#(!04%2 %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE ,INDA!*OYCE53$!&OREST3ERVICE2OCKY-OUNTAIN2ESEARCH3TATION -ARTHA.UNGESSER-!4#/- )NTRODUCTION %ARLIER ANALYSES OF THE SUPPLY AND DEMAND OF TIMBER ASSUMEDTHECONTINUATIONOFHISTORICALCLIMATEANDTHUS DIDNOTEXPLICITLYINCORPORATEFACTORSSUCHASTEMPERATURE ORPRECIPITATIONINTOTHEPROJECTIONSOFTIMBERGROWTH&OR ESTS ARE ADAPTED TO LOCAL CLIMATES AND CHANGES IN THESE CLIMATES ARE LIKELY TO IMPACT FUTURE FOREST GROWTH AND TIMBER OUTPUTS 7ITHIN THE STRATEGIC PLANNING PROCESS OF THE&OREST3ERVICE*OYCEETALTHEANALYSISOFECO SYSTEMPRODUCTIVITYASINmUENCEDBYCLIMATICFACTORSHAS BEEN IDENTIlED AS A CRITICAL QUESTION IN ORDER TO ADDRESS THECHALLENGINGPROBLEMSASSOCIATEDWITHCLIMATECHANGE s 7HAT ARE THE LIKELY EFFECTS OF INCREASING ATMOSPHERIC CARBONDIOXIDEANDPROSPECTIVECLIMATECHANGEONECO SYSTEM PRODUCTIVITY AS MEASURED BY CHANGES IN NET PRIMARYPRODUCTIVITY )NTHELAST20!!SSESSMENT53$!&OREST3ERVICE THECONSEQUENCESOFELEVATEDCARBONDIOXIDEANDCLIMATE CHANGEONNETPRIMARYPRODUCTIVITYOFFORESTSWEREEXAM INED USING CLIMATE MODEL SCENARIOS AND AN ECOLOGICAL MODEL 4%- *OYCE ET AL 4HESE RESULTS WERE THEN USED TO EXAMINE THE IMPACT OF CLIMATE CHANGE ON THE SUPPLYANDDEMANDFORTIMBERPRODUCTSONPRIVATETIM BERLANDS IN THE 5NITED 3TATES SEE lG *OYCE AND "IRDSEYTHISVOLUME)NTHISANALYSISMOSTOFTHEFOREST PRODUCTIVITYCHANGESACROSSTHE5NITED3TATESWEREPOSI TIVELEADINGTOINCREASESINTHETIMBERINVENTORIES7ITH THIS CHANGE IN HARVESTABLE INVENTORIES TIMBER HARVESTS ACROSS THE 5NITED 3TATES SHIFTED AS DEMAND IN VARIOUS REGIONS ADJUSTED TO TAKE ADVANTAGE OF LOWER COST RAW MATERIALS3INCETHESELAST20!ANALYSESNEWEXPERIMEN TALDATAANDMODELINGANALYSESENHANCETHEPICTUREOFTHE POTENTIAL IMPACT OF CARBON DIOXIDE AND CLIMATE CHANGE ONFORESTS)NTHISCHAPTERWEPRESENTTHERESULTSOFSUCH RESEARCH .EWEXPERIMENTSONTHEIMPACTOFCARBONDIOXIDEON VEGETATIONANDMETAANALYSESOFTHEACCUMULATEDRESEARCH HAVEDEMONSTRATEDTHEIMPACTOFCARBONDIOXIDEONPLANT PROCESSES)NTHISCHAPTERWECOMPARETHEIMPACTOFELE VATEDCARBONDIOXIDEONECOSYSTEMPRODUCTIVITYASDETER MINEDFROMRECENTEXPERIMENTALDATAWITHPRODUCTIVITY 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n RESULTSFROMTHELAST20!ANDOTHERECOLOGICALMODELANAL YSES)NADDITIONWEEXPLOREWHETHERTHESEPOTENTIALPRO DUCTIVITYSHIFTSAREWITHINTHERANGEOFPRODUCTIVITYSHIFTS THATTIMBERMANAGEMENTTREATMENTSCOULDINDUCEINMAN AGEDFORESTS7ECOMPARETHERESULTSOFECONOMICRESEARCH ANALYZING THE POTENTIAL TO INCREASE TIMBER GROWTH AS AN INCENTIVETOSTORECARBONWITHTHESEPROJECTEDCHANGESIN PRODUCTIVITYFROMCLIMATECHANGE )NTHELAST20!!SSESSMENTTWOOFTHEMODELINGASSUMP TIONSWERECLIMATEANDVEGETATIONWEREINEQUILIBRIUM ANDTHESPATIALSCALEOFTHEECOLOGICALMODELWASANADE QUATE SCALE FOR NATIONAL LEVEL ANALYSES 2ECENT MODELING STUDIESHAVEEXAMINEDTHESEASSUMPTIONS )N MOST ECOLOGICAL MODELS EVEN NOW VEGETATION IS REPRESENTED AS PRISTINE MATURE VEGETATION RATHER THAN THEACTUALVEGETATIONOFMANYDIFFERENTAGECLASSESSUC CESSIONAL STAGES INTRODUCED SPECIES AND MANAGEMENT HISTORIES &URTHER FOR BOTH THE ECOLOGICAL AND THE ECO NOMIC MODELS A BROAD RANGE OF HABITATS AND SPECIES WEREAGGREGATEDINTOECOSYSTEMSORTIMBERMANAGEMENT TYPES RESPECTIVELY FOR MODELING PURPOSES 4HE VEGETA TION AGGREGATION SCHEMES AND THE NATURE OF THE SPATIAL EXTRAPOLATION DIFFER BETWEEN THE ECOLOGICAL AND THE ECO NOMIC MODELS 4HE IMPLICATIONS OF AN ECOLOGICAL VERSUS ATIMBERMANAGEMENTCLASSIlCATIONSCHEMEAREDISCUSSED INTHISCHAPTER #OMPUTATIONAL PROBLEMS ARISE WHEN INTEGRATING OR LINKING MODELS THAT OPERATE AT DIFFERENT TEMPORAL AND SPATIALSCALES#OMPUTATIONALLIMITATIONSFORCEATRADEOFF BETWEENTHESPATIALEXTENTOFTHEANALYSISOFTENDICTATED BY POLICY CONSIDERATIONS AND THE GRAIN OF THE ANALYSIS #LIMATE MODELS IN ORDER TO COMPUTE GLOBAL DYNAMICS OPERATE AT LARGE SPATIAL SCALES GRID CELLS RANGE FROM DEGREES TO DEGREES RESULTING IN COARSE RESOLUTION OF MOUNTAINOUS REGIONS AND SMALL SCALE CLIMATE FEATURES -ESOSCALE CLIMATE MODELING NOW OFFERS A lNER DEPIC TIONOFCLIMATEFEATURESANDTHEPOSSIBILITYTOINCORPORATE THEEFFECTSOFVEGETATIONANDLANDUSEFEEDBACKSONTOTHE ATMOSPHERIC PROCESSES BUT AT LARGE COMPUTATIONAL COST 'IVENTHISOPPORTUNITYITISIMPORTANTTOUNDERSTANDTHE UTILITYOFGOINGTOAlNERSCALEINTHEECOLOGICALANALYSES BECAUSETHECOMPUTATIONALINTENSITYINCREASESBYANORDER OFMAGNITUDEWHENAKMSCALEISUSEDINSTEADOFTHE TRADITIONAL DEGREE SCALE )N THIS CHAPTER WE REPORT RESEARCHRESULTSQUANTIFYINGTHECLIMATECHANGERESPONSES INECOSYSTEMPRODUCTIVITYATlNERVERSUSCOARSERSPATIAL SCALES *OYCEAND.UNGESSER %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE 0LANTAND%COSYSTEM 0RODUCTIVITY )MPACTOF%LEVATED#ARBON$IOXIDEON 0RODUCTIVITY .ETPRIMARYPRODUCTION.00ISTHEPROCESSBYWHICH THEVEGETATIONINANECOSYSTEMCAPTURESCARBONFