINOCYAM US PUSILLUS
EDW A RDSIA CLAPAREDII
+
+
+
+
+
+
+
+
+
EBALCRAN
+
+
+ ECH ICO RD
ECHIPUSI
+
+
+
EDW ACLAP
+
ENSIAMER
E N S IS AMERICANUS
ENSIARCU
E N S IS ARCUATUS
+ ENSIENSI
E N S IS EN SIS
E N S IS PHAXOIDES
ENSIPHAX
E T EO N E BARBATA
ETEOBARB
ETEO LON G
E T EO N E LONGA
+
+
EUDORELLA TRUNCATULA
+
E U D O R EL LO PSIS DEFORMIS
+
+ + +
+
EU D O D EFO
EUMISANG
+
+
EUMIDA SANGUINEA
EUDOTRUN
+
+
EU SPC A T E
E U SPIR A CATENA
+
E U SPIR A NITIDA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ EUSPNITI
EUZOFLAB
EU Z O N U S FLABELLIGERUS
EX O GHEBE
+
E X O G O N E H EBES
GOULDIA MINIMA
GOULMINI
GARI COSTULATA
G A RICO ST
GARIFERV
GARI FE R V E N SIS
+ +
GATTYANA CIRROSA
+
G ATTCIRR
GLYCLAPI
GLYCERA LAPIDUM
GLYCROUX
G LYCERA ROUXI
+
+ +
+
G LYCERA S P E C . JUV.
G L Y C SPE C
GLYCNORD
GLYCINDE NORDMANNI
+
G OLFINGIA ELONGATA
+
GOLFINGIA VULGARIS
+ +
GONIADA MACULATA
GOLFELO N
+
+ +
+ +
+
+ +
+ + + + + + +
GOLFINGIA PRO CERA
GO LFPRO C
GOLFVULG
+ + + +
+
+
GONIADELLA BOBRETZKII
+ + + + +
G Y PT IS C A PEN SIS
+
+ +
+
+
GONIMACU
GO NIBOBR
G Y PTC A PE
HARM OTHOE GLABRA
HARMGLAB
HARM OTHOE IMBRICATA
HARMIMBR
HARM OTHOE LJUNGMANI
HARM U U N
HARM OTHOE LUNULATA
HARMLUNU
HARM OTHOE S P E C . JUV.
HARPINIA ANTENNARIA
+ + + + +
+ + + +
+
H A RM SPEC
HARPANTE
+ + + + + + + +
H ET ER O M A ST U S FILIFORMIS
HETEFILI
HIATELLA ARCTICA
H IPPO M ED ON DENTICULATUS
HIATARCT
HYALA VITREA
HIPPD EN T
+
+
+ +
+
+
HYALVITR
HYPERIID
HYPERIIDAE S P E C .
IONE THORACICA
+ + +
+
+
+
IONETHOR
IPHIOBES
IPHIMEDIA O B ESA
KELLIA SUBORBICULARIS
+
+
+
52
+
+
+
IPHINOE TR ISPIN O SA
+
IPHITRIS
KELLSUBO
Appendix-1 Biomonitoring 2001 (+=presence)
Oyster Ground
«0
S
19
1
(0 ce
5“ S
ce
S
$
20
21
23
24
22
ce
§
ce
S
1
ce
$
1
1
§
§
§
25
26
27
28
29
30
31
32
33 34
1
ce
S
ê
ce
ce
$
$
35
36
37 38 39
1
ce
ce
S
<1
$
40
41
42
Species nam e
C ode
+
LANICE CONCH ILEG A
+
LANICONC
LANICE S P E C . JUV.
LANIJUVE
LEM BOS L O N G IPES
LEPTON SQ UAM OSUM
+
LEMBLONG
+
LE U CO TH O E INCISA
+
+
+
+
+
LEPTSQ U A
+
LEUCINCI
LEVINSENIA GRACILIS
LEVIGRAC
LIOCARCINUS S P E C . JUV.
LIO C S PE C
LUMBRINERIS FRAGILIS
+
LUMBRINERIS LATREILLI
+
+
+
LUMBFRAG
+
+
+
+
LUMBLATR
+
LYSILLA LOVENI
+
LYSILOVE
MACOMA BALTHICA
MACOBALT
MACTRA CORALLINA
+
MAGELONA ALLENI
+
MAGELONA MIRABILIS
+
M EDIOM ASTUS FRAGILIS
MACTCORA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
MAGEALLE
+
+
+ MAGEMIRA
+
MEDIFRAG
M EG A LU R O PU S AGILIS
MEGAAGIL
MELITA O B TU SA TA
MELITOBTU
M IC R O PR O T O P U S MACULATUS
MICRMACU
MODIOLUS S P E C . JUV.
M O D ISPEC
MONTACUTA TENELLA
M ONTTENE
MYA TRUNCATA
MYATRUNC
MYRIOCHELE DANIELSSENI
MYSELLA BIDENTATA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
MYRIHEER
+ MYSEBIDE
MYSIA UNDATA
MYSIUNDA
NEMATODA
NEMERTINI
+
+
+
+
+
+
+
+
+
+ + +
+
+
+
+
NEPH TY S ASSIMILIS
+
+
+
NEMATODA
+ NEMERTIN
+
NEPH TY S CA ECA
NEPH TY S C IR R O SA
NEPH TY S HOMBERGII
+
+
+
+
+
+
+
NEPH TY S INCISA
+
+
+
+
N EPHA SSI
+
N EPH C IR R
+
+
+
+
+
+
+
+
+
+ +
+
NEPH TY S S P E C . JUV .
+
+
N O TO M A STU S LATERICEUS
NUCULA NITIDOSA
NEPHHOM B
NEPHINCI
+
+
+
+
+
+
+
+
+
+ +
+ +
+ + + +
+
+ + +
+ + + + + + +
+ + + +
N ER EIS LONGISSIM A
N EPH C A EC
NEPHSPEC
NERELO N G
NOTOLATE
+ + + + +
NUCUNITI
NUCULA TE N U IS
NUCUTENU
ODOSTO M IA S P E C .
OD O SSPEC
OLIGOCHAETA
OLIGOCHA
OPHELIA LIMACINA
OPHELIMA
+
OPHELINA ACUMINATA
O PH IO D R O M U S FLEXUOSUS
+
OPHIURA ALBIDA
+
+
+
+
+
+ +
O PH EA CU M
+
+ +
OPHIFLEX
+
OPHIALBI
OPHITEX T
OPHIURA TEXTURATA
OPHIURA S P E C . JUV.
+
+
+
+ +
+
+
+
ORBINIA SERTU LA TA
O R CH O M EN E HUMILIS
O R CH O M EN E NANA
ORCHHUMI
+
+
+
+
ORCHNANA
O R CH O M EN E S P E C . JUV.
OW ENIA FU SIFO R M IS
PA RAONIS FU LG EN S
O P H IS P E C
O R B ISER T
O R CH SPEC
+
+
+
53
+
+
+
+ +
O W EN FU SI
PARAFULG
Appendix-1 Biomonitoring 2001 (+=presence)
O /ster Ground
d
d
d d
19
20
21
22
m
d
d
O d
d d d d d d d
23
24
25
27
26
28
29
30
31
32
33
m
d
d d d d d d d d
34
35
36
37
38
39
40
41
42
C ode
S pecies nam e
PECTINARIA KORENI
+
+
PECTINARIA AURICOMA
+
+
+
+
+
PE R IO C U L O D E S LONGIMANUS
PHO LO E MINUTA
+
PHORONIDA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ +
+
+
+
+
+
+
+
+
PECTAURI
+
PE C T K O R E
+
+
+
+
+
+
PERILONG
+
+
+
+
PHOLMINU
+
+
+
+
+ PHO RO N ID
PHY LG RO E
+
PHYLLODOCE GROENLANDICA
PHYLMACU
PHYLLODOCE MACULATA
PH Y LR O SE
+
PHYLLODOCE R O SE A
PHYLLODO
PHYLLODOCIDAE
PODAHELG
P O D A R K EO PSIS HELGOLANDICA
+
PO ECILO C H A ETU S SE R P E N S
POLYDORA S P E C .
+
+
+
+
+
+
+
+
+
+
+
+ POECSERP
+
+
POLYDORA
POLYKINB
+
POLYNOE KINBERGI
P O N T ALTA
P O N TO C R A TES ALTAMARINUS
P O N T B IS P
+
PO N TO PH ILU S B ISPIN O SU S
+
P R IO N O SPIO CIRRIFERA
PR IO CIRR
+
PR O C ED C R
+
P R O C E S S A EDULIS C R A SSIPE S
PR O C N O H O
P R O C E S S A NOUVELI HOLTHUISI
PR O C PA RV
+
P R O C E S S A PARVA
PSEU LO N G
+
+
PSEU D O CU M A LONGICORNIS
ROXAUTRI
ROXANIA UTRICULUS
SABELLA PENICILLUS
SABEPENI
SCALINFL
+
SCALIBREGMA INFLATUM
SCOLBONN
SC O LE LE PIS BONNIERI
+
+
+ +
+
+
+
SC O L O PL O S ARMIGER
+
SCOLARMI
+
SEMINITI
SEMIERYCINA NITIDA
SIGALION MATHILDAE
S P IO FILICORNIS
SPIO PH A N E S BOMBYX
+
+
+
+
+
+
SIGAMATH
+
+
+
+
+
+
+
+
+
+ SPIOFILI
+ SPIO BO M B
SPIO K R O E
+
SP IO PH A N E S KROEYERI
SPISULA ELLIPTICA
SPISELLI
+
SP ISSO L I
SPISULA SOLIDA
S P IS S U B T
+
+
SPISULA SUBTRUNCATA
S P IS S P E C
+
SPISU LA S P E C . JUV.
+
STHENELAIS LIMICOLA
+ +
+ +
+
+ + + + +
+
STHELIMI
SYLLIDAE
SYLLIDAE
SYNCHAPL
SYNCHELIDIUM HAPLOCHELES
+ +
+
+
+
SYNCHELIDIUM MACULATUM
+
+
SYNELMIS KLATTI
TELLIMYA FERU G IN O SA
+ +
TELLINA FABULA
+ +
+
+ +
+
+
+
+
SYNCMACU
SYNEKLAT
TELLFERU
+ +
TELLFABU
TELLINA PYGMAEA
TELLPYGM
TELLINA T EN U IS
TELLTENU
THARYX KILLARIENSIS
THARKILL
THIA SCUTELLATA
THRACIA CONVEXA
THIASCUT
THRACONV
THRACIA PHASEOLINA
THYASIRA FLEXUOSA
TRACHYTHYONE ELONGATA
+ +
+
+ + +
+ +
+ +
+ + +
+
+
+
+
TH RA PH A S
THYAFLEX
TRACELON
TR A V FO RB
TRAVISIA FO RBESII
54
Appendix-1 Biomonitoring 2001 (+=presence)
¡ o /ster Ground
ñ
«
w
V)
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
m
w
6
d d d d ó
36
37
38
d d d d d d d d d d d d d d d d d
V)
39
40
41
42
Species nam e
C o de
TRIDTRIA
TRIDONTA TRIANGULARIS
TURBELLARIA
+
TURBELLA
+
TURBONILLA PUSILLA
UPOGEBIA DELTAURA
+
+
TU R B PU SI
U PO G D ELT
+ +
+
UPOGEBIA STELLATA
U PO G ST E L
+
+
UR OTH O E BREVICORNIS
UR OTH O E PO SE ID O N IS
VITREOLINA ANTIFLEXA
+
+
55
UROTBREV
U ROTPO SE
VITRANTI
Appendix-1 Biomonitoring 2001 (+=presence)
0 1 F is h o r e a r e a
g
1
o
g S
2
3
8
8
8
8
4
5
6
7
8
+
+
8
9
S g g g S g S g
10
11
12
13
+
+
14
15
16
17
5
g g
18
19
g
20 21
g g g g g
22
23
24
25
ABRA ALBA
26
C ode
Species n am e
+
ABRAALBA
A B RA PRIS
ABRA PRISM ATICA
ACANECHI
ACANTHOCARDIA ECHINATA
ACID O BES
+
ACIDOSTOM A OBESU M
ACROCNIDA BRACHIATA
A C RO BRAC
ALTENAEUM DAWSONI
ALTEDAW S
AMPELISCA BREVICORNIS
AM PEBREV
AM PELISCA TEN U ICORN IS
AM PETENU
A M PHARETE FINMARCHICA
AMPHFINM
AM PHILSP
AM PHILOCHUS S P E C .
+
AMPHIOXUS LANCEOLATUS
+
AMPHIURA CHIAJEI
+
+
+ +
+
+
AMPHLANC
+
AMPHCHIA
AMPHFILI
AMPHIURA FILIFORMIS
+ +
ANTHOZOA
ANTHOZOA
A ONIDES PAUCIBRANCHIATA
AONIPAUC
A PH ELO CH A ETA MARIONI
APHEMARI
A PH ER B IS
A PH ER U SA B ISPIN O SA
APHEOVAL
+
A PH ER U SA O VALIPES
A PH ER U SA S P E C .
APH ESPEC
A PH RO D ITE ACULEATA
APHRACUL
APLA C O PH O R A
APLACO PH
ARCTISLA
ARCTICA ISLANDICA JUV.
+
ARICIDEA MINUTA
A STER IA S R U B E N S
A S T R O P E C T E N IRREGULARIS
+
ATYLUS FA LCATUS
+
+
+
+
+
+
A ST ER U B E
ATYLFALC
BATHYPOREIA ELEGANS
+
+
BATHYPOREIA GUILLIAMSONIANA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ + + + +
+
+
+
+
+
+
+
+
BATHELEG
+
+
+
BATHGUIL
BIVAINDE
BIVALVE INDET.
BRISLYRI
B R IS S O P S IS LYRIFERA
CALLIANASSA S P E C . JUV.
+
CALLIANASSA SU BTERRANEA
+
C A LU U V E
+
CALLSUBT
CALLTYRR
CALLIANASSA TYRRHENA
CAPICAPI
CAPITELLA CAPITATA
+
+
CAPRELLIDAE
CAPRELLI
CERILLOY
CERIA N TH U S LLOYDII
CHAEVARI
+ +
+ + + + +
+
+
+
CHAMELEA STRIATULA
+
C H A E SE TO
CHAM STRI
C H A M SPEC
CH ON D U N E
CHAMELEA S P E C . JUV.
CH ON E DUNERI
C O RO PH IU M INSIDIOSUM
ATYLSWAM
+
+
CORBULA GIBBA
ARICMINU
A STRIRR E
ATYLUS SWAMMERDAMI
C H A E T O PT E R U S VARIOPEDATUS
C H A ETO ZO N E S E T O S A
+
+
+
+
+
+
C O RBG IBB
+
C O RO IN SI
C O R Y S T E S CASSIVELAUNUS
C O R Y C A SS
C RA N G O N C RA N G O N
CRANCRAN
CUCUMARIA FR O N D O SA
C U C U FR O N
CU LTPELL
CULTELLUS PELLUCIDUS
CYLICHNA CYLINDRACEA
DIASTYLIS BRADYI
+
+ + +
CYLICYLI
+
DIASBRAD
56
Appendix-1 Biomonitoring 2001 (+=presence)
0 F fsh o re a re a
S s
5
5
E E S B B B B B S E
2
3
4
5
1
6
7
8
9
10
h
12
13
14
O
E E B S E E S S B B B
15
16
17
18
19
20
21 22
23
24
25
26
S p ecies nam e
C odé
DIPLO CIRRU S GLAUCUS
DIPLGLAU
DONAX VITTATUS
+
+
+
+
+ +
+
+
+
DOSINIA EXOLETA
DOSIEXOL
+
DOSINIA LUPINUS
DOSILUPI
EBALIA CRANCHII
EBALCRAN
+
ECHINOCARDIUM CORDATUM
ECHINOCYAMUS PU SILLUS
DONAVITT
+
+ +
+
+ +
+
+
+
+
+
ECHICORD
+
ECHIPUSI
EDW ARDSIA CLAPAREDII
EDW ACLAP
+
EN SIS AMERICANUS
+
ENSIAMER
+
EN SIS ARCUATUS
ENSIARCU
EN SIS EN SIS
ENSIENSI
EN SIS PHAXOIDES
ENSIPHAX
ET EO N E BARBATA
ETEOBARB
+
ET EO N E LONGA
+
+
EUDORELLA TRUNCATULA
EUDOTRUN
EU D O R EL LO PSIS D EFO R M IS
EUMIDA SANGUINEA
EU D O D EFO
+ + +
+
+
EUMISANG
EU SPIR A CATENA
EU SPIR A NITIDA
ETEO LON G
EU SPC A T E
+
+ + + +
+
+
+
+
+
EU ZO N U S FLABELLIGERUS
EX O G O N E H EBES
+
+
EUSPNITI
EUZOFLAB
EX O G H EBE
GOULDIA MINIMA
GOULMINI
GARI CO STULATA
G A RICO ST
GARI FE R V E N SIS
GARIFERV
GATTYANA C IR RO SA
G ATTCIRR
+
GLYCERA LAPIDUM
GLYCLAPI
+
GLYCERA ROUXI
GLYCROUX
+
GLYCERA S P E C . JUV.
+
G LY C SPE C
GLYCINDE NORDMANNI
GLYCNORD
GOLFINGIA ELONGATA
GOLFELO N
GOLFINGIA PR O C ER A
G O L FP R O C
GOLFINGIA VULGARIS
GONIADA MACULATA
GOLFVULG
+ + +
+ + + +
GONIMACU
GONIADELLA BOBRETZKII
G O NIBOBR
+
G Y PTIS C A PE N SIS
G Y PTC A PE
HARM OTHOE GLABRA
HARMGLAB
HARM OTHOE IMBRICATA
HARMIMBR
+
HARM OTHOE LJUNGMANI
+
HARM OTHOE LUNULATA
HARM OTHOE S P E C . JU V .
HARMLJUN
HARMLUNU
+
H A RM SPEC
+
HARPINIA ANTENNARIA
H ETER O M A STU S FILIFORM IS
HARPANTE
HETEFILI
HIATARCT
HIPPD EN T
HIATELLA ARCTICA
H IPPO M ED ON DENTICULATUS
HYALA VITREA
HYALVITR
HYPERIIDAE S P E C .
HYPERIID
IONE THORACICA
IONETHOR
IPHIMEDIA O B ESA
IPHINOE TR ISPIN O SA
IPHIO BES
+
+
IPHITRIS
KELLIA SUBO RBICU LA RIS
KELLSUBO
57
Appendix-1 Biomonitoring 2001 (+=presence)
0 1 F fs h o r e a r e a
S
B B B
1
2
3
4
5
B B B B B s B B B B B B S B B B
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
S
B B B B
22
23
24
25
LANICE CONCHILEGA
+
+
+
LANICONC
LANIJUVE
+
LANICE S P E C . JUV.
LEMBLONG
LEMBOS LO N G IPES
LE PT SQ U A
LEPTON SQUAMOSUM
LEU CO TH OE INCISA
26
C ode
Species nam e
+
+
+
+
+
+
+
LEUCINCI
LEVINSENIA GRACILIS
LEVIGRAC
LIOCARCINUS S P E C . JUV.
LIO C S PE C
LUMBRINERIS FRAGILIS
LUMBFRAG
LUMBRINERIS LATREILLI
LYSILLA LOVENI
LUMBLATR
MACOMA BALTHICA
MACOBALT
MACTRA CORALLINA
M ACTCORA
LYSILOVE
MAGEALLE
MAGELONA ALLENI
MAGELONA MIRABILIS
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
MAGEMIRA
MEDIOM ASTUS FRAGILIS
+
M EGALUROPUS AGILIS
MELITA OBTUSATA
M IC R O PR O TO PU S MACULATUS
+
+
+
+
MEDIFRAG
+ MEGAAGIL
MELITOBTU
+
MICRMACU
+
MODIOLUS S P E C . JUV.
M O D ISPE C
MONTACUTA TENELLA
M ON TTENE
MYATRUNCATA
MYATRUNC
M YRIHEER
MYRIOCHELE DANIELSSENI
+
MYSELLA BIDENTATA
+
MYSEBIDE
+
+
MYSIUNDA
MYSIA UNDATA
+
NEMATODA
NEMERTINI
+
+
+
+
+
+
NEMATODA
+
+
+
+
+
+
+
+
+
+
+
N EPH TY S CAECA
+
+
N EPH TY S C IRRO SA
N EPHTY S HOMBERGII
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ + + + +
+
+
+
+
+
+
+
+
N EPHH O M B
NEPHINCI
N EPHTY S INCISA
N EPHTY S S P E C . JUV.
NER EIS LONGISSIMA
NOTO M A STU S LATERICEUS
+ N EPH C A EC
+ N EPH C IR R
+
+
NEM ERTIN
N EPH A SSI
+
N EPH TY S ASSIMILIS
+
+
+
+
N EPHSPEC
N ER ELO N G
+
N O TOLATE
+
NUCUNITI
+
NUCULA NITIDOSA
NUCULA TENUIS
N UCUTENU
ODOSTOM IA S P E C .
O D O SSPEC
O LIGOCHA
OLIGOCHAETA
+
+
OPHELIA LIMACINA
OPHELINA ACUMINATA
O PH IFLEX
O PH IO D RO M U S FLEX U O SU S
OPHIURA ALBIDA
OPHIURA TEXTURATA
+
OPHIALBI
+
O PH ITEX T
+
+
OPHIURA S P E C . JUV.
+
+
+
O P H IS P E C
+
O R B ISE R T
ORBINIA SERTULATA
ORCHHUM I
O R CH O M EN E HUMILIS
OR CH O M EN E NANA
+
+
+
+
OR CHNANA
ORCH SPEC
OR CH O M EN E S P E C . JUV.
OW ENIA FUSIFORM IS
OPHELIM A
O PH EA C U M
O W E N FU S I
+
+
PARAONIS FULGENS
58
+
+ PARAFULG
Appendix-1 Biomonitoring 2001 (+=presence)
01ff s h o re a r e a
S B B S B B B S B B
1
2
3
4
5
6
7
8
9
10
s
S B B B B B B B B S S B B B
11
12
13
14
15
16
17
18
19 20
21
22
23
24
25
S pecies n a m e
C ode
PECTINARIA AURICOMA
PECTAURI
PECTINARIA KORENI
PE R IO C U L O D E S LONGIMANUS
PECTKORE
+
+
PH O LO E MINUTA
PHORONIDA
+
+
+
+
+
+
+
+
PERILO N G
+
PHOLMINU
+
+
+
PHO RO N ID
PH Y LLODOCE GROENLANDICA
PHY LG RO E
PHY LLO D O CE MACULATA
+
PHYLMACU
PH Y LLODOCE R O SE A
PH Y LR O SE
PHYLLODOCIDAE
PHYLLODO
P O D A R K E O PSIS HELGOLANDICA
P O E C ILO C H A E TU S S E R P E N S
POLYDORA S P E C .
+
+
+
+
PODAHELG
+
+
PO ECSERP
POLYDORA
PO LYN O E KINBERGI
P O N T O C R A T E S ALTAMARINUS
5
26
POLYKINB
+
PO N T ALTA
PO N TO PH ILU S B ISPIN O SU S
P O N T B IS P
P R IO N O SP IO C IR RIFER A
PR IO C IR R
P R O C E S S A EDULIS C R A S SIPE S
PR O C E D C R
P R O C E S S A NOUVELI HOLTHUISI
PR O C N O H O
P R O C E S S A PARVA
PR O C PA R V
PSEU D O CU M A LONGICORNIS
+
+
+
+
+
+
+
+
+
+
+
+
PSEU LO N G
ROXANIA U TRICULUS
ROXAUTRI
SABELLA PENICILLUS
SCALIBREGM A INFLATUM
SCALINFL
SA BEPEN I
SC O L E L E PIS BONNIERI
+
S C O L O P L O S ARMIGER
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
SCOLARMI
SEM IERYCINA NITIDA
SEMINITI
SIGALION MATHILDAE
S P IO FILICORNIS
SP IO P H A N E S BOMBYX
+
+
+
+
+
+
+
+
+
SIGAMATH
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
SP IO P H A N E S KROEYERI
SPIOFILI
SPIO BO M B
SP IO K R O E
SPISU LA ELLIPTICA
+
SPISELLI
SPISU LA SOLIDA
+
SPISU LA SU BTRU N CA TA
+
SP ISSO L I
+
S P IS S U B T
SPISU LA S P E C . JUV .
S P IS S P E C
STHEN ELA IS LIMICOLA
+
STHELIMI
SYLLIDAE
+
SYNCHELIDIUM H APLOCHELES
SYLLIDAE
+
SYNCHELIDIUM MACULATUM
SYNCHAPL
+
+
+
+ + + +
+
+
SYNCMACU
SYNELMIS KLATTI
SYNEKLAT
TELLIMYA FE R U G IN O SA
+
+
TELLINA FABULA
+
+
+
+
+
+
+
+
+
+
+
+
+
TELLFERU
+
TELLFABU
TELLPYGM
TELLINA PYGM AEA
TELLINA T E N U IS
+
TELLTENU
+
THARKILL
THIASCU T
THARYX KILLARIENSIS
THIA SCUTELLATA
+
+
+
THRACIA CO NVEXA
THRACIA PHA SEO LIN A
SCO LB O N N
TH RACONV
+
+
+
TH R A PH A S
THYASIRA FLEX U O SA
+
THYAFLEX
TRA CELON
TRA CH YTH Y O NE ELONGATA
TRAVISIA FO R B ESII
+
59
TRA V FO RB
Appendix-1 Biomonitoring 2001 (+=presence)
01f f s h o r e a r e a
S b B B B B B B B
1
2
3
4
5
6
7
8
9
5
B B B B B B B B S S g S S S g g
10 h
12
13
14 15 16 17 18 19 20 21
22
23 24
25 26
Species n am e
C ode
TRIDONTA TRIANGULARIS
TRIDTRIA
+
TURBELLA
TURBELLARIA
+
TURBONILLA PUSILLA
TU R B PU SI
UPOGEBIA DELTAURA
U PO G D ELT
U PO G ST E L
UPOGEBIA STELLATA
U R OTH O E BREVICORNIS
UROTH O E PO SE ID O N IS
VITREOLINA ANTIFLEXA
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
U ROTBREV
U ROTPO SE
VITRANTI
60
Appendix-1 Biomonitoring 2001 (+=presence)
01 f s h o r e a r e a
C o a s ta l a re a
§ § B Q B B B B B B I a 8 a a a
27 28 29 30 31
32 33 34 35 36
1
2
3
4
5
6
%
7
8
a a a a a a
9
10 h
12 13
14
S pecies nam e
%
15
C ode
ABRA ALBA
+
ABRAALBA
ABRA PRISMATICA
ABRA PRIS
ACANTHOCARDIA ECHINATA
ACANECHI
A CIDOSTOM A O B ESU M
ACID O BES
ACROCNIDA BRACHIATA
ACRO BRAC
ALTENAEUM DAWSONI
AMPELISCA B REV ICO RN IS
ALTEDAW S
AM PEBREV
AMPELISCA T EN U IC O R N IS
AM PETENU
AMPHARETE FINMARCHICA
AMPHFINM
AMPHILOCHUS S P E C .
AM PHILSP
+
AMPHIOXUS LANCEOLATUS
AMPHIURA CHIAJEI
AMPHLANC
AMPHCHIA
AMPHIURA FILIFORMIS
AMPHFILI
ANTHOZOA
ANTHOZOA
A ONIDES PAUCIBRANCHIATA
APHELOCHAETA MARIONI
AONIPAUC
APHEMARI
A PH ERU SA B ISPIN O SA
APH ER B IS
A PH ER U SA OVALIPES
APHEOVAL
A PH ER U SA S P E C .
APH ESPEC
A PHRO D ITE ACULEATA
APHRACUL
A PLACO PH O RA
A PLACO PH
ARCTICA ISLANDICA JUV.
ARCTISLA
ARICIDEA MINUTA
+
+
+
+
ARICMINU
A STERIA S R U BEN S
A STER U BE
A ST R O PE C T E N IRREGULARIS
A STRIRR E
ATYLUS FALCATUS
+
ATYLUS SWAMMERDAMI
+
BATHYPOREIA ELEGANS
BATHYPOREIA GUILLIAMSONIANA
ATYLFALC
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
ATYLSWAM
+
+
+
+ BATHELEG
+ BATHGUIL
+
BIVALVE INDET.
BIVAINDE
B R IS S O P S IS LYRIFERA
CALLIANASSA S P E C . JUV.
BRISLYRI
+
+
CALUUVE
CALLIANASSA SU B TERR A N EA
CALLSUBT
+
CALLIANASSA TY RRH EN A
CALLTYRR
CAPITELLA CAPITATA
+
+
CAPRELLIDAE
+
+
+
+
+
+
+
CAPRELLI
C ERIANTHUS LLOYDII
CERILLOY
C H A E T O PT E R U S VA RIO PED A TUS
CHAEVARI
C H AETO ZON E S E T O S A
CHAMELEA STRIATULA
CAPICAPI
+
+
+
+
+
+
+
+ C H A E SE T O
CHAM STRI
CHAMELEA S P E C . JUV.
C H A M SPEC
CHONE DUNERI
C H ON D U N E
CORBULA GIBBA
C O RBG IBB
CO RO PH IU M INSIDIOSUM
C O RO IN SI
+
C O R Y ST E S CASSIV ELA U N U S
CRANGON CRA N G O N
+
C O R Y C A SS
CRANCRAN
CUCUMARIA FR O N D O SA
C U C U FR O N
CULTELLUS PELLUCIDUS
CU LTPELL
CYLICHNA CYLINDRACEA
CYLICYLI
+
DIASTYLIS BRADYI
61
+
DIASBRAD
Appendix-1 Biomonitoring 2001 (+=presence)
C o a s ta l a re a
8 8 8 8 B B B B B B a
28 2 9 30 31 32 33 3 4 35 36
i
27
COA
01P fs h o re a r e a
2
a a
3
4
a
5
a a a a %a a 3 3 a
6
7
8
9 10 11
12
13
14
15
C ode
S p ecies nam e
DIPLGLAU
DIPLO CIRRU S GLAU CU S
+
DONAX VITTATUS
+
+
+
+
+
DONAVITT
DOSINIA EXOLETA
D OSIEXOL
DOSINIA LUPINUS
DOSILUPI
EBALCRAN
EBALIA CRANCHII
+
ECHINOCARDIUM CORDATUM
ECHINOCYAMUS PUSILLUS
+
+
+
+
+
+
+
+
+
+
+
EC H IC O R D
ECH IPUSI
+
ED W ACLAP
EDW ARDSIA CLAPAREDII
EN SIS AM ERICANUS
+
+
+
+
+
+
+
+
+
+ ENSIAM ER
EN SIS ARCUATUS
ENSIARCU
EN SIS EN SIS
EN SIEN SI
EN SIS PHAXOIDES
ENSIPHAX
ETEO B A RB
ETEO N E BARBATA
+
ETEO N E LONGA
+
+
+
+
+
EUDORELLA TRUNCATULA
E U D O D E FO
EU D O R EL LO PSIS D EFORM IS
EUMIDA SANGUINEA
+
+
EUMISANG
+
+
E U S PC A T E
+
EU SPIR A CATENA
EU SPIRA NITIDA
ET EO L O N G
EUDO TRU N
+
+
+
+
+
+
+
+
+
+
+
+
+ EUSPNITI
EUZOFLAB
EU ZO NU S FLABELLIGERUS
E X O G H EB E
+
EX O G O N E H EB ES
GOULDIA MINIMA
GOULMINI
GARI CO STULATA
G A R IC O ST
GARI FE R V E N SIS
G A RIFER V
GATTYANA C IR R O SA
G A TTCIRR
GLYCERA LAPIDUM
GLYCLAPI
GLYCROUX
GLYCERA ROUXI
+ +
GLYCERA S P E C . JU V .
G LY CSPEC
GLYCINDE NORDMANNI
G LY CN O RD
GOLFINGIA ELONGATA
G O LFELO N
GOLFINGIA P R O C E R A
GO LFPRO C
GOLFVULG
GOLFINGIA VULGARIS
+
GONIADA MACULATA
GONIMACU
G O N IB O BR
GONIADELLA BOBRETZKII
+
G Y PTIS C A PE N SIS
G Y PT C A PE
HARMGLAB
HARM OTHOE GLABRA
HARM OTHOE IMBRICATA
+
HARMIMBR
H A RM U U N
HARM OTHOE LJUNGMANI
+ +
+ +
HARM OTHOE LUNULATA
+
HARM OTHOE S P E C . JUV .
+
H A R M SPEC
HETEFILI
HIATARCT
H IPPD EN T
HIPPO M ED ON DENTICULATUS
HYALA VITREA
HARMLUNU
H ARPANTE
HARPINIA ANTENNARIA
H ETER O M A STU S FILIFORMIS
HIATELLA ARCTICA
+
+
HYALVITR
HYPERIIDAE S P E C .
HYPERIID
IONE THORACICA
IO N ETH O R
IPH IO BES
IPHIMEDIA O B ESA
IPHINOE TR IS PIN O SA
+
IPHITRIS
KELLSUBO
KELLIA SUBO RBICU LA RIS
62
Appendix-1 Biomonitoring 2001 (+=presence)
Olffshore area
Coastal area
<o
E 8 8 S 8 8 8 8 8 8 3 %
O8 3 3 O
3 3 3 1 ? ? ? 8 3
27 28
29
30
31
32
33
34 35
+
+
36
1
2
3
4
5
6
7
8
9
10 11
12
13
14
15
Species name
Code
+ +
LANICE CONCHILEGA
+
+ +
LANICONC
LANICE S P E C . JUV.
LANIJUVE
+
LEMBOS LO N G IPES
LEMBLONG
LEPTON SQUAM OSUM
L E PT SQ U A
+
LEU CO TH O E INCISA
+
LEUCINCI
LEVINSENIA GRACILIS
LEVIGRAC
LIOCARCINUS S P E C . JUV.
L IO C S PE C
LUMBRINERIS FRAGILIS
LUMBFRAG
LUMBLATR
LUMBRINERIS LATREILLI
LYSILLA LOVENI
LYSI LOVE
+
MACOMA BALTHICA
+
+ +
MACOBALT
+
MACTRA CORALLINA
M ACTCORA
MAGELONA ALLENI
MAGELONA MIRABILIS
MAGEALLE
+ +
+
+ +
+ + + +
+ +
+
+
MEDIOM ASTUS FRAGILIS
MAGEMIRA
M EDIFRAG
+
M EG A LU RO PU S AGILIS
+ + + +
+
MELITA OBTU SA TA
MEGAAGIL
M ELITOBTU
M IC R O PR O T O P U S MACULATUS
MICRMACU
MODIOLUS S P E C . JUV.
MONTACUTA TENELLA
M O D ISPEC
M ONTTENE
+
MYA TRUNCATA
MYATRUNC
MYRIOCHELE DANIELSSENI
M YRIHEER
+
+
MYSELLA BIDENTATA
+
+ +
+ +
MYSIA UNDATA
MYSIUNDA
NEMATODA
NEMERTINI
N EPH TY S ASSIMILIS
N EPH TY S CAECA
N EPH TY S C IR R O SA
N EPH TY S HOMBERGII
M YSEBIDE
NEMATODA
+ + +
+
+
+
+
+
+
+
+
+ + + + + + + + + + + +
+
+
+ + +
+ + +
+
NEMERTIN
N EPH A SSI
+
+
+ + +
+ + + + + + + + +
+ + + +
+ +
+
+ +
N EPH TY S INCISA
N EPH C A EC
N EPH C IR R
N EPHH O M B
NEPHINCI
+
N EPH TY S S P E C . JUV .
+
+ +
+
N EREIS LONGISSIM A
+ + +
+
+
+
NO TO M A STU S LA TERICEU S
+ +
+
+ +
NUCULA NITIDOSA
N EPHSPEC
N ER ELO N G
N O TOLATE
NUCUNITI
NUCULA TEN U IS
NUCUTENU
ODOSTOM IA S P E C .
O D O SSPEC
+ +
OLIGOCHAETA
+
OPHELIA LIMACINA
OPHELINA ACUMINATA
+
O PH IO D R O M U S FL EX U O SU S
O PHIURA ALBIDA
+
O PHIURA TEXTURATA
OPHIURA S P E C . JUV.
OPHELIMA
OPH EA CU M
O PHIFLEX
+
+
+
+
+
+
+ +
+
O R B ISE R T
O R CH O M EN E HUMILIS
ORCHHUM I
+
+
+
+ + +
ORCHNANA
O R CH O M EN E S P E C . JUV .
OW ENIA FU SIFO RM IS
PARAONIS FU LGEN S
OPHIALBI
O PH ITEX T
O P H IS P E C
ORBINIA SERTULATA
O R CH O M EN E NANA
OLIGOCHA
ORCH SPEC
+
+
O W EN FU SI
PARAFULG
63
Appendix-1 Biomonitoring 2001 (+=presence)
Olffs hore area
8 B B B B B B
S B
27
Coastal area
s
28 29 30 31
32
33
34 3 5 36
t
1
§
%
2
3
i
4
%
5
%
6
s
7
8
8
aa
9
10
s
11
i
12
aaa
13
14
15
Species name
Code
PECTINARIA AURICOMA
PECTA U RI
+
PECTINARIA KORENI
PE R IO C U L O D E S LONGIMANUS
+
PH OLOE MINUTA
+
PHORONIDA
+
PE C T K O R E
+
PE R ILO N G
PHOLMINU
+
+
PH O R O N ID
+
PHYLLODOCE GROENLANDICA
+
+
PH Y LG R O E
PHYLMACU
PHYLLODOCE MACULATA
+
PHYLLODOCE R O SEA
PH Y L R O SE
+
PHYLLODOCIDAE
PHYLLODO
PO D A H ELG
PO ECSER P
PO D A R K EO PSIS HELGOLANDICA
PO ECILO CHA ETU S S E R P E N S
+
+
+
PO LYDORA
POLYDORA S P E C .
POLYKINB
POLYNOE KINBERGI
+ + + +
PO N TO C R A TES ALTAMARINUS
PO N T ALTA
PO N TO PH ILU S BISPIN O SU S
PO N T B IS P
PR IO N O SPIO CIRRIFERA
PR IO C IR R
PRO CED CR
P R O C E S S A EDULIS C R A S S IP E S
+
P R O C E S S A NOUVELI HOLTHUISI
PR O C N O H O
P R O C E S S A PARVA
+
+
+
+ + + +
PR O C PA R V
ROXANIA UTRICULUS
PS E U L O N G
ROXAUTRI
SABELLA PENICILLUS
SA B EPEN I
PSEU D O CU M A LONGICORNIS
SCALINFL
SCALIBREGMA INFLATUM
+
SC O LE LE PIS BONNIERI
SC O L O PL O S ARMIGER
+
+ +
+ +
+ + +
+
+ +
+
+
+
+ + +
+
+
SIGAMATH
SIGALION MATHILDAE
+ +
+
S P IO FILICORNIS
+
+ + + + + +
+
+ +
+ +
+ + +
+
+
+ +
+ +
+
+
SPISULA SOLIDA
SPISU LA SUBTRUNCATA
+
+ +
SP ISSO L I
+
+ +
S P IS S U B T
+
SPISULA S P E C . JUV.
S P IS S P E C
+
+
STHENELAIS LIMICOLA
+
+
SYLLIDAE
STHELIMI
SYLLIDAE
SY N C H A PL
SYNCHELIDIUM HAPLOCHELES
+
+ + +
+
SYNCHELIDIUM MACULATUM
+ + + + +
SYNCM ACU
SYNEKLAT
SYNELMIS KLATTI
TELLIMYA FERU G IN O SA
+ +
+
+
+
+ +
+ +
+ + + + +
+ + + +
+ + +
TELLFER U
+ +
TELLFABU
TELLPYGM
TELLTEN U
TELLINA T EN U IS
THARKILL
THARYX KILLARIENSIS
+
THIA SCUTELLATA
+
TH IASCU T
TH RA CON V
THRACIA CONVEXA
+
TH R A PH A S
THYAFLEX
THYASIRA FLEXUOSA
TR A C ELO N
TRACHYTHYONE ELONGATA
TRAVISIA FORBESII
SPIO B O M B
SPISELLI
SPISU LA ELLIPTICA
THRACIA PHASEOLINA
SPIO FILI
SP IO K R O E
SPIO PH A N E S KROEYERI
TELLINA FABULA
TELLINA PYGMAEA
SCOLARMI
SEMINITI
SEMIERYCINA NITIDA
SPIO PH A N E S BOMBYX
SC O LB O N N
+
+
T R A V FO R B
64
Appendix-1 Biomonitoring 2001 (+=presence)
Offshore area
Coastal area
S
O 8 S S S S 5 8 8
27 28 29 30 31 32 331 341 35 36
3
5
1 2
<
o
3 4
S
S p ecies nam e
TRIDONTA TRIANGULARIS
TURBELLARIA
<O <o : <o <o <o
O! OI OI OI O
a a s
5
7 8 9 10 11 12 13 14 15
8
3
S
6I
+
C ode
TRIDTRIA
TURBELLA
TURBONILLA PUSILLA
TU R B PU SI
U PO G EBIA DELTAURA
U PO G D ELT
U PO G EBIA STELLATA
U PO G ST E L
U R O TH O E BREVICORNIS
U R O TH O E PO SE ID O N IS
VITREOLINA ANTIFLEXA
+ +
+
+
+
+
+
+
+
+
+ + + +
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+ UROTBREV
+
+ U R O TPO SE
VITRANTI
65
A ppendix 2 , B iom onitoring 2001
station
DOG 1
Crustacea
N
atylfalc
12.8
batheleg
641.5
bathguil
12.8
caprelli
coroinsi
38.5
corycass
diasbrad
12.8
ebalcran
harpante
hyperiid
iphiobes
iphitris
leuci nci
12.8
liocspec
12.8
megaagil
12.8
orchhumi
orchnana
perilong
12.8
procparv
pseulong
12.8
syncmacu
25.7
urotpose
513.2
Echinodermata
acrobrac
12.8
amphchia
echicord
12.8
echipusi
ophialbi
ophispec
295.1
ophitext
12.8
Mollusca
arctisla
12.8
chamspec
chamstri
25.7
cultpell
cyticyli
dosiexol
dosilupi
ensiensi
ensiphax
25.7
euspniti
89.8
gariferv
12.8
goulmini
mactcora
12.8
mysebide
12.8
mysiunda
12.8
tellfabu
102.6
tellferr
25.7
thraphas
51.3
thyaflex
turbpusi
vitranti
Polychaeta
chaeseto
25.7
eteolong
gonimacu
51.3
gyptcape
harmspec
laniconc
mageaile
12.8
magemira
115.5
nephhomb
nephspec
B
0.004
0.192
0.004
0.012
DOG 2
N
B
115.5
0.035
12.8
25.7
0.004
0.008
DOG 3
N
25.7
295.1
25.7
B
0.008
0.089
0.008
102.6
12.8
0.031
0.387
DOG 4
N
B
DOG 5
N
B
1295.8
128.3
0.389
0.038
12.8
12.8
0.004
0.004
38.5
12.8
0.012
1.311
12.8
0.004
12.8
0.647
25.7
25.7
0.008
0.008
51.3
0.015
12.8
0.004
12.8
0.106
12.8
0.004
77.0
346.4
25.7
6.222
0.251
16.855
12.8
0.001
12.8
0.038
12.8
0.657
25.7
0.004
25.7
12.8
77.0
0.000
0.003
0.010
0.004
12.8
25.7
0.004
0.004
0.004
0.004
0.004
0.008
0.154
0.506
115.5
0.035
38.5
0.004
0.008
12.8
12.8
0.004
0.004
25.7
0.008
0.012
51.3
0.015
25.7
12.8
128.3
0.008
0.004
0.038
12.8
1.103
12.8
0.000
128.3
0.038
154.0
0.046
64.2
25.7
5.732
0.003
230.9
1.115
38.5
0.015
12.8
64.2
0.465
0.001
25.7
0.001
12.8
12.8
0.000
0.000
25.7
12.8
0.317
0.030
12.8
0.078
12.8
12.8
0.354
0.034
9.057
0.010
2.539
0.001
0.005
0.489
0.295
0.001
0.006
0.001
0.004
0.546
0.030
0.005
0.012
0.115
0.208
0.051
38.5
0.006
243.8
0.053
25.7
12.8
25.7
0.020
0.010
0.151
12.8
0.019
12.8
0.607
38.5
0.079
77.0
0.008
25.7
12.8
0.107
0.001
12.8
25.7
0.002
0.002
102.6
0.033
179.6
25.7
12.8
51.3
12.8
0.236
0.021
0.001
0.010
0.018
25.7
0.001
12.8
0.038
25.7
0.012
12.8
0.012
51.3
0.042
25.7
25.7
12.8
51.3
0.134
0.012
0.005
1.028
51.3
0.046
12.8
25.7
0.010
0.022
25.7
38.5
12.8
141.1
38.5
12.8
0.024
2.075
0.039
0.097
0.036
0.012
243.8
12.8
38.5
0.203
0.154
0.032
12.8
474.7
0.010
0.083
102.6
0.046
12.8
0.010
77.0
0.091
66
A ppendix 2, Biom onitoring 2001
nerelong
notdate
ophelima
orbisert
owenfusi
pectkore
pholminu
phylrose
scolamii
scolbonn
25.7
sigamath
spiobomb
25.7
spiofili
sthelimi
Miscellaneous taxa
nemertin
25.7
edwaclap
12.8
cerilloy
amphlanc
phoronid
sum
2335.1
diversity
nspc
35
SH-W
2.51
Simp
0.14
station
DOG6
Crustacea
N
ampebrev
12.8
atylfalc
12.8
atylswam
batheleg
603.0
bathguil
141.1
calljuve
callsubt
coroinsi
64.2
diasbrad
ebalcran
eudodefo
harpante
iphitris
orchnana
12.8
orchspec
perilong
64.2
pseulong
upogspju
urotbrev
urotpose
77.0
Echinodermata
acrobrac
25.7
269.4
amphchia
amphfili
asterube
brislyri
25.7
echicord
ophispec
12.8
12.8
0.125
0.010
12.8
1.458
51.3
12.8
0.022
0.239
12.8
25.7
0.503
0.012
0.207
0.012
0.008
0.014
14.6
0.181
0.042
0.019
0.987
0.024
12.8
12.8
0.012
0.012
12.8
0.012
243.8
12.8
0.220
0.012
51.3
2.827
25.7
0.234
51.3
0.042
25.7
0.022
12.8
89.8
0.434
0.075
38.5
0.239
12.8
51.3
0.544
0.042
77.0
0.029
12.8
0.173
12.8
0.323
38.5
0.006
64.2
0.051
64.2
12.8
0.164
0.062
25.7
12.8
12.8
0.040
0.042
4.787
12.8
25.7
1693.6
0.034
0.003
8.9
38.5
1796.2
0.019
6.6
128.3
2873.9
0.152
7.9
64.2
1886.0
0.010
31
2.69
0.11
B
0.004
0.004
51.3
25.7
DOG7
N
12.8
12.8
38.5
538.9
102.6
77.0
25.7
35
3.06
0.06
B
0.004
0.004
0.012
0.162
0.031
0.023
0.008
37
2.39
0.22
OYS 1
N
B
25.7
0.008
12.8
0.450
25.7
12.8
0.008
0.010
64.2
0.019
12.8
0.035
40.7
41
3.13
0.07
OYS 2
N
12.8
B
0.004
25.7
12.8
38.5
12.8
0.008
0.004
0.720
1.120
25.7
12.8
0.008
0.004
12.8
0.004
OYS 3
N
25.7
B
0.008
25.7
0.008
12.8
0.004
25.7
12.8
12.8
25.7
0.008
0.004
0.004
0.008
25.7
0.008
2052.8
8.891
12.8
12.8
10.451
3.949
0.004
0.019
12.8
12.8
25.7
0.004
0.004
0.008
0.023
12.8
0.004
0.349
0.011
192.5
0.011
1988.7
12.8
12.8
8.214
0.003
7.502
5.071
67
590.2
0.217
12.8
12.8
8.411
0.000
Appendix 2, B iom onitoring 2001
Mollusca
abraalba
aplacoph
arctisla
12.8
chamstri
corbgibb
cultpell
12.8
cylicyti
dosiexol
12.8
ensiensi
12.8
gariferv
12.8
hyalvitr
12.8
kellsubo
monttene
mysebide
89.8
nucuniti
12.8
nucutenu
roxautri
tellfabu
51.3
tellferr
12.8
telltenu
25.7
thyaflex
tridtria
Polychaeta
aonipauc
aphemari
chaeseto
12.8
chaevari
diplglau
12.8
gattcirr
gonimacu
25.7
gyptcape
harmspec
lysilove
magemira
230.9
medifrag
nephcaec
nephcirr
25.7
nephspec
notolate
ophiflex
owenfusi
320.8
pectauri
12.8
pectkore
pholminu
phylspec
12.8
poecserp
12.8
polydora
polykinb
scolarmi
scolbonn
25.7
sigamath
64.2
spiobomb
77.0
spiofili
sthelimi
Miscellaneous taxa
turbella
nemertin
12.8
12.8
0.002
25.7
25.7
0.008
0.063
423.4
0.148
38.5
0.007
38.5
0.019
25.7
0.020
38.5
38.5
795.5
0.002
0.007
0.098
38.5
0.014
12.8
12.8
0.038
0.006
12.8
0.037
12.8
0.007
25.7
0.014
0.003
25.7
0.002
0.001
12.8
0.040
0.014
11.872
0.000
0.001
12.8
0.014
12.8
0.001
0.006
0.002
0.001
0.006
0.004
0.007
12.8
12.8
38.5
0.001
0.007
38.5
64.2
0.051
0.186
12.8
12.8
12.8
0.112
0.010
0.010
0.108
154.0
0.203
0.986
0.007
25.7
38.5
38.5
12.8
0.005
0.000
38.5
0.010
12.8
12.8
51.3
12.8
12.8
0.003
1.160
0.102
0.725
0.003
12.8
0.698
25.7
0.007
0.008
12.8
0.021
0.025
12.8
0.025
25.7
0.014
282.3
0.559
12.8
25.7
0.007
0.014
12.8
0.911
12.8
0.025
25.7
38.5
0.014
1.394
38.5
0.708
38.5
12.8
115.5
0.217
0.203
0.061
12.8
25.7
0.098
0.014
12.8
25.7
0.005
0.102
0.020
0.073
25.7
0.020
12.8
0.010
128.3
0.034
12.8
12.8
0.103
0.003
0.007
0.005
0.146
1.194
0.036
0.054
0.046
0.005
0.058
230.9
12.8
12.8
77.0
25.7
12.8
0.010
0.113
0.061
0.020
0.044
12.8
0.069
12.8
0.011
12.8
0.014
68
51.3
12.8
25.7
0.102
0.025
0.051
89.8
12.8
0.178
0.025
12.8
0.062
A ppendix 2, Biomonitoring 2001
edwaclap
phoronid
sum
diversity
nspc
SH-W
Simp
154.0
2578.8
0.034
20.6
36
2.78
0.10
station
OYS 4
Crustacea
N
ampebrev
ampetenu
12.8
amphilsp
12.8
batheleg
89.8
bathguil
38.5
calljuve
calisubt
caprelli
12.8
corycass
12.8
diasbrad
ebalcran
eudotrun
harpante
12.8
hippdent
12.8
hyperiid
ionethor
iphitris
leucinci
25.7
megaagil
orchnana
perilong
12.8
syncmacu
12.8
urotpose
12.8
Echinodermata
acrobrac
12.8
amphfili
307.9
astrirre
ophialbi
Mollusca
abraalba
102.6
aplacoph
chamstri
25.7
corbgibb
cultpell
12.8
cylicyli
dosiexol
dosilupi
12.8
euspniti
hyalvitr
kellsubo
25.7
mactcora
modispec
12.8
monttene
myatrunc
mysebide
89.8
mysiunda
nucuniti
64.2
tellfabu
397.7
thraphas
64.2
thyaflex
179.6
vitranti
38.5
1744.9
0.007
1.5
35
2.74
0.12
B
0.004
0.004
0.027
0.012
0.030
19.5
30
1.67
0.40
OYS 5
N
B
12.8
0.004
OYS 6
N
25.7
12.8
795.5
2424.9
0.017
3.397
17.3
27
2.15
0.19
B
0.008
OYS 7
N
38.5
12.8
64.2
89.8
B
25.7
77.0
0.008
0.023
12.8
25.7
0.004
0.008
12.8
0.004
0.008
6.285
0.004
0.037
102.6
0.031
25.7
12.8
0.004
0.004
0.004
0.264
0.045
0.025
1180.4
3.251
12.8
0.037
307.9
0.083
295.1
0.081
0.634
0.004
2617.3
12.8
15.252
0.011
0.031
12.8
0.013
25.7
0.009
12.8
166.8
51.320
0.016
25.7
0.011
OYS 8
N
12.8
B
0.004
51.3
0.015
12.8
179.6
0.012
25.7
25.7
51.3
0.006
0.008
0.015
102.6
0.043
166.8
0.128
141.1
0.015
102.6
12.8
0.023
0.014
51.3
0.042
38.5
0.002
64.2
0.255
8.475
64.2
0.019
538.9
0.835
12.8
25.7
0.005
0.032
12.8
12.8
0.014
0.001
0.059
0.025
12.8
25.7
12.8
26.1
0.004
12.8
77.0
0.010
0.008
0.004
0.008
12.8
0.082
0.012
0.004
25.7
12.8
12.8
12.8
12.8
38.5
3707.9
36
1.77
0.35
0.797
3.474
0.004
2.367
0.004
0.004
115.5
3233.2
0.049
0.647
0.003
0.001
0.001
0.012
0.244
0.048
0.026
0.246
1706.4
0.182
12.8
0.001
25.7
0.049
69
12.8
0.002
51.3
12.8
12.8
12.8
0.006
0.025
0.004
0.001
A ppendix 2, Biom onitoring 2001
Polychaeta
amphfinm
aonipauc
aphracul
chaeseto
chaevari
diplglau
eumisang
gonimacu
gyptcape
harmlunu
laniconc
lumbfrag
lumblatr
magealle
magemira
medifrag
myriheer
nephcaec
nephcirr
nephhomb
notolate
opheacum
ophiflex
owenfusi
paragrac
pectkore
12.8
0.008
102.6
0.068
25.7
89.8
38.5
0.191
0.474
1.324
7.894
12.8
154.0
218.1
sthelimi
25.7
syneklat
12.8
Miscellaneous taxa
nemertin
38.5
anthozoa
edwaclap
phoronid
89.8
golfproc
golfvulg
sum
3169.0
diversity
nspc
40
SH-W
2.89
Simp
0.09
station
OYS 9
Crustacea
N
ampebrev
ampetenu
aphespec
batheleg
51.3
25.7
calljuve
callsubt
64.2
diasbrad
ebalcran
12.8
eudotrun
12.8
25.7
6.964
12.8
12.8
25.7
12.8
0.017
0.051
0.000
51.3
0.034
38.5
38.5
0.030
0.025
269.4
25.7
0.178
0.017
12.8
0.034
12.8
0.008
25.7
0.017
12.8
38.5
0.008
0.025
25.7
12.8
10.051
0.007
12.8
12.8
38.5
12.8
12.8
25.7
12.8
0.007
0.100
0.007
25.7
77.0
0.014
1.089
51.3
0.027
12.8
0.003
0.003
0.017
0.034
0.108
12.8
0.298
12.8
12.8
12.8
25.7
0.017
0.112
0.017
0.088
25.7
12.8
51.3
0.550
12.8
0.042
12.8
359.2
0.317
0.190
6.322
0.019
0.025
pholminu
phylmacu
poecserp
polydora
sabepeni
scolarmi
spiobomb
spiofili
0.017
0.017
0.008
12.8
38.5
718.5
12.8
12.8
0.008
51.3
0.068
25.7
0.034
12.8
0.061
0.102
0.178
0.361
0.008
12.8
0.017
89.8
102.6
0.362
0.054
12.8
12.8
0.066
0.007
12.8
0.003
12.8
0.003
12.8
12.8
0.003
0.003
0.325
0.041
15.8
25.7
12.8
12.8
2745.6
0.014
0.517
0.014
67.6
37
2.32
0.21
N
12.8
12.8
38.5
0.310
0.044
128.3
0.044
295.1
0.144
5658.0
25.8
1090.6
7.4
1911.7
20.2
OYS 11
B
B
12.8
0.004
12.8
38.5
77.0
38.5
0.004
0.090
3.237
0.012
25.7
12.8
0.090
1.468
0.010
0.004
12.8
0.004
OYS 13
OYS 12
N
0.004
0.015
0.017
1.357
29
2.88
0.07
23
2.08
0.26
35
1.74
0.31
OYS 10
B
0.110
70
N
B
12.8
0.004
12.8
51.3
0.015
2.496
N
B
12.8
0.004
12.8
38.5
0.008
0.012
A ppendix 2, Biomonitoring 2001
harpante
77.0
¡onethor
leucinci
12.8
perilong
upogdelt
urotbrev
12.8
urotpose
25.7
Echinodermata
amphfili
179.6
cucufron
echicord
12.8
echipusi
38.5
ophialbi
Mollusca
abraalba
77.0
abrapris
altedaws
chamstri
115.5
corbgibb
410.6
cultpell
cylicyli
euspniti
38.5
hyalvitr
mactcora
mysebide
25.7
nucuniti
359.2
seminiti
spiselli
teiltenthraconv
thraphas
12.8
thyaflex
Polychaeta
chaeseto
38.5
chaevari
gattcirr
gonlmacu
gyptcape
harmlunu
lumbfrag
lumblatr
magealle
12.8
magemira
179.6
medifrag
myriheer
12.8
nephcaec
nephhomb
nephspec
notolate
ophiflex
38.5
parafulg
pectkore
25.7
poecserp
pholminu
phylmacu
12.8
polydora
scolarmi
sigamath
38.5
spiobomb
51.3
spiofili
12.8
spiokroe
12.8
0.023
51.3
0.015
0.004
25.7
38.5
25.7
12.8
0.008
0.012
0.008
2.148
102.6
0.359
38.5
38.5
0.012
0.012
12.8
2.234
141.1
12.8
12.8
0.331
0.351
0.283
77.0
0.023
975.1
5.123
12.8
2.783
25.7
12.8
0.159
0.032
12.8
12.8
25.7
12.8
0.001
0.002
0.147
0.017
397.7
64.2
0.047
0.068
12.8
0.011
89.8
0.009
38.5
0.015
51.3
0.061
38.5
0.015
12.8
0.244
128.3
0.051
12.8
0.005
12.8
0.005
0.004
0.008
0.296
1193.2
4.173
9.737
0.003
0.002
0.009
0.173
0.853
744.1
0.170
12.8
0.001
500.4
0.369
38.5
0.010
12.8
77.0
12.8
12.8
25.7
25.7
12.8
0.002
0.131
0.013
0.031
0.002
0.006
0.001
0.416
0.004
0.774
77.0
692.8
141.1
0.081
0.089
38.5
25.7
487.5
0.002
0.001
0.188
320.8
0.321
12.8
12.8
0.004
12.8
0.001
38.5
12.8
12.8
0.054
1.782
1.172
0.003
0.020
12.8
12.8
12.8
38.5
0.056
0.088
0.007
12.8
12.8
12.8
0.005
0.005
0.005
25.7
0.124
12.8
0.740
0.009
0.007
0.012
0.020
51.3
0.097
38.5
51.3
0.020
0.027
12.8
12.8
0.005
77.0
0.030
12.8
0.029
12.8
10.475
12.8
0.014
25.7
12.8
0.193
0.014
38.5
0.041
12.8
12.8
12.8
0.737
0.102
0.014
12.8
25.7
12.8
12.8
0.014
0.027
0.014
0.014
0.007
0.198
0.014
0.001
0.007
0.142
0.027
0.007
0.071
12.8
25.7
0.007
0.014
12.8
25.7
0.014
12.8
12.8
25.7
71
0.014
0.014
0.014
0.027
A ppendix 2, Biomonitoring 2001
sthelimi
syneklat
Miscellaneous taxa
turbella
nemertin
38.5
anthozoa
edwaclap
phoronid
25.7
golfelon
sum
2065.6
diversity
nspc
32
SH-W
2.81
Simp
0.09
station
OYS 14
Crustacea
N
ampebrev
ampetenu
38.5
batheleg
bathguil
calljuve
12.8
callsubt
diasbrad
eudodefo
eudotrun
harpante
38.5
ionethor
leucinci
64.2
melitobtu
perilong
pseulong
syncmacu
upogdelt
upogstel
urotpose
Echinodermata
amphfili
12.8
echicord
echipusi
ophialbi
ophispec
Mollusca
abraalba
25.7
arctisla
12.8
chamstri
12.8
corbgibb
256.6
cultpell
cylicyli
euspniti
garicost
gariferv
hiatarct
hyalvitr
474.7
kellsubo
12.8
leptsqua
monttene
mysebide
12.8
nucuniti
51.3
odosspec
12.8
tellferr
12.8
0.042
12.8
0.007
0.158
0.019
13.8
12.8
5.253
51.3
0.015
2553.2
15.5
27
1.80
0.29
OYS 15
N
12.8
0.012
12.8
B
0.010
0.012
25.7
0.056
564.5
12.8
2784.1
0.166
0.039
8.0
0.014
0.027
12.8
0.025
38.5
0.022
12.8
12.8
0.003
0.006
25.7
64.2
0.037
2219.6
8.9
77.0
1642.2
0.952
19.5
34
2.63
0.14
34
2.47
0.15
B
0.004
0.004
12.8
25.7
OYS 16
N
B
25.7
0.008
38.5
51.3
0.123
1.821
12.8
51.3
12.8
0.004
3.578
0.004
12.8
77.0
25.7
0.004
0.023
0.008
77.0
51.3
0.023
0.015
OYS 17
N
12.8
0.022
28
2.16
0.23
B
0.004
OYS 18
N
B
0.023
0.008
0.058
12.8
0.004
77.0
25.7
89.8
77.0
12.8
25.7
0.023
0.004
0.008
12.8
12.8
0.004
0.004
12.8
0.004
12.8
0.004
0.019
12.8
12.8
25.7
0.004
0.004
0.000
0.001
0.175
0.475
0.001
0.001
0.040
0.006
243.8
25.7
25.7
1.341
1.593
0.417
12.8
564.5
25.7
0.004
38.5
12.8
0.012
0.004
0.117
0.911
0.340
141.1
0.042
12.8
0.004
89.8
12.8
12.8
0.034
10.451
0.001
641.5
0.792
64.2
0.001
64.2
0.001
12.8
0.000
12.8
0.000
564.5
12.8
25.7
51.3
0.138
0.204
0.003
35.608
12.8
12.8
0.001
1.566
64.2
0.011
0.014
0.011
12.8
12.8
0.204
0.013
12.8
0.011
12.8
0.004
1064.9
12.8
0.253
0.016
64.2
0.064
12.8
0.003
38.5
25.7
0.006
0.006
25.7
0.019
72
A ppendix 2, Biomonitoring 2001
thraconv
12.8
thraphas
Polychaeta
aphemari
chaeseto
25.7
chaevari
diplglau
eteobarb
eumisang
12.8
exoghebe
gatteirr
glycnord
gonimacu
12.8
gypteape
12.8
harmglab
harmspec
lumbfrag
25.7
lumblatr
magemira
medifrag
nephcaec
25.7
nepheirr
nephhomb
12.8
nephspec
notolate
51.3
opheacum
opheliju
ophiflex
38.5
parafulg
51.3
pectauri
pectkore
12.8
pholminu
poeeserp
25.7
scolarmi
scolbonn
sigamath
spiobomb
25.7
spiokroe
25.7
spiofili
sthelimi
syneklat
tharkill
12.8
Miscellaneous taxa
nemertin
cerilloy
12.8
phoronid
77.0
golfvulg
25.7
sum
1539.6
diversity
nspe
33
SH-W
2.70
Simp
0.13
station
OYS 19
Crustacea
N
ampebrev
ampetenu
12.8
calljuve
12.8
callsubt
12.8
diasbrad
12.8
eudodefo
5.697
0.020
0.010
0.010
0.010
12.8
77.0
38.5
12.8
12.8
25.7
12.8
25.7
0.012
0.071
8.259
0.012
0.012
0.400
0.047
0.024
12.8
12.8
38.5
0.028
38.5
0.025
25.7
0.020
51.3
0.034
12.8
25.7
0.010
38.5
0.030
12.8
12.8
0.024
436.2
0.594
38.5
0.030
12.8
0.032
384.9
12.8
12.8
0.450
205.3
0.659
1.208
0.020
0.249
0.010
0.054
12.8
0.012
12.8
0.012
25.7
12.8
77.0
25.7
0.100
0.012
0.984
0.134
12.8
38.5
0.098
0.036
12.8
128.3
0.108
38.5
0.115
12.8
12.8
0.008
0.071
38.5
0.025
12.8
64.2
0.008
0.095
0.003
0.513
0.032
2.520
0.193
0.041
12.8
12.8
0.003
0.019
38.5
0.335
0.010
12.8
0.020
12.8
0.003
0.012
0.020
0.020
51.3
12.8
12.8
0.014
0.044
0.012
0.010
0.276
12.8
64.2
0.120
0.042
12.8
12.8
0.044
0.008
77.0
0.071
102.6
0.206
0.010
11.214
0.036
0.200
21.6
51.3
25.7
1154.7
0.014
2.295
18.5
36
3.15
0.06
12.8
0.020
51.3
0.020
269.4
0.112
2283.7
7.3
1193.2
13.0
3323.0
39.4
27
1.92
0.28
B
OYS 20
N
B
0.004
0.006
0.289
0.004
77.0
38.5
25.7
0.210
3.747
0.008
31
3.09
0.05
OYS 21
N
B
12.8
154.0
0.015
6.693
73
OYS 22
N
12.8
25.7
12.8
12.8
31
2.58
0.11
B
0.004
0.718
0.004
0.004
OYS 23
N
B
12.8
0.004
12.8
0.206
A ppendix 2, Biom onitoring 2001
eudotrun
harpante
12.8
ionethor
leucinci
orchnana
pontbisp
procparv
syncmacu
upogdelt
Echinodermata
amphfili
513.2
asterube
astrirre
12.8
brislyri
12.8
cucuelon
cucufron
12.8
echicord
ophialbi
25.7
ophispec
Mollusca
abraalba
25.7
chamstri
corbgibb
38.5
cultpell
cylicyli
38.5
dosiexol
dosilupi
euspniti
hyalvitr
12.8
kellsubo
mactcora
mysebide
115.5
nucuniti
spissubt
tellfabu
thyaflex
12.8
Polychaeta
chaeseto
diplglau
glycspec
gonimacu
gyptcape
lumblatr
magealle
magemira
medifrag
myriheer
nephhomb
38.5
nephspec
nerelong
notolate
ophiflex
12.8
owenfusi
25.7
pectkore
pholminu
25.7
poecserp
polydora
scolarmi
sigamath
spiobomb
12.8
spiokroe
0.004
25.7
89.8
25.7
0.008
0.027
0.008
12.8
25.7
12.8
12.8
12.8
0.004
0.008
0.004
0.004
0.046
154.0
51.3
12.8
1.669
474.7
2.689
77.0
22.503
590.2
2.107
12.8
0.739
12.8
38.5
3.249
0.297
372.1
12.8
0.046
0.015
141.1
0.042
77.0
0.023
12.8
0.004
898.1
10.242
25.7
0.003
25.7
12.8
0.017
0.000
12.8
12.8
12.8
25.7
0.000
0.370
0.007
0.049
25.7
0.006
0.214
0.968
0.003
0.013
7.215
0.618
0.000
0.002
0.007
0.007
25.7
12.8
12.8
12.8
38.5
0.006
0.000
0.014
0.004
0.042
51.3
0.019
0.013
0.015
12.8
64.2
128.3
0.001
0.002
0.013
436.2
0.079
89.8
0.031
25.7
0.036
77.0
25.7
0.039
0.026
51.3
25.7
38.5
0.006
0.070
0.070
0.001
12.8
0.349
0.000
0.000
0.009
0.004
0.034
12.8
0.002
64.2
89.8
0.006
0.096
641.5
38.5
0.089
0.016
38.5
0.043
51.3
179.6
0.001
0.087
12.8
0.019
154.0
0.061
12.8
0.019
12.8
0.019
38.5
12.8
0.015
0.005
25.7
0.037
38.5
0.015
38.5
0.274
25.7
12.8
0.552
0.005
12.8
64.2
0.019
0.093
115.5
12.8
0.046
0.005
38.5
12.8
12.8
0.056
0.244
0.019
64.2
12.8
64.2
0.102
0.393
0.025
0.034
25.7
0.068
12.8
12.8
0.034
0.034
51.3
0.337
38.5
0.877
0.100
25.7
38.5
0.020
0.110
51.3
975.1
38.5
38.5
12.8
51.3
12.8
12.8
0.008
0.014
0.191
0.007
0.081
1.800
0.029
0.036
0.073
0.027
102.6
0.014
12.8
12.8
12.8
12.8
51.3
12.8
25.7
0.042
0.003
0.034
74
A ppendix 2, Biomonitoring 2001
sthelimi
syneklat
Miscellaneous taxa
nemertin
anthozoa
edwaclap
12.8
phoronid
38.5
golfvulg
sum
1052.1
diversity
nspc
23
SH-W
2.14
Simp
0.25
station
OYS 24
Crustacea
N
ampebrev
ampetenu
batheleg
51.3
bathguil
12.8
calljuve
callsubt
128.3
ebalcran
eudodefo
harpante
ionethor
iphitris
leucinci
orchnana
perilong
pseulong
12.8
upogspju
urotpose
Echinodermata
amphfili
12.8
astrirre
echicord
25.7
ophispec
Mollusca
abraalba
372.1
arctisla
12.8
chamstri
corbgibb
12.8
cultpell
12.8
cylicyti
dosilupi
euspniti
12.8
hyalvitr
12.8
leptsqua
mysebide
nucuniti
spissubt
spisspec
tellfabu
38.5
tellferr
thraphas
thyaflex
Polychaeta
chaeseto
chondune
eumisang
12.8
12.8
38.5
0.034
0.102
12.8
0.056
0.015
38.5
0.019
10.6
1372.8
8.4
26
2.58
0.14
B
OYS 25
N
603.0
12.8
3746.4
0.637
0.156
39.3
2.023
0.060
1.007
0.008
230.9
25.7
0.069
0.008
12.8
12.8
10.249
38.5
0.030
1321.5
13.3
32
2.82
0.10
OYS 26
N
B
51.3
0.015
0.015
0.004
38.5
38.5
25.7
0.037
0.019
0.107
35
2.50
0.13
B
25.7
12.8
38.5
1.007
12.8
0.004
12.8
0.004
12.8
0.004
0.054
12.8
0.017
12.8
0.011
64.2
0.061
12.5
33
2.45
0.16
OYS 27
N
12.8
B
0.004
38.5
0.012
12.8
77.0
12.8
0.004
0.023
OYS 28
N
B
12.8
0.004
38.5
0.012
12.8
25.7
0.004
0.008
12.8
0.004
12.8
0.004
12.8
0.000
51.3
0.002
0.004
0.004
12.8
0.032
0.013
1167.5
25.7
38.5
12.8
0.663
0.022
0.130
0.000
1026.4
2.284
0.003
12.8
0.005
1809.0
0.311
0.096
0.320
89.8
12.8
0.020
0.169
564.5
0.160
12.8
0.030
12.8
0.064
12.8
51.3
25.7
38.5
0.020
0.005
0.035
0.012
6.733
0.004
0.013
38.5
0.038
25.7
12.8
0.002
0.002
2.082
12.8
0.004
243.8
25.7
38.5
64.2
0.122
0.674
0.503
0.001
64.2
3297.3
0.002
0.880
51.3
64.2
0.004
0.019
12.8
12.8
12.8
0.010
0.002
0.013
12.8
0.004
77.0
0.020
64.2
0.106
12.8
0.001
12.8
12.8
0.003
0.045
0.002
25.7
12.8
0.024
0.012
0.029
75
12.8
12.8
0.002
0.012
12.8
0.007
A ppendix 2, Biom onitoring 2001
glycspec
gonimacu
38.5
0.030
gyptcape
12.8
0.020
lumbfrag
lumblatr
89.8
0.254
magemira
myriheer
nephcaec
nephcirr
nephhomb
nephinci
nephspec
nerelong
12.8
0.085
notolate
449.1 21.542
ophiflex
owenfusi
pectauri
pectkore
0.014
pholminu
12.8
phylgroe
poecsetp
spiobomb
51.3
0.041
sthelimi
syneklat
Miscellaneous taxa
nemertin
25.7
0.130
anthozoa
0.054
edwadap
12.8
phoronid
golfproc
golfvulg
33.7
sum
1437.0
diversity
nspc
23
SH-W
2.23
0.17
Simp
station
OYS 29
N
Crustacea
ampetenu
12.8
batheleg
12.8
calljuve
callsubt
corycass
12.8
diasbrad
12.8
B
0.004
0.004
12.8
38.5
12.8
0.037
12.8
0.012
12.8
0.012
12.8
0.144
38.5
0.019
2065.6
2.8
29
1.94
0.33
102.6
12.8
51.3
0.015
0.024
38.5
0.015
64.2
12.8
12.8
0.708
0.005
0.105
12.8
12.8
0.034
0.049
12.8
0.005
12.8
25.7
12.8
0.007
0.076
0.007
25.7
0.098
205.3
0.125
744.1
0.547
12.8
12.8
12.8
1.755
0.007
0.034
12.8
0.012
25.7
0.014
12.8
0.037
0.217
0.037
0.012
0.012
12.8
0.005
0.014
0.025
38.5
12.8
12.8
OYS 30
N
12.8
12.8
166.8
38.5
25.7
4169.8
0.132
0.307
0.098
5.9
OYS 31
N
12.8
0.004
12.8
77.0
6.788
64.2
0.007
0.378
0.047
12.8
0.007
12.8
0.012
12.8
0.025
25.7
12.8
0.040
9.109
51.3
0.054
12.8
0.007
4477.7
4.8
1270.2
10.5
32
1.33
0.55
28
1.82
0.27
B
12.8
12.8
12.8
B
0.004
0.004
0.160
1.080
10.034
0.004
76
23
1.87
0.35
OYS 32
N
B
12.8
0.004
25.7
2.229
OYS 33
N
B
25.7
38.5
0.033
1.214
A pp en d ix 2, Biomonitoring 2001
harpante
lemblong
orchnana
perilong
pseulong
upogstel
Echinodermata
amphfili
141.1
echicord
25.7
ophispec
Mollusca
abraalba
372.1
acanechi
12.8
arctisla
12.8
chamstri
38.5
corbgibb
12.8
cultpell
12.8
cylicyli
12.8
euspniti
kellsubo
12.8
mysebide
25.7
nucunlti
102.6
tellfabu
128.3
tellferr
25.7
thraphas
38.5
thyaflex
230.9
Polychaeta
aphemari
chaeseto
chaevari
diplglau
12.8
gatteirr
gonimacu
25.7
gyptcape
laniconc
lumblatr
magemira
320.8
nephcaec
25.7
nepheirr
25.7
nephhomb
12.8
nephspec
12.8
nerelong
notolate
opheacum
ophiflex
parafulg
pectauri
pectkore
pholminu
25.7
phylrose
poecserp
12.8
priocirr
scolarmi
128.3
sigamath
12.8
spiobomb
38.5
spiofili
12.8
sthelimi
Miscellaneous taxa
turbella
nemertin
25.7
anthozoa
0.031
19.878
0.174
11.417
0.001
0.001
0.002
0.111
0.089
0.001
0.003
0.216
0.009
0.034
0.006
0.226
154.0
0.046
12.8
25.7
38.5
0.004
0.008
0.012
2027.1
89.8
38.5
6.326
6.544
0.056
38.5
0.042
64.2
0.039
25.7
12.8
0.008
2.366
12.8
0.000
12.8
0.000
128.3
0.069
782.6
4.608
38.5
25.7
0.019
0.112
12.8
0.354
154.0
64.2
0.019
0.467
12.8
12.8
38.5
0.007
7.467
38.5
12.8
0.174
0.007
12.8
0.008
25.7
0.041
0.024
12.8
12.8
0.008
0.008
38.5
25.7
0.022
0.007
12.8
0.090
12.8
38.5
12.8
0.027
0.008
0.835
51.3
0.562
25.7
12.8
12.8
0.007
0.301
0.234
25.7
0.825
12.8
0.044
25.7
0.095
12.8
0.420
102.6
0.027
12.8
77.0
0.003
0.020
64.2
25.7
0.017
0.007
0.024
0.032
0.269
0.286
0.036
0.012
0.200
12.8
12.8
0.008
0.008
102.6
0.130
12.8
0.023
Tl
12.8
0.027
0.004
12.8
0.004
12.8
0.597
667.2
1.854
25.7
0.000
12.8
51.3
0.759
0.009
12.8
12.8
0.003
0.024
38.5
0.002
25.7
38.5
12.8
0.007
0.030
12.412
12.8
0.642
12.8
0.557
12.8
12.8
0.703
0.003
12.8
12.8
0.301
0.085
51.3
25.7
25.7
0.014
0.029
0.347
12.8
0.003
0.001
0.012
0.212
0.793
0.046
0.032
0.012
89.8
12.8
0.007
12.8
0.046
12.8
0.058
12.8
25.7
12.8
0.014
1.561
1.866
12.8
0.014
A ppendix 2 , Biomonitoring 2001
edwaclap
phoronid
golfvulg
golfelon
sum
diversity
nspc
SH-W
Simp
12.8
64.2
0.023
0.064
102.6
0.076
38.5
0.010
2027.1
44.3
680.0
14.4
3412.8
10.4
36
2.86
0.09
station
OYS 34
Crustacea
N
ampetenu
apheoval
apherbis
aphespec
batheleg
bathguil
64.2
calljuve
callsubt
154.0
corycass
diasbrad
eudotrun
25.7
harpante
25.7
ionethor
25.7
iphitris
leuci nci
12.8
perilong
12.8
proceder
upogdelt
Echinodermata
amphfili
38.5
cucuelon
echicord
ophialbi
Mollusca
abraalba
1950.2
altedaws
chamstri
corbgibb
25.7
cyllcyli
12.8
dosilupi
euspnlti
25.7
hyalvitr
leptsqua
25.7
mysebide
nucuniti
25.7
spiselli
tellferr
thraphas
turbpusl
Polychaeta
chaeseto
chaevari
dlplglau
exoghebe
gattclrr
glycspec
gonimacu
12.8
15
2.39
0.09
B
OYS 35
N
12.8
12.8
25.7
33
1.94
0.36
B
0.004
12.8
12.217
0.008
0.008
0.008
12.8
166.8
0.004
0.050
0.004
0.004
12.8
0.004
154.0
0.144
0.168
0.009
0.004
0.038
0.005
0.016
OYS 36
N
B
25.7
0.008
30
2.39
0.21
B
25.7
0.008
12.8
12.8
0.004
0.004
38.5
2.866
12.8
0.004
12.8
0.004
12.8
0.004
0.008
0.015
12.8
0.004
38.5
12.8
0.435
2.817
320.8
0.555
12.8
0.001
25.7
0.262
154.0
19.078
25.7
12.8
423.4
12.8
0.001
0.000
0.132
0.017
12.8
25.7
0.001
0.343
12.8
0.047
12.8
38.5
38.5
0.013
0.046
0.005
89.8
0.313
12.8
0.000
51.3
0.015
12.8
0.018
12.8
0.010
12.8
1.721
89.8
12.8
1.180
2.845
25.7
0.516
372.1
0.064
0.017
38.5
12.8
0.005
0.019
0.010
0.041
0.009
0.014
0.007
B
25.7
51.3
25.7
25.7
12.8
OYS 38
N
0.004
0.012
0.004
0.069
0.064
0.014
20.0
12.8
38.5
12.8
25.7
12.8
25.7
1488.3
25.7
0.017
0.003
0.025
0.068
0.178
0.179
2.340
77.0
25.7
OYS 37
N
102.6
77.0
38.5
51.3
0.183
51.3
12.8
0.014
22
1.81
0.32
384.9
25.7
25.7
0.246
0.307
0.083
22.3
0.004
0.008
0.275
11.141
0.088
243.8
51.3
12.8
1437.0
25.7
25.7
12.8
0.014
0.760
12.8
12.8
0.203
0.007
0.010
0.063
78
A ppendix 2, Biom onitoring 2001
gyptcape
25.7
lumblatr
38.5
magealle
magemira
12.8
medifrag
12.8
myriheer
nephassi
nephhomb
25.7
nephspec
nerelong
notolate
89.8
ophiflex
orbisert
parafulg
pectauri
pectkore
pholminu
poecserp
12.8
polydora
priocirr
12.8
spiobomb
sthelimi
Miscellaneous taxa
nemertin
12.8
phoronid
307.9
goltvulg
sum
2989.4
diversity
nspc
25
SH-W
1.54
Simp
0.44
station
OYS 39
Crustacea
N
ampetenu
38.5
atylswam
batheleg
bathguil
calljuve
callsubt
12.8
coroinsi
corycass
diasbrad
ebalcran
12.8
eudodefo
eudotrun
25.7
harpante
115.5
iphitris
leuci nci
micrmacu
perilong
pontalta
pseulong
syncmacu
urotbrev
urotpose
Echinodermata
acrobrac
amphfili
1103.4
0.049
0.032
0.005
0.005
12.8
154.0
12.8
12.8
0.005
0.005
25.7
89.8
12.8
0.010
0.036
0.020
38.5
25.7
12.8
0.015
0.064
0.063
0.025
0.090
25.7
12.8
0.014
0.007
38.5
12.8
0.388
0.007
0.259
25.7
12.8
38.5
25.7
12.8
0.174
2.042
0.020
0.112
0.044
51.3
0.020
12.8
64.2
0.007
0.034
12.8
0.005
25.7
12.8
0.014
0.036
12.8
115.5
89.8
1693.6
0.079
0.081
1.016
9.6
0.030
3.253
0.005
25.7
0.014
12.8
0.014
12.8
25.7
0.020
12.8
0.010
25.7
0.020
12.8
0.010
12.8
0.005
603.0
22.8
0.005
141.1
12.8
0.075
0.037
0.056
0.168
25.7
141.1
0.102
0.149
15.6
1565.3
13.5
27
2.63
0.10
B
0.012
282.3
25.7
1488.3
0.149
0.097
9.7
29
2.71
0.11
OYS 40
N
B
12.8
0.004
36
2.82
0.11
OYS 41
N
B
0.362
25.7
21
2.62
0.09
OYS 42
N
B
77.0
0.023
12.8
25.7
0.229
0.008
OFF 1
N
B
12.8
102.6
51.3
115.5
25.7
25.7
0.004
0.031
0.015
0.114
0.905
0.008
12.8
0.004
51.3
89.8
12.8
25.7
12.8
12.8
0.015
0.027
0.004
0.008
0.004
0.004
12.8
0.004
1.674
0.012
0.008
0.035
51.3
0.015
89.8
0.027
12.8
3.217
461.9
0.004
4.487
25.7
0.008
12.8
0.004
64.2
51.3
2.123
0.010
79
12.8
0.004
38.5
77.0
0.012
0.023
A ppendix 2, Biom onitoring 2001
echicord
echipusi
ophialbi
ophispec
Mollusca
abraalba
25.7
abrapris
aplacoph
12.8
arctisla
bivainde
12.8
chamstri
64.2
corbgibb
cultpell
25.7
cylicyli
12.8
dosilupi
ensiensi
euspniti
kellsubo
mysebide
1141.9
nucuniti
12.8
tellfabu
tellferr
thraphas
thyaflex
12.8
vitrant'
Polychaeta
aphemari
12.8
chaeseto
25.7
diplglau
12.8
eumisang
gonibobr
gonimacu
25.7
gyptcape
harmspec
laniconc
lysilove
12.8
magealle
magemira
12.8
nephcaec
nephhomb
38.5
nephspec
12.8
notolate
opheacum
12.8
ophiflex
owentusi
12.8
parafulg
12.8
pectauri
25.7
pectkore
282.3
pholmlnu
269.4
poecserp
polykinb
12.8
scalinfl
scolarmi
spiobomb
spiofíli
12.8
sthelimi
12.8
syneklat
38.5
12.8
0.008
6.142
64.2
0.002
38.5
25.7
0.003
0.017
25.7
0.001
12.8
0.000
12.8
0.424
12.8
89.8
0.001
0.003
12.8
0.000
25.7
0.112
25.7
0.091
25.7
0.052
12.8
25.7
0.766
0.013
38.5
0.027
25.7
12.8
25.7
0.002
0.002
0.162
38.5
0.027
25.7
12.8
12.8
0.003
0.004
0.009
25.7
0.024
12.8
0.014
51.3
0.056
25.7
0.029
12.8
51.3
0.014
5.181
397.7
0.567
12.8
1.736
0.003
0.007
0.014
0.112
0.004
0.131
0.099
0.001
0.014
0.010
0.005
25.7
564.5
1.237
0.130
25.7
12.8
0.007
0.010
12.8
0.047
102.6
25.7
0.011
0.115
12.8
38.5
0.001
0.033
25.7
0.010
12.8
0.025
25.7
0.010
25.7
0.684
0.005
12.8
25.7
0.005
0.010
0.784
0.005
38.5
38.5
0.352
0.015
0.019
25.7
12.8
166.8
154.0
154.0
0.011
0.001
0.022
0.828
0.003
25.7
192.5
0.587
0.197
12.8
38.5
25.7
0.008
0.071
0.027
25.7
0.212
1.763
25.7
282.3
12.8
25.7
12.8
0.229
0.102
0.039
0.154
0.014
102.6
0.129
0.093
25.7
0.005
0.005
0.010
2.506
0.107
12.8
38.5
0.010
0.005
0.015
12.8
0.014
12.8
0.014
12.8
0.012
38.5
0.024
12.8
0.014
64.2
51.3
0.059
0.047
128.3
0.141
0.545
12.8
38.5
38.5
0.005
0.049
0.015
51.3
0.046
0.015
0.129
25.7
0.027
0.210
12.8
25.7
12.8
0.014
0.163
0.014
80
A ppendix 2, B iom onitoring 2001
Miscellaneous taxa
nemertin
phoronid
166.8
sum
3643.7
diversity
34
nspc
SH-W
2.15
Simp
0.20
station
OFF 2
Crustacea
N
acidobes
batheleg
89.8
bathguil
12.8
calljuve
caprelli
coroinsi
diasbrad
12.8
iphitris
leucinci
megaagil
orchnana
perilong
pseulong
synchapl
syncmacu
12.8
thiascut
urotbrev
urotpose
141.1
Echinodermata
amphchia
echicord
12.8
echipusi
ophitext
ophispec
Mollusca
abraalba
corbgibb
cylicyli
dosilupi
ensiamer
ensiarcu
euspniti
25.7
mysebide
spissubt
25.7
tellfabu
205.3
teilten’
179.6
telltenu
thraphas
12.8
tridtria
Polychaeta
chaeseto
77.0
eteolong
102.6
eumisang
166.8
gonimacu
12.8
magemira 2168.3
nephcaec
nephcin
12.8
nephhomb
0.176
10.1
B
0.027
0.004
0.004
12.8
128.3
2040.0
0.008
0.161
9.5
25.7
64.2
1693.6
32
2.60
0.14
33
2.97
0.07
OFF 3
N
51.3
OFF 4
N
38.5
0.015
12.8
38.5
12.8
0.835
0.004
25.7
0.008
77.0
0.023
12.8
0.004
B
0.004
0.017
0.063
11.5
12.8
0.005
0.628
4.550
0.106
B
0.012
0.004
25.7
12.8
0.008
0.004
25.7
12.8
12.8
0.008
0.004
0.004
38.5
0.012
0.282
0.042
9.6
33
2.93
0.08
B
179.6
51.3
0.054
0.015
12.8
12.8
0.004
0.004
12.8
12.8
0.004
0.004
12.8
12.8
12.8
0.004
0.004
0.004
38.5
0.012
25.7
0.128
333.6
12.8
22.536
0.002
12.8
0.002
12.8
0.002
25.7
0.005
25.7
102.6
0.198
0.214
12.8
0.000
0.060
25.7
85.830
8.526
38.5
0.085
141.1
12.8
1398.5
25.7
0.313
0.029
2.806
0.842
12.8
1.636
12.8
0.001
25.7
25.7
0.048
0.004
38.5
0.040
166.8
577.4
0.183
0.183
64.2
25.7
0.339
0.005
12.8
0.009
0.009
0.142
0.190
0.031
0.740
4.056
OFF 5
N
64.2
179.6
1757.7
OFF 6
N
B
12.8
0.004
12.8
0.004
25.7
25.7
102.6
0.044
0.008
0.031
25.7
0.012
12.8
25.7
12.8
4.918
0.025
0.001
25.7
12.8
0.005
0.000
115.5
0.013
25.7
0.301
0.032
12.8
500.4
0.285
0.076
2.9
24
2.97
0.05
0.042
11.200
25.7
64.2
846.8
25.7
0.022
205.3
0.176
12.8
4.210
0.024
81
38.5
0.029
12.8
0.010
230.9
0.361
12.8
0.171
A ppendix 2, Biom onitoring 2001
nephspec
notolate
ophelima
38.5
owenfusi
pholminu
podahelg
poecserp
192.5
scolarmi
115.5
scolbonn
sigamath
spiobomb
12.8
spiofili
Miscellaneous taxa
nematoda
nemertin
38.5
anthozoa
phoronid
amphlanc
sum
3669.4
diversity
nspc
22
SH-W
1.74
Simp
0.36
station
OFF 7
Crustacea
N
atylfalc
12.8
batheleg
bathgui!
caprelli
89.8
leucinci
megaagil
590.2
melitobtu
orchnana
923.8
perilong
pseulong
syncmacu
12.8
urotbrev
25.7
urotpose
192.5
Echinodermata
astenjbe
12.8
echicord
89.8
echipusi
ophispec
51.3
Mollusca
donavitt
euspniti
12.8
nucuniti
spiselli
spissubt
12.8
tellfabu
102.6
tellferr
38.5
turbpusi
Polychaeta
aricminu
chaeseto
102.6
glycroux
gonimacu
gyptcape
12.8
0.029
25.7
25.7
0.022
0.022
25.7
25.7
0.022
0.022
51.3
0.044
12.8
128.3
12.8
154.0
12.8
102.6
12.8
2296.6
0.071
0.745
1.165
12.8
51.3
12.8
0.029
0.113
0.743
0.024
12.8
102.6
0.029
0.227
1.279
51.3
1.392
25.7
2655.8
104.2
23
1.78
0.32
B
0.004
OFF 8
N
12.8
0.010
25.7
25.7
51.3
0.019
0.019
0.037
0.549
0.110
0.012
12.8
295.1
0.600
0.217
1.104
2.470
0.017
0.040
32.5
77.0
12.8
38.5
0.618
4.053
0.007
1437.0
7.0
32
2.70
0.11
B
115.5
38.5
0.035
0.012
12.8
0.004
64.2
0.007
12.8
0.124
5.6
526.0
16
2.43
0.10
32
2.83
0.09
OFF 9
N
0.025
B
OFF 10
N
B
89.8
25.7
0.027
0.008
115.5
51.3
0.035
0.015
25.7
0.008
12.8
0.004
38.5
12.8
0.012
0.004
718.5
0.216
OFF 11
N
B
269.4
0.081
12.8
0.004
25.7
12.8
0.002
38.5
0.339
51.3
0.008
51.3
0.045
12.8
0.010
12.8
0.066
0.027
0.177
0.277
12.8
12.8
0.004
0.004
12.8
0.004
0.004
0.008
0.058
51.3
641.5
0.015
0.192
38.5
64.2
0.012
0.019
37.741
22.389
38.5
0.156
0.001
128.3
18.750
0.002
0.143
0.637
0.056
0.169
179.6
25.7
12.8
7.305
12.8
115.5
25.7
15.040
0.081
12.8
0.817
0.011
205.3
0.100
12.8
0.076
0.030
0.015
82
0.001
A ppendix 2, Biom onitoring 2001
magemira
128.3
nephcirr
12.8
nephspec
notolate
parafulg
poecserp
scolarmi
89.8
scolbonn
sigamath
spiobomb
25.7
spiofili
102.6
Miscellaneous taxa
nematoda
38.5
nemertin
64.2
phoronid
amphlanc
sum
2732.8
diversity
nspc
22
SH-W
2.26
Simp
0.17
station
OFF 12
Crustacea
N
apheoval
12.8
batheleg
154.0
bathguil
12.8
harpante
megaagil
perilong
syncmacu
thiascut
urotbrev
51.3
urotpose
89.8
Echinodermata
25.7
amphchia
echicord
51.3
ophialbi
ophispec
Mollusca
abraalba
12.8
chamstri
25.7
donavitt
51.3
euspniti
25.7
mysebide
tellfabu
25.7
tellferr
thyaflex
38.5
Polychaeta
aricminu
38.5
chaeseto
gonimacu
12.8
harmlunu
12.8
magemira
51.3
nephassi
nephcaec
nephcirr
102.6
25.7
nephspec
notolate
0.213
0.022
359.2
12.8
0.425
0.046
51.3
51.3
0.213
0.208
12.8
51.3
0.007
0.110
0.007
12.8
12.8
0.010
0.010
166.8
12.8
0.339
0.007
64.2
0.047
859.6
1.5
0.003
0.040
25.7
0.229
38.5
0.040
38.5
12.8
0.316
0.010
62.5
12.8
2052.8
0.040
11.3
564.5
19.5
1642.2
17.2
21
2.13
0.17
B
0.004
0.046
0.004
0.015
0.027
0.001
0.698
13
2.30
0.10
16
2.17
0.16
B
OFF 14
N
B
OFF 15
N
B
38.5
0.012
102.6
0.031
12.8
0.004
12.8
0.004
25.7
12.8
25.7
0.008
0.004
0.008
25.7
0.008
38.5
205.3
0.012
0.062
12.8
128.3
564.5
0.647
0.038
0.169
12.8
38.5
25.7
0.002
37.186
0.038
0.004
51.3
0.005
0.000
12.8
0.000
0.011
0.071
0.098
12.8
3.123
12.8
12.8
0.001
0.000
19
2.03
0.23
OFF 13
N
12.8
0.000
0.200
12.8
0.042
0.169
0.053
0.186
0.010
0.007
0.030
0.015
0.655
1.236
0.755
0.015
12.8
12.8
0.273
0.176
12.8
25.7
12.8
25.7
25.7
397.7
12.8
0.264
192.5
64.2
12.8
12.8
12.8
51.3
0.013
0.052
38.5
12.8
0.023
0.001
12.8
25.7
0.007
0.080
12.8
0.098
89.8
0.488
64.2
51.3
12.8
0.137
0.115
0.029
38.5
0.054
12.8
0.006
64.2
0.041
OFF 16
N
B
89.8
25.7
0.027
0.008
12.8
64.2
0.004
0.019
12.8
0.000
25.7
0.015
38.5
0.227
0.004
0.047
0.015
0.015
0.063
0.176
0.030
25.7
0.008
12.8
0.030
12.8
12.8
12.8
38.5
64.2
0.039
0.005
0.305
0.093
0.022
83
A ppendix 2, Biom onitoring 2001
pholminu
scolarmi
77.0
scolbonn
12.8
spiobomb
25.7
spiofili
sthelimi
Miscellaneous taxa
nemertin
amphlanc
12.8
sum
949.4
diversity
nspc
23
SH-W
2.85
Simp
0.06
station
OFF 17
Crustacea
N
batheleg
bathguil
megaagil
pseulong
12.8
thiascut
urotbrev
urotpose
12.8
Echinodermata
echipusi
Mollusca
donavitt
euspniti
Polychaeta
aricminu
chaeseto
glyclapi
glycspec
laniconc
magemira
12.8
nephcaec
nephcirr
12.8
parafulg
77.0
scolarmi
scolbonn
spiobomb
spiofili
51.3
syllidae
Miscellaneous taxa
nematoda
nemertin
amphlanc
sum
179.6
diversity
nspc
6
SH-W
1.48
Simp
0.23
station
OFF 22
Crustacea
N
atylswam
12.8
batheleg
25.7
bathguil
25.7
0.339
0.015
0.030
0.011
1.8
38.5
384.9
0.135
3.8
12.8
12.8
0.007
0.029
12.8
0.041
51.3
25.7
12.8
0.115
0.058
0.152
12.8
0.041
25.7
0.319
846.8
1.4
22
2.69
0.09
16
2.57
0.06
1052.1
14
1.72
0.31
B
OFF 18
N
B
OFF 19
N
0.004
12.8
12.8
38.5
0.004
0.004
0.012
77.0
38.5
OFF 20
N
12.8
25.7
0.023
51.3
0.012
38.5
0.012
12.8
0.004
0.004
12.8
38.5
0.032
0.3
12.8
25.7
38.5
230.9
0.063
0.117
0.042
0.063
2.4
12.8
0.009
38.5
0.042
12.8
0.015
25.7
12.8
12.8
0.127
0.317
0.024
12.8
0.100
25.7
282.3
0.030
0.7
11
2.18
0.10
9
2.08
0.08
B
0.004
0.008
0.008
B
OFF 23
N
B
25.7
25.7
0.008
0.008
OFF 24
N
B
0.004
0.008
0.015
12.8
0.008
359.2
1.1
OFF 21
N
B
12.8
51.3
38.5
12.8
0.004
0.015
0.012
0.344
25.7
0.003
166.8
0.134
12.8
0.015
141.1
25.7
12.8
0.139
0.025
0.315
25.7
12.8
0.693
0.076
38.5
0.273
77.0
0.357
38.5
179.6
89.8
0.608
0.176
0.088
25.7
25.7
0.271
0.030
25.7
0.029
25.7
25.7
12.8
538.9
0.005
0.034
0.008
1.3
654.3
2.7
12
2.10
0.14
B
0.771
9
1.96
0.13
2.067
0.008
0.224
0.047
38.8
77.0
14
2.28
0.13
OFF 25
N
B
25.7
12.8
0.008
0.004
OFF 26
N
B
A ppendix 2, Biom onitoring 2001
leucinci
megaagil
orchnana
12.8
pseulong
thiascut
urotbrev
urotpose
205.3
Echinodermata
echicord
64.2
Mollusca
donavitt
ensiamer
euspniti
spissoli
Polychaeta
aricminu
eteolong
eumisang
harmljun
lanlconc
nephcaec
nephcirr
51.3
nephhomb
nephspec
ophelima
parafulg
scolaimi
scolbonn
spiobomb
spiofili
12.8
travforb
Miscellaneous taxa
nemertin
12.8
phoronid
77.0
sum
500.4
diversity
nspc
10
SH-W
1.83
Simp
021
station
OFF 27
Crustacea
N
batheleg
bathguil
51.3
calljuve
lemblong
megaagil
orchnana
perilong
procnoho
pseulong
syncmacu
urotbrev
urotpose
89.8
Echinodermata
echicord
echipusi
12.8
ophialbi
ophispec
25.7
25.7
12.8
0.008
0.004
12.8
0.117
12.8
0.004
115.5
0.035
51.3
0.015
25.7
12.8
0.008
0.004
25.7
0.008
12.8
0.000
12.8
25.7
0.633
0.051
12.8
0.010
89.8
0.7
0.004
0.062
12.8
0.004
19.630
12.8
13.669
25.7
25.7
12.195
0.025
12.8
0.196
38.5
25.7
166.8
282.3
0.051
0.034
0.217
12.053
102.6
0.874
12.8
12.8
12.8
0.017
0.066
0.017
205.3
1013.6
0.188
25.9
51.3
12.8
25.7
16
2.17
0.15
B
0.073
17.0
7
1.77
0.13
OFF 28
N
B
64.2
0.019
0.015
25.7
141.1
12.8
12.8
12.8
0.022
0.022
12.8
0.022
25.7
0.423
12.8
12.8
0.022
0.078
12.8
25.7
89.8
12.8
0.107
0.044
0.156
0.022
12.8
0.155
500.4
1.3
18
2.52
0.09
OFF 29
N
12.8
38.5
12.8
38.5
B
0.004
0.012
0.015
0.012
12.8
0.004
12.8
0.300
0.008
0.027
0.001
0.009
0.200
0.247
0.010
0.954
0.034
21.1
12.8
2.766
5
1.55
0.10
OFF 30
N
166.8
89.8
B
0.050
0.027
25.7
0.008
12.8
51.3
25.7
384.9
0.004
0.015
0.008
0.115
OFF 31
N
141.1
38.5
B
0.042
0.012
51.3
0.015
12.8
1.852
6.664
218.1
115.5
0.142
0.022
0.000
85
Appendix 2, B iom onitoring 2001
Mollusca
chamstri
donavitt
ensiamer
12.8
euspniti
38.5
hyalvitr
madcora
tellfabu
tellpygm
thraphas
tridtria
Polychaeta
aricmlnu
chaeseto
eteolong
eumisang
12.8
gonimacu
gyptcape
harmimbr
12.8
harmspec
hetefili
12.8
laniconc
12.8
magemira
nephassi
nephcaec
12.8
nephcirr
25.7
nephhomb
nephspec
opheacum
parafulg
pholminu
poecserp
64.2
scolarmi
scolbonn
spiobomb
25.7
spiofili
sthelimi
travforb
Miscellaneous taxa
nemertin
phoronid
25.7
amphlanc
sum
436.2
diversity
nspc
15.0
SH-W
2.47
Simp
0.08
station
OFF 32
Crustacea
N
atylswam
batheleg
25.7
bathguil
calljuve
calltyrr
iphitris
leucinci
megaagil
12.8
melitobtu
12.8
23.998
0.038
115.5
12.8
0.031
12.8
0.023
25.7
12.8
0.027
0.002
128.3
0.757
12.8
1.589
12.8
0.002
25.7
12.8
0.012
115.5
0.339
25.7
0.010
449.1
3.9
(
64.2
141.1
0.024
0.074
0.001
51.3
0.042
0.012
25.7
0.012
25.7
12.8
0.120
0.005
0.005
0.076
0.383
38.5
0.056
397.7
13.425
12.8
179.6
12.8
0.491
0.449
2.173
38.5
0.461
25.7
0.080
12.8
25.7
0.019
0.037
12.8
0.005
25.7
12.8
25.7
12.8
0.124
0.628
0.012
0.005
38.5
0.056
1360.0
5.3
0.024
0.024
0.342
0.047
12.8
0.303
38.5
12.8
25.7
0.291
0.368
0.030
0.147
25.7
12.8
0.019
12.8
64.2
0.019
0.376
89.8
77.0
12.8
1616.6
0.361
0.041
0.322
15.8
0.240
0.047
12.8
0.023
256.6
8.0
0.027
25.1
11
2.22
0.08
OFF 33
N
12.8
25.7
0.008
12.8
12.8
25.7
25.7
102.6
0.004
12.8
12.8
B
24
2.50
0.12
23
2.61
0.10
B
0.004
0.008
0.004
0.012
3.999
0.008
0.031
0.004
0.004
OFF 34
N
B
179.6
12.8
0.054
0.004
51.3
0.015
86
10
1.90
0.17
OFF 35
N
B
25.7
0.008
12.8
12.8
0.004
0.004
OFF 36
N
A ppendix 2, Biom onitoring 2001
perilong
procparv
pseulong
12.8
syncmacu
thiascut
urotbrev
38.5
urotpose
307.9
Echinodermata
echicord
12.8
ophialbi
ophispec
Mollusca
donavitt
euspniti
12.8
myatrunc
spisspec
tellpygm
Polychaeta
aricminu
77.0
chaeseto
eteolong
51.3
eumisang
exoghebe
glycspec
hetefili
laniconc
nephassi
nephcaec
nephcirr
64.2
nephspec
phylrose
poecserp
scolarmi
