This paper not to be cited without prior rcference to the author. INTERNATIONAL COUNCIL FOR TUE EXPLORATION OF TUE SEA C.M. 19861G: 90 Demersal Fish Committee DUTCU GROUNDFISU SURVEY: DATA ON COD AND WUITING (1980-85) by Henk J.L. Heessen. Netherlands Institute for Fishery Investigations P.O. Box 68,1970 AB IJmuiden The Netherlands. INTERNATIONAL COUNCIL FOR TUE EXPLORATION OF THE SEA c.Ml 1986/G:90 \. Demersal Fish Committee DUTCH GROUNDFISH SURVEY: data on eod and whiting (1980-1985) by Henk J. L. Heessen • Netherlands Institute for Fishery Investigations P.O. Box 68, 1970 AB IJmuiden The Netherlands Summary Data on distribution and abundanee of young eod and whiting in the south-eastern North Sea in the period 1980-1985 aie presented. Comparisons are made between the eatehes of the same ycar elasses in eonseeutive years." For eod eorrelation diagrams between indices of year elass strength from thc Duteh Groundfish Survey and the VPA are also given. Introduetion • During 1980-1982, groundfish survcys wcre eonduetcd in all quarters, following the standard proeedurcs used in the International Young Fish Survey in February. Data from these survcys on the distribution and abundanee of young eod and whiting were analysed by Heessen (1983) and a distinet ycarly migration pattern of youngeod in the Gcrman Bight emcrged • Partieularly during the first quarter of the ycarj when the IYFS is eondueted, young eod usually reaeh thc highest degree of eoneentration in the shallow eoastal zonc. Bceause of this aggregation it is diffieult to obtain an aeeuratc estimate of thc abundanec of young eod of age groups 1 and 2 at this time of the ycar. Three months carlier, in Oetober/Novcmbcr, young eod are mueh less eoneentrated and also they are relatively less abundant in shallow water. Besides, such information might be morc effieicntly used for management purposes beeausc it would beeome available earlier in time. Thercfore, a shift in timing of the young fish survcy from Fcbruary to November, might result in more accurate estimates of year elass strength of eod. llowever, the IYFS serves other purposcs then only reeruitment estimates of North Sea eod and for logistie reasons therc is no possibility to transfer rescareh vessel cffort from February to November (Anon., 1985). Still,. beeause of"the problems eneountcred in estimating the strength of year elasses of young eod from the IYFS (see also Anon., 1983 and 1985) we deeided in 1983 to eontinue the surveys in Oetober/Novembcr. Although during thls survey only the south-eastern part of the North Sea 1s f1shed, the ma1n distribution area of young cod 1s covered. - 2 Material and methods The methods used in the Dutch Groundfish Survey are the same as during the IYFS (see IYFS manual, Anon.,1986a). The gear uscd is a high opening bottom trawl (GOV-trawl) and the haul duration is 30 minutes. The area covered in the years 1980-1982 corrcsponded largely with roundfish area 6 and one haul was made per statistical rectangle. From 1983 onwards the area was slightly extended. In the castern and for young cod the most important part two hauls were,made per rectangle and in the western part only one haul per rectangle. Apart from the standard stations fished by thc research vessel "Tridens" (in 1980 und 1982 by the chartered commercial trawler KW 34 "Rose Marie"), in 1984 aud 1985 additional hauls were made in the coastal zone by research vessel "Isis". In Figure 1 the number of hauls per statistical rectangle is given for the different years; the total number of hauls is given in Table 1. • As described in the IYFS manual the whole catch is sampled for length compositions of all species encountered. Otolith sampies are collected for cod, whiting and haddock. The "standard-area" (Figure 2) used for the calculation of DGFS-indices comprises 31 statisticalrectangles, all within roundfish area 6. To correct for rectangles that were not fished in a certain year, the rectangles in the standard-area that were fished during all 6 years (18 out of the 31 rectangles) were used as a reference. The coverage of the standard area in different years is indicated in Table 1. Results and discussion In Figure 3-9 the mean number per rectangle for cod (0-, 1- and 2-group) and whiting (0-, 1-, 2-, and 3-group) is shown for the years 1980-1985. In Table 2 the indices for the same age groups are given for the standard area. Comparisons between catches of the same year classes of cod and whiting in consecutive years are shown in Figure 10 and 11. There is a high correlation between the indices of O-group cod and the index of the same year class one year later (Figure 10). In the data for cod the absence of year class 1984 is very striking, both as 0- and as l-group the catches were very low. The O-group index for year class 1981 is strongly influenced by one very large haul. During the first three years of the survey only one haul was made per statistical rectangle. Over these years in 4 cases over 40% of the total annual catch of a certain age-group was eaught in a single haul. The effect of eliminating these hauls is especially noteworthy in the case of O-group cod in 1981, as indicated in Figure 10. A remarkable feature of the average catch of O-group cod and the catch of the same year class one year later (Figure 10) is that both are of,the same order of magnitude, whereas natural mortality is probably still high during this stage of life. Tbe results of the Multispecies Assessment Working Group indicated a predation mortality of l-group cod of approximately 40% (Anon.; 1986c). An explanation for this feature might be found in the observation that in Oetober/November a high proportion of Q-group eod is present in very shallow waters, where it is hardly possible to fish with a GOV-trawl. Thus, the largest catch of O-group has been made in 1985 in a shallow gully in reetangle 36F8 (see Figure 3). At this station only a 15 minutes haul ean be made. It is likely that in the easternmost part of thc German Bight comparable concentrations may occur. -------- - 3 Comparison of 1- and 2-group eod (Figure 10) revea1s a sharp deerease in numbers which may partly result from a high fishing morta1ity but probab1y also from emigration of 2 year o1d fish out of the investigated area. Hardly any f1sh of age group 3 and older are caught, indicating that there are not many o1der eod 1eft in the area. A1though the time series is rather short the DGFS indices for cod seem to be reasonably well correlated with VPA numbers (Figure 12). • For whiting year class 1983 was remarkably strong in the investigated area. High abundance of this year class was found as 0-, 1- and 2-group. From the camparisan of of the var10us age groups of whiting (Figure 11) it 1s clear that the correlations between 1- and 2-group and between 2- and 3-group are much better than for the 0- and l-group. Taking into account the size distribution of O-group it seems likely that in the fourth quarter not all O-group whiting have recruited to the gear. Comparisons of VPA numbers for whiting and DGFS indices have not been presented because the correlations are very poor. References Anonymus, 1983 - Report of the International Gadoid Survey Working Group, IJmuiden 6-12 July 1983. ICES C.M. 1983/G:62. Anonymus, 1985 - Report of the Working Group on International Young Fish Surveys in the North Sea, Skagerrak and Kattegat. ICES C.M. 1985/H:2. Anonymus, 1986a - Manual for the International Young Fish Surveys in the North Sea, Skagerrak and Kattegat. ICES C.M. 1986/H:2. Anonymus, 1986b - Report of the North Sea Roundfish Working Group. ICES C.M. 1986/Assess:16. Anonymus, 1986c - Report of the ad hoc Multispecies Assessment Working Group. ICES C.M. 1986/Assess:9. Heessen, H.J.L., 1983 - Distribution and abundance of young cod and whiting in the south-eastern North Sea in the period 1980-1982. ICES C.M. 1983/G:30. - 4 - Tab1e 1. surveys. year 1980 1981 1982 1983 1984 1985 Number of hau1s and number of rectangles covered during subsequent number of rectangles covered in standard area number of hau1s in standard area total number of hau1s 28 24 31 27 31 26 28 24 31 SO 83 79 31 34 31 56 92 87 • Tab1e 2. Indices of year c1ass strength of young cod and whiting from the Dutch Groundfish Survey. The indices are the arithmetic mean number per hour traw1ing per rectangle in the standard area. COD • WHITING year class age 0 1978 1979 1980 1981 1982 1983 1984 1985 43.2 176.8 26.9 121.5 1.3 143.6 year c1ass age 0 1977 1978 1979 1980 1981 1982 1983 1984 1985 166.4 1,393.2 165.9 2,649.1 142.8 859.3 age 1 163.8 46.9 83.0 21.8 121.3 3.6 age 1 329.8 205.1 639.8 430.9 1,330.1 783.0 age 2 4.5 11.2 1.6 2.3 1.6 3.1 age 2 61.5 130.5 105.2 223.5 140.7 893.3 age 3 26.8 9.1 52.1 12.4 90.8 31.6 Number of hauls per statistical rectangle in different years. Figure 1 Fa Fl F2 F3 F4 F5 F6 F7 Fa F9 FQ fJ!hV " 42 ~ , 41 , '\ ) , S3 '\ • 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 l 1 1 1~ ~'~ ~ 33 31 , F2 F3 F4 F5 , F6 F8 , , 1 1 " , ) • 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 '\ ';, f{fi 42 ~ 41 ~ ~ 40 1 .... 1•• 1~ 37 '~ r( 1 "0' 35 34 F2 F3 31 . F5 F6 Fa ~ , F8 F9 42 4 " 1 1 1 1 1 1 • "rI '\ 1 1 ;,r}) 1 F1 I. ~ '.\ A 38 M ~~ ~ 33 ~~ 32 ~~ nT . . Fa V .... 1 ~ 1~ 1 "(I i: .{ 31 1981 . , 30 6 F2 F3 F4 F5 10 F6 F7 F8 F9 f!fi 42 "- , 1 , , 2 2 2 2 2 2 2 2'.\ 2 3 2~ · 1 1 2 2 P\ 1 1 2 2 ·) 1 1 3 I. Iw' ) 1 J , · -~ ~ nT . 2 ~ 40 39 38 37 ~~ 36 35 2r}) 1 , .' ~t '<..C'l\J- :'., 41 34 33 ~ ~~ 32 ~.:a 31 1983 , 30 , 6 10 F9 fJ!h , 42 42 41 41 !>.. 56 v • ~ .) 'IJ '" j 1 ~. -~ v-fT . , o ~ ~ 40 1 1 1 1 1 1 2 2 4 4 ., 1 1 2 2 2 4 3 '.\ 38 1 1 2 2 4 5 4 ~ 37 1 1 2 4 1 1 51, 1 1 , i 1 o \Q F7 " 53 1 · ~l 30 , 6 F4 1 51 . o 1 32 1982 1 57 33 ~~ 1 ~ 36 1- nr Fl 1 38 .::; ~ -~ FQ 1 39 '.\ 'r}) 1 j _?f ~l 1 1 o F9 ~"' 1-/ , , -~ , ~I v , , j 10 F7 F7 \ , 1 i 30 , F6 1 32 1980 , F5 1 ·) 5- ~ Fl ; M '0' 5' V 38 , 53 6 Fa , ::\ ~ ~~ ~ o A 1 "( F4 fJ!h , ,1 -~ ~r , --u: . 1 F3 V 'r}) 1 j , 51 ~I 1 1 1 V 1 F2 , v , F1 " 1 6.r(4 o.c .~:~ ~ ,4 -:1 l:>- ~~ 4 , . 6 37 36 35 35 6!f:l 1 39 1984 34 34 33 33 32 32 31 3' 1985 30 10 o' 30 10 Figure 2 Standard area used for the calculation of DGFS indices. ES • E6 1E1 EB ES FO ~ ~= ---_ . __J,_", lrJ fl - f2 -- f3 . f4 F5 f6 F7 Fa .,-9 GO C1 -l-~ --:.~ 52 5' ... 50 ~j- -~!~~--~-- _-~ _ ... L_>--_ I_~ I - f - ._- G2 49 48 ~$ -{ J') / / ~ l't 45 Figure 3 FO t Fl Arithmetic mean number of O-group cod per rectangle. F2 F3 F4 F5 F6 F7 Fa · .5S ... ·IV) 0 2 2 3 16 13 ,,'., .\ 70 32 22~ 0 0 o 2 0 0 1 0 '1 6 0 FO F3 31 F4 F5 F7 0 · 0 ,,;~ 0 0 0 37 88 i 0 " $ ~( 0 0;: Fl 41 40 39 ~2·.\ 119~ 38 .~ l( 68- 35 33 F7 0 0 0 0 0 0 0 1 0 0 0 0 0 6 0 0 0 0 0 13 0 0 0 0 0 . "( 2 s' 42 . 41 .. ·) V 0 '\ j 2 :;f' -~ ~r . o 0 0 0 '0 °A 0 '-C v 1 '~ ~ 9~ 40 39 38 37 ~.~ 36 b- 35 ~rv · FO 34 F1 ~~ 31 1984 · · 30 10 ~v 42 \ 4 \r 6 o ß32 20 760.. 821l ~T'''' 154 76~ 37 1"~32 576 ~ 36 '-'V' 35 ~16rp 40 4 34 76 33 5~ 32 ~el 31 1981 . 30 · F2 F3 F4 F5 F6 F7 2 0 42 3 5 62 10 1 33 259 49 217 · ·I'J) " 0 2 18 2 4 0 3 224 0 4 9 I, 2 o j · $ 4 10 Fa F9 42 /'-C FO Fl ~189 ~ 41rp ~ ;}1, 35~\ 37 \0~ 36 35 33 ~~ 32 31 1983 . F2 F3 F4 F5 30 · F6 F7 10 F8 F9 42 ~ 667 · • 0 j ~ -~ 0 0 :, ~ . o 40 39 2 43 528 0 1 74 411 19'-:,\ 38 0 3 8 37 0 81 8 945~ 54.. r~29 _~2 ~ 1 I, 103 1174 U JE IV' 28f};l )4 ...:: l>- ~!el I' rr1( 41 0 6 0 "'\ 38 34 · ·f'J) 39 569~ ~ i'\ 40 66 ~( · 950.. 41 V ~~ -~ o 38 ~ 32 ~~ F9 39 · 33 6 4 Fa ~' 2 . F7 SI !\ 0 F6 2 o F9 ~ l\ 4 o. 10 Fa 2 ~ · ~( 30 · F6 10 -~ S' 1982 F5 42 120 j i 31 F4 2 "\ .51 ~~ F3 6 4 S3 34 ~'tl F2 0 Po ~ · 12 36 32 . '0 37 2rtJ • 1 FO 0 ... '-C · I" 42 \. ~ 34 F5 .51 0 Il51 ~~ o 4 0 126 0 j t ,n 0 IV " n 0 ·) .5) n 0 1\ .53 F9 ~ n F4 4 0 'W 10 Fa · • 30 · F6 F3 .5' ~v · ·) 32 · .5S 35 ~~ 1980 F2 ... 33 ~~ Fl 8 36 , o F2 · 34 , Fl · 37 64.. ~ro ~.~ M( . t 38 :rtJ o -~ 39 "'\.Clb- Oll 0 -~ 420 40 ~ 100 .56 • 41 8 0 j · SI .5\ 42 ~ v FO · .51 (JJk · • F9 1985 , 6 · 10 Figure 4 · FO Fl F2 F3 F4 F5 F6 F7 Fe F9 Wh · 2 18 4 10 7 0 0 4 14 2 " 8 6 64/l 58 .~ ~T , . 608 960 74 F1 33 . F4 F5 f'J F6 F7 27 9 o '/ 4 6 2 8 93 6 o "..\ 4 6 12 6 2 ce 10 7aff) 4 1?5 ~. ~( FO Fl . F3 0 i'> · f'J) ; 149~ 37 ~"~ 36 35 34 31 F5 F7 10 Fe F6 F7 0 0 0 0 2 4 3 4 0 9 0 0 0 18/, 51 2 J FO F1 F9 " · 8 6 10 20 24 12 0 13 41 0 4 'il 8 ~9 489 2 10 33 13 1\ 8 6 ·) 0 0 22 0 " 2" '\ j .~ -~ ./1T o ~, ~ 40 41 " 56 r-...J 42 67 64 84 7311 217 700 441.. Pc65 <C ~ 39 4 ./ ~ " .\ 38 422~ 37 ~r~ 36 35 22 306 351}) n 23 33 ...: l>- 32 31 . 6 1984 · , 30 10 , 36 " 35 34 32 31 1981 F3 · 6 F4 F5 30 . F6 F7 10 F8 F9 (Ilkv 42 ~ 8 0 160 2 1 15 2 3 4 139 2 0 0 7 3 35 0 0 50 • ·) f'J )" "' .-:i ; 0 4 17 6 10 85 155 81 .. j( -u. Oll FO F1 F2 • 1\ 0 ) 0 · --:; 'nT o 74 38 ~ 37 "~ 36 34 1;ff) 33 32 31 1983 . F3 · ~ ".~ 2 35 0 30 · F4 F5 10 F6 F7 16 "'\ j 39 litJ- · i'J ~.. ~ 40 ...: ~ · ~( . o. · 41 ~~ .~ 2 ~~ 37 33 . F2 o 337~ ..;; ~ · 34 4 61.. j~61_40 .... · S3" ~ 38 51 Wh " 42 ~~ ~T . . · (IlkV 28 2 -~ " F9 \r 62ff) 0 Fa 39 I 30 · F6 0 0 i'> 1982 F4 39 33 6 F5 2 " .~ 32 . F2 40 38 " 51 l' 41 ~ o F4 51 42 I<" .~ · 39 181 1298 4 46~ .. "( 16 F3 o F9 ~, ~ 33 ) .. 53 Fe 40 '\ 8 • 10 4 8 36 11 2' • 30 · 6 " ·) · 31 1980 F2 · 32 · i'> 37 34 Wh :- · 38 Fl " 53 6c/}) 2 F3 39 36 fOJ: " . F2 40 35 56 .. ~ 69~ "(70 \N"~ 51 S3 \~ ~~ o FO 203 1046 41 ~~ .~ · 28 74 0 25 2 j · 52 • 4 0 · V) 51 42 ~ · FO " 51 " re~tangle Arithmetic mean number of l-group cod per 0 2 7 11 8 10 15 10 2 5 0 3 3 O .. rl OA ° 31}) r~ ~ Fe ~, 42 41 40 o ., 39 1 0'..\ 38 0 27 o ... " F9 ~ 3_~ 37 36 , 35 34 0 ,1 33 "" ~ 32 ~~ 31 1985 · 6 30 10 Arithmetie mean number of 2-group eod per reetangle Figure 5 ,. FO F1 F2 F3 F4 F5 F6 F7 'I 0 0 0 10 0 1 0 0 0 35 0 0 0 0 1 25 0 0 0 3 · V) i '\ 0 0 11 2 % i('7 FO F1 F3 1980 · F5 F6 · 1 0 3 · 2 0 ·) · 51 '" -~ Fl 2 o .~ 38 ~ 37 v ~.~ 5~ 31 F2 F3 · F4 F5 30 · F6 F7 · 0 2 0 2 0 0 0 2 5S i" 0 0 0 0 1 2 0 0 1 0 2 10 (ffiV 42 . 4 0 0 0 1 0 '" -~ j -~ 4 0 0 0 211 6 13. "U' o 0 40 ~ ~ 39 0 0 0 1 0 0 4 0 0 40 0'0;.\ 38 ~ 37 r( 1 ~'~ 36 i!yo 35 '" _.f FO 41 40 39 30f}) 0 ß 5 89 6.• ~1 -u 38 T3~ 37 1 36 ~ v 35 34 33 5~ K''8 F1 32 31 1981 F2 F3 F4 F5 30 · · F6 F7 10 F8 F9 0 0 18 0 0 1 0 0 2 1 3 · 0 0 0 'I 9 0 0 · v) 0 0 111 0 Ff) · '" -~ 3 J 2 -~ _5 ~ 32 31 1984 30 IQ 42 V 40 · FO F1 39 15 1'.\ 38 11 4 ~ 37 ~~ 36 ~(o 4 33 ~~ 32 31 1983 . F2 F3 F4 F5 · F6 F7 37 5 10 1 2 5 0 4 3rp ~. J · ---, .~J o 1 11 4 3~ 8 2 :" 13 6 0 42 38 2 ·) fffi 0'0;.\ 2 4 F9 1 2 0 F8 ~'.. ~ 3 0 10 40 0 '\ 30 · · 6 18 '\ 35 34 · V ~ ~ 2., . ?"lr :';} 4 5 41 ~ MT . 2 ~~ ff!A :" 33 4 · 42 ~., ~ · 34 · 211 16 0 55 3rp 6 F9 '.\ ~T . · $1 ../"l( , F8 0 51 o ., 8 0 o F9 V 0 0 -~ · 10 F8 0 ) i 5\ 1982 6 0 0 33 · V v 2 36 35 :" .) ·) 53 56 PI 'I 32 . F7 5S 34 51 i 40 39 5. -(5 -u 41 ~, ~ o F6 51 42 o ., 10 F5 , o F9 ~~ ~T FO 0 l' o 53 Fa :f}) 0 .~ · 0 1 11 0 J 1 0 0 0 ~ 1 2 0 0 0 1\ • 1 · 10 0 5S · 30 · F7 F4 V 32 ff!A :2 38 31 F4 1 39 33 51 · 40 35 ~~ F2 F3 :" 34 . F2 · 36 ~~ n(. F1 f(th 37 01f}) , o SI ~ :~ 2 811 J -~ · 18 • o~'.~ ~ -U-:';} 53 53 41 V · FO 42 \ 5S • ,. F9 ff!A · SI F8 39 ~ '. r( 5 'v-I\J- 36 35 34 33 3 ~~ 32 ~'1:1 31 1985 · · 30 IQ ----------------- Figurc 6 ,. Fa Fl Arithmetic mean number of Q-group whiting per rectangle F2 F3 F4 F5 F6 F7 954 244 268 420 5!J., .~ 257 353 162 278 422 6 72 27 32 49 224 29 79 4 56 114 282 · V) 208 65 lQ1 " 3 j i -~ -~ · . F3 F4 F5 F7 103 41 145 69 '\ o 652 308 51 145 265 8 31- ~( 40 0 ~ ~T . .~ F2 F3 32 31 F5 30 · F6 F7 F9 --$'- -~ ·M( 3 55 463 28;, 39 102 122 35 105 511 13q\ 38 158 320 86 88 62 34 83~ 37 o 499 720 36 3':. r( 14 0 7~ o ) 42 "ir' ~ m .~ 35 o ~'5 33 ~~ 32 ~'<:t 31 1984 · 118~ F3 35 32 31 1981 · F4 F5 F6 F7 F8 219 952 1364 10~?' 37 383~ 40 84 43~ 135 f"U -1'31\51 30 10 i 135rp 35 34 33 c; ~ 32 ~~ 31 1983 · o 30 · 6 F2 132 ~( 36 4<;8 t686 18 _.~ ~ ~ F3 F4 F5 10 F6 F7 F8 375 727 2138 33 8 19 796 675 680 2779 664 1127 388 234 _~261.. 412 13 A . '"\.(""~ F9 fJlk " 42 V 40 41 ,,~ ~ 39 253~ 37 55:.\ ]22 1037 611 2200 -~ 39 483~ 173 , ~, ~ 40 41 675 1105 2775 545 ß224 4768 · '\ j 42 1616 2867 · V) F9 fitk 976 280 · -\ 10 38 · -./~ r · F1 30 . 6 51'~\ 304 FO 37 050 2801 _-:f · 38 33 . F2 39 36 _5~ j i .~ 40 34 240 707 1293 7292 '\ 4 ~~ 55 34 6 357 401 610 · V) 36 1~~ . "" · 51 I o v~ ~ 40 53 107 ... 56 38 o .J 41 . 24 75 , 42 0 5' 47~ 29 51 0 56 544 o 10 F8 -,.513-m ~I 1982 F4 188~ 5~ 33 6 496 3363 · 53 !l, "'\ 38 34 · V 39 35 fltk " ·) 40 42 165~ 835 56 -\ '"ic-~ . F1 36 ~~ o FO · 37 5~ .;f" -~ 53 39'~\ 45 ~ rtJ 10 j \" 41 F9 \~ ~ 57 42 24 710 468 4q,., 18~ 320 r~ ~ F9 Fa 1196 MT . · 10 F8 F7 27 732 j i 30 · F6 F6 2 1113 138 16 575 " 31 158 "\ • · V) _;f 6 F5 132 972 1013 447 1331 16 -\ .~ 225 7 Fl 943 235 .~ 36 1980 F4 6 419 32 . F2 92 ~~ 0 1028 Fa · 37 34 F3 ~ 33 34 ,. 69~ ~ F2 · ~, · • 38 Fl 55 ~~ 5~ i A FO 53 · · V) 39 35 ~6 ~J ~ · 3) MT . F1 "l.r'"~ 40 9 6f};J o Fa .-( 41 ~ \" 201 -\ " 42 ~ v · • F9 fltk · ~, Fa ~ 21~ 38 36 35 33 114 ~~ 32 ~~ 31 1985 · 6 30 10' Figure 7 FO t Fl F2 Arithmetic mean number F3 F4 F5 F6 F7 F8 F9 10e , 72( 44 141 159 454~ 72' 320 643 41 34 211 107 68 186 14 298 6e 8'/1 'I ) , $) ~ j i • " 36 . ~ Fl '~ F3 35 33 32 31 , F4 F5 F6 F7 F8 13 1126 625 685 618 2200 637 465 763 384 39~ 615 865; " _;f -~ d FO Fl 34 ~ 32 F5 30 F6 F7 F8 SI , , 133E f\ " j , -~ -~ ·nT 626 42 444 356 252 129' 51~ 1 P05'~ 36 580 "?';;1(94"f-~ 2}'-U- ~ 35 39rp i $ ~~ 1984 6 F2 '"J r FO 35 33 32 31 , F3 F4 '0 F5' F6 F7 30 10 F8 F9 55 Fl 301 41 40 39 173 29 5 5 312 37 61 75 ~ ~ 4'~\ 38 845 60 184 60 27~ 37 23 18 28 ~ h 144 228 696 866, 554 ""U' 162 581}: ~ 33 ~ 32 ~ 31 1983 , 30 , F2 F3 F4 F5 10 F6 F7 F8 2508 964 629 • fI 40 ~ 63 383 12 11 19 88 27 1 ~ nr , o 2159 1957 670 v" 4o'l 999 1337 218 j 35 34 360 626 " 36 202 63ft) 241 460 F9 ~ ~- 42 , 97 i 30 , , 32 31 36 ~ 6 ·V ~ 2 ~~ 37 151 , 174 rv 33 , , ,I) ). 94 ß~6 23~ ~ ~ 34 s' 37 38 22~ 39 463 2461 1018 558~ I o. ,;., ~ 2778 691 1221 210i 15034 t:> 68 40 38 692 F-J 'il? 671 2349 1221 79'0\ 2280 ) 41q 46l S6 323 2146 621 • 'il? 163 293 42 V 0 39 1981 675 o 41 40 .;; ~ 2601 t Wh S6 89 P. F\ , F9 4 fi1i • '0 42 ~~ Fl I , , SS FO 33 31 F4 n . r l 140 87 t' 35 159 81 12 39 257 F3 · 165 -~ 37 36 F9 '.\ 55 s· 1982 F2 277 38 ~ 6 , 148 664 39 ~ nT 158 40 '~ 82 F8 ~,~ ., 4 W' j i J"'""\r F7 0 303 296/; ~~ o 42 ~ 243 0", " 19~\ :) F6 6' 189 273 420 881 3072 357 382 920" , l~ 1 991 rv ') j i $ F9 v •f\ _91 ,} '0 ~ 43 0 36 451 592 I" S' 30 Wh 45 225 575 , l s' 2q 36 1980 F2 , 37 6 FO , 16~ 34 , F5 ~ 38 ~~ , F4 fi1i SS ~~ nT, F3 , 39 13.3 -~ F2 S) ,; • '~\ 346 214 202 80 30] I( u. '0' Fl 40 8i 18rp o " 41 ~ ,;., 337 FO S6 v I 42 ~ SS · S7 Wh , oE l-group whiting per rectangle 1~9 ?7 1'i," 128 42 41 ~ 39 38 1"\ 837 29~ 571 8 3 , 36 37 '""\r'-'--t li~ 35 rp. 34 33 1$7 ~ 32 ~ , 40 31 . 6 1985 , 3C 10 Figure 8 ,. FO F1 Arithmetic mean number of 2-group whiting per rectungle F2 F3 F4 F5 F6 F1 · 6 lQ 0; 352 84 49 3 5 19 4 4 ? 53 363 139 21 "'11 141. "( 5' .." ~ J' " 8 j ~ i • n 60 n ( Fl 151 133 fU v 7rp F3 f!tA 42 ~., ~ 40 41 39 ._~ 38 4~ ~.~ 31 30 · F1 F8 F9 · 56 3 . 8 f\ ·) 5 • rv ~ j i _.,f" 26 114 22y 41 :'., ~ 40 F1 '\~ 28 41 33 0-:.\ 104 ~ 55 26 22~ 85 6 24 p9 67 38 37 .~ r( 36 'V ~'" 35 13~ .:::~ 32 1982 F5 30 · · 6 F4 F6 F7 . 56 Fe 414 • 42 v 41 f\ 384 .1 ) 68 ~ "') j 2 -~ -~ n7 . o 52 40 :., ~ 11 1'\ 6 1 10 16 14 96 261 63 86 78 2·-:.\ 38 14~ 37 362 156 29 507 267 29 642 56 24l r(393 119 ~ ~ 58 6~ 1 39 63~ 36 35 75 34 $ ~ · FO F1 F9 33 189 115 100 34 33 "" ~ 32 31 1981 F3 F4 F5 10 F6 F1 3 11 23 169 35 61 85 1456 50 671 41 117 1\ 1387 136 224 478 .) 196 161 236 198 · ) 37; ~~ 1984 30 10 42 ~ 41 <, ~ 40 39 3 .\ 38 ~ 37 v 150 39 7~: 11'78-~ ~ /'"V: 36 ~ 35 531}) 33 32 31 1983 F2 F3 6 F4 F5 F6 30 . · · F1 10 F8 F9 f!tA 42 ~ · 246 · 284 538 507 8 1249 1669 447 91 fI ·) 622 134 3651. ~ "') -~ J '.~ nr . o 400 435 41 ~, ~ 40 39 ;.~~ 38 '2, 376 138 63 183 3796 3267 1360 28~ 1(48 9 ~ 2550 1383 995 • 32 31 {!!k ~ ~ 2 ~'<::t F9 ~ -'Ir , Fl F8 241 159 $ FO 30 · · 55 · 21~ 36 24rp 110 " 31 35 28 '\ 38~ ~~ F2 o 38 53 f'.J 4 39 · · 42 V,\ ~' 0 56 33 ~ · F8 170 243 'n-r . 5J 58 121 · F1 118 347 368.. tf 123 6061. ,..--U- 'V 55 ll1rp 6 F6 341 111 o F9 f{tAy V 19 23 j · 10 0 65 1 319 " i 31 ... · f"-I i . F3 • .2 12 30 148 229 251 f\ 0 5J ~~ F2 460 203 34 51 · 39 33 o r ~ 11 102 ~T , . FO 58 333 49 ~ · 17 82 J " 42 V 64 F5 51 {!!k 2 4(, o 10 l' 5' · 36 33 F6 F4 · 5J · F3 ...f!tA 31 34 F5 F2 55 35 6 Fl 56 1980 F4 FO 32 . F2 · 51 ~~ lr"li' '\:t o FO 17 F9 ?,.1 -~ .. 10; "'I4/' 24 F8 "L.C" 37 ~ 36 ~ 35 89fP 34 294 33 --5 ~ 32 ~~ 31 1985 · · 30 10 Figure 9 FO F1 Arithmetic mean number of 3-group whiting per rectangle F2 F3 F4 F5 F6 F7 Fe F9 FO 7· (ifi 42 · " 41 · ,~' ~ 40 56 1 0 2 10 20 13 75 67 0 0 0 26 235 92 1 0 0 20 -\ · v) "\ 1 7 4 47 j · ~ , o · FO 1 3 3 . rp 1~ 37 "\.. 35 34 33 ,.:; ~ F4 F5 F1 1980 F2 F3 · · 6 F4 F5 F6 30 F7 Fe 42 " · 1 23 0 25 1 7 53 21 229 12 ;., 9 26 0-;,\ 9 45 20 7~ 15 31 5' -\ · V) i 71 '\ J 22 .~ ~ F6 F7 F8 F9 ~~ 31 1982 F3 F4 F5 F6 30 · F7 F9 41 32 43 2 7 14 49 176 32 39 32 :..1'-;,\ ~ 244 150 145 17 336 180 98~ PI 76 12 441 31 · V) 8 · 5 i "'\ j -~. ~ :/1 ( . o 51 34 0 52 ;l 13~ ;t'"U" 8(rtJ 12 )00 ~ '\.. 40 39 1 5 21 6 '\ 5 · FO ~ 33 1 ~~ 32 31 1981 · F2 F3 F4 FS F6 F7 6 28 38 35 5~ 31 o '.~ 38 41 o 32 2 6 16 2 ~ 37 18 4 11 46 5? ·(4 2 8 15 6 ·u 14 Il F2 • 1984 30 10 ~ 35 34 ,.;;; ~ 32 ~~ 31 1983 · F3 F4 FS 10 F6 F7 F8 42 "'\ 41 v ,', 14 0 29 52 18 7 o '~ 28 52 34 24 59 3 150 51 2". r(30 26 23 31 32 7"U" 83rp 241 j .,; !v- F9 f{tA 4 .1) 30 · 2 120 ~ ~ 40 39 38 37 36 ti' 35 34 ~1 33 >- 32 ~:l 31 I' ·· 36 33 8 o ~ 39 18 ~3 1\ :/1 ( V" 1 ,., 49f}) · .~ ~ 40 11 10 -~ 41 5 · 32 ~~ 42 2 · n( . · 2 F9 7 5' r....r '\ r F8 , , -~ Fl 10 0 ) FO 30 . · 6 · " 35 34 · d 36 ~~ F1 39 37 O'~ 2!oo .( 7 '-" 40 38 8~ 24 56 · V) ~ 25f}) 16 ----1T . " 54/; lQ 55 36 !v- · 41 "~\ ~ 56 33 · 16 -~ 34 6 5 37 53 ~9 · 0 0 42 22 55 Q .~ l 0 22 0 o 10 r F8 0 1 l' ~":l F2 2q 'I 34 · 2 36 35 6 3 37 ~ · 53 38 561}) · 0 55 .~ }7 56 • 39 32 . Fl 65;1 42 K ~, 3 5 6 o FO -..r 40 33 f-/1( l 18~. 54 41 ~ 3 15 0 o F9 v 0 j i '0 iffi 55 · V) 31 · 9 -\ 32 · " 36 16 • 53 56 • 38 ~~ r. · 55 ~ 57 · F3 (ffi 39 ~ 56 r(81 u 29 1 16 .~. -~ 5' F2 6" v 55 • F1 7· 1985 . · · 6 10 Flgure 10 - Correlatlon between DGFS indices of the same year classes of cod in consecutlve years. DGFS eOD 1 83 • 100o~ so - 81 - - - - - - - - - • 80 • 82 • 8i ..8 l. • I SO 100 • 150 DGFS eOD 0 DGFS eoo 2 79 • 10- • 583 • 81 82 ~t. • 80 • • I I I 50 100 150 OGFS eOD 1 Figure 11 - Correlation between DGFS indices of the same year classes of whiting in consecutive years. DGFS WHI 1 83 • 1000 84 • 81 • 500 - 82 • 80 • • 8l I I 1000 2000 DGFS WHIO OGFS WHI2 1000 - 8~ 500 81 • 82 80 79 • • • • 8{ I r 500 1000 1500 DGFS WHI 1 DGFS WHI3 100- 81 • 79 50- • 82 • • /.00 • 800 DGFS WHI 2 Figure 12 Correlation between DGFS indices for cod and numbers in the - VPA (Anon., 1986b) VPA1r - - - - - - - - - - - - - - - - - - - . eOD o (- _ ~3. ~ 500 - 82 80 • • 250- ~L. 8f 1 • I I 100 SO 150 DGFS eOD 0 VPA eOD2 79 • 30081 82 8.0 • 100- r • 8.3 • 200- T 100 50 • 150 DGFS eOD 1 VPA COD3 .79 6078 • 81 • /.0- • 80 20- 82 ~ • 5 • 10 DGFS eoo 2