25.7
scolbonn
spiobomb
spiofili
syllidae
travforb
12.8
Miscellaneous taxa
nemertin
phoronid
sum
654.3
diversity
nspc
12
SH-W
1.84
Simp
0.24
station
COA 1
Crustacea
N
atylfalc
batheleg
115.5
bathguil
25.7
caprelli
crancran
diasbrad
25.7
orchnana
pontalta
12.8
syncmacu
urotbrev
12.8
urotpose
102.6
0.004
0.012
0.092
6.936
12.8
12.8
12.8
12.8
38.5
38.5
166.8
0.004
0.252
0.004
0.004
0.924
0.012
0.050
51.3
25.7
33.725
0.026
0.009
12.8
12.8
12.8
0.000
0.002
0.002
38.5
38.5
25.7
25.7
0.034
0.034
0.073
0.022
38.5
89.8
0.034
5.419
12.8
25.7
25.7
128.3
12.8
0.120
0.022
0.022
0.293
0.012
141.1
0.125
0.020
0.014
0.418
0.146
12.8
12.8
0.004
0.004
141.1
0.042
25.7
0.001
12.8
25.7
0.001
0.027
51.3
25.7
12.8
12.8
12.8
51.3
0.008
0.004
0.019
0.004
0.015
25.7
0.079
0.117
12.8
0.015
25.7
0.004
12.8
0.003
25.7
0.030
12.8
0.015
12.8
0.003
0.263
0.073
0.625
0.960
25.7
0.007
0.478
0.046
0.015
12.8
0.003
102.6
1.8
64.2
0.158
12.8
12.8
12.8
128.3
25.7
25.7
51.3
12.8
0.058
0.337
0.117
0.029
38.5
38.5
12.8
0.450
8.1
38.5
102.6
1398.5
0.875
0.041
46.9
35
3.17
0.05
B
0.035
0.008
COA 2
N
89.8
12.8
705.7
1.1
15
2.31
0.12
B
COA 3
N
12.8
0.004
0.031
0.031
500.4
2.8
19
2.62
0.08
B
0.004
COA 4
N
12.8
12.8
12.8
25.7
12.8
218.1
744.1
6
1.73
0.07
B
0.027
0.004
0.008
0.004
12.8
0.008
0.004
0.065
0.223
307.9
12.8
0.092
0.004
102.6
346.4
0.031
0.104
87
12.8
51.3
449.1
COA 5
N
B
51.3
12.8
0.015
0.004
12.8
0.004
25.7
0.008
128.3
0.038
0.004
0.681
0.004
0.004
0.015
0.135
A ppendix 2, Biom onitoring 2001
Echinodermata
echicord
Mollusca
donavitt
ensiamer
12.8
euspniti
macobalt
mysebide
spissoli
12.8
spissubt
77.0
tellfabu
346.4
teilten
Polychaeta
capicapi
12.8
chaeseto
25.7
eteolong
12.8
eumisang
hannlunu
hatmspec
laniconc
magemira
1039.2
nephcaec
12.8
nephcirr
25.7
nephhomb
nephspec
ne relong
ophelima
64.2
pectkore
phylgroe
poecserp
38.5
scolarmi
641.5
scolbonn
12.8
spiobomb
spiofili
12.8
Miscellaneous taxa
nemertin
77.0
sum
2720.0
diversity
nspc
22
SH-W
1.98
Simp
0.22
station
COA 6
Crustacea
N
atylswam
batheleg
25.7
bathguil
12.8
caprelli
orchnana
syncmacu
38.5
urotbrev
25.7
urotpose
38.5
Echinodermata
echicord
12.8
ophitext
Mollusca
donavitt
ensiamer
12.8
euspcate
euspniti
38.5
35.245
25.7
2.635
35.228
154.0
0.060
0.011
7.881
64.2
577.4
243.8
12.8
12.8
38.5
12.808
12.8
14.569
12.8
12.8
0.513
0.169
51.3
40.617
25.7
77.0
0.706
0.010
218.1
359.2
7.454
5.849
38.5
0.027
12.8
12.8
12.8
51.3
38.5
0.008
0.008
0.008
3.338
0.027
38.5
115.5
0.190
2.318
3.758
1.479
0.239
0.022
0.042
0.022
2.473
0.113
0.117
12.8
12.8
397.7
320.8
51.3
3.332
28.866
5.613
0.011
12.8
12.8
0.015
0.015
12.8
0.015
12.8
25.7
102.6
25.7
12.8
0.073
0.046
1.316
0.032
2.530
25.7
64.2
0.699
0.178
0.169
0.073
0.610
0.007
25.7
12.8
0.034
19.361
51.3
25.7
0.781
0.037
25.7
0.080
295.1
0.511
12.8
12.8
0.058
0.483
12.8
0.017
25.7
12.8
0.034
0.097
0.107
0.064
2.117
0.022
0.022
282.3
0.051
77.0
25.7
0.095
0.032
38.5
166.8
0.027
0.120
12.8
0.017
0.469
49.4
2553.2
44.6
38.5
2052.8
0.700
58.0
12.8
1834.7
0.886
77.9
12.8
769.8
0.065
21.8
16
2.08
0.17
COA 7
N
25
2.46
0.12
0.008
0.004
COA 8
N
12.8
128.3
51.3
B
0.004
0.038
0.015
0.012
0.008
0.012
12.8
12.8
12.8
205.3
0.004
0.004
0.004
0.062
12.8
B
4.538
64.2
12.8
25.7
B
0.019
0.004
0.008
18
2.21
0.18
23
2.44
0.12
COA 9
N
B
COA 10
N
B
12.8
0.004
12.8
12.8
0.004
0.004
25.7
0.008
243.8
962.3
0.073
0.289
51.3
192.5
0.015
0.058
0.349
25.7
25.7
14.615
2.822
12.8
11.200
102.6
13.708
12.8
0.003
38.5
34.681
25.7
7.293
12.8
0.047
11.872
25.7
88
0.046
A ppendix 2, Biom onitoring 2001
macobalt
192.5
mysebide
12.8
spissubt
218.1
tellfabu
192.5
tellferr
12.8
Polychaeta
capicapi
51.3
chaeseto
12.8
harmlunu
laniconc
magemira
808.3
nephcirr
nephhomb
12.8
nephspec
nerei ong
phylspec
12.8
scolarmi
64.2
spiobomb
89.8
spiofili
410.6
sthelimi
Miscellaneous taxa
nemertin
sum
2258.1
diversity
nspc
20
SH-W
2.11
Simp
0.18
1.471
0.010
10.005
4.972
0.023
25.7
0.042
0.010
12.8
0.437
1129.0
25.7
0.026
0.007
2.142
0.135
25.7
25.7
0.213
0.016
25.7
0.005
25.7
0.596
12.8
0.005
64.2
0.330
0.769
0.010
0.053
0.073
0.332
34.8
1295.8
2.3
7
0.59
0.76
station
COA 11
COA 12
Crustacea
N
B
N
batheleg
25.7
bathguil
12.8
corycass
12.8
urotbrev
25.7
0.008
urotpose
320.8
0.096
12.8
Echinodermata
echicord
12.8
12.808
ophialbi
12.8
0.002
ophitext
12.8
1.558
Mollusca
abraalba
ensiamer
64.2
euspniti
38.5
0.032
mysebide
12.8
0.003
12.8
tellfabu
tellferr
38.5
0.033
Polychaeta
capicapi
chaeseto
eteolong
12.8
0.003
magemira
25.7
nephcaec
12.8
nephcirr
51.3
0.183
38.5
nephhomb
12.8
nephspec
12.8
nerelong
notolate
owenfusi
phylgroe
757.0
22.7
16
2.31
0.13
B
0.008
0.004
19.684
COA 13
N
B
25.7
0.008
63.931
0.003
12.8
12.8
12.8
12.8
12.8
12.8
12.8
0.080
0.010
0.010
0.476
0.010
8.624
12.8
0.046
12.8
6119.9
0.023
291.2
20
1.14
0.51
0.004
0.088
1.790
1.880
0.044
0.044
243.8
0.080
4259.6 261.485
115.5 2.452
77.0
0.040
0.176
1.324
1.961
0.056
12.8
12.8
77.0
89.8
51.3
12.8
0.014
0.378
0.208
0.334
0.738
0.014
38.5
0.039
12.8
0.014
25.7
859.6
0.065
51.2
19
2.56
0.09
COA 14
N
B
12.8
89.8
COA 15
N
25.7
12.8
B
0.008
0.004
0.004
0.027
25.7
205.3
0.008
0.062
115.5
2.725
12.8
0.037
269.4
128.3
2.868
0.160
1950.2
102.6
0.318
0.311
25.7
25.7
51.3
77.0
23.744
0.545
0.011
2.119
64.2
38.5
0.073
0.044
12.8
0.017
192.5
0.249
77.0
128.3
12.8
0.130
5.436
0.017
25.7
12.8
0.256
0.196
12.8
0.015
102.6 2.415
25.7
0.029
12.8
1.243
1077.7 21.376
12.8
89
25.7
115.5
38.5
0.015
A ppendix 2, Biom onitoring 2001
scolarmi
12.8
scolbonn
spiobomb
spiofili
sthelimi
Miscellaneous taxa
nemertin
12.8
oligocha
sum
564.5
diversity
nspc
12
SH-W
1.65
Simp
0.33
0.229
25.7
154.0
0.088
0.723
12.8
0.015
166.8
102.6
12.8
0.188
0.115
0.232
166.8
4477.7
0.188
32.5
0.090
15.1
423.4
13
2.12
0.16
88.3
38.5
0.2
2
0.64
0.33
20
1.89
0.26
90
12.8
25.7
115.5
0.017
0.034
0.149
64.2
12.8
1154.7
1.056
0.017
34.5
21
2.60
0.09
CONTENTS
1. SUMMARY...........................................................................................................................................................
1
2. SAMENVATTING................................................................................................................................................ 3
3. INTRODUCTION................................................................................................................................................. 5
4. MATERIAL AND METHODS............................................................................................................................... 6
4.1. Sampling.................................................................................................................................................6
4.2. Sample treatments............................................................................................................................... 6
4.3. Ashfree Dry weight............................................................................................................................. 7
4.4. Statistics..................................................................................................................................................7
4.5. Sediment analysis................................................................................................................................ 8
5. RESULTS AND DISCUSSION.......................................................................................................................... 9
5.1. Sediment composition........................................................................................................................ 9
5.2. Distribution of the macrobenthic fauna in 2001 .............................................................................. 10
5.2.1. Diversity, density and biomass .........................................................................................10
5.2.2. Temporal variation in density and biomass of individual species............................... 11
7. ACKNOWLEDGEMENTS................................................................................................................................
15
8. REFERENCES..................................................................................................................................................... 16
Tables and Figures.................................................................................................................................................. 18
Appendices.............................................................................................................................................................. 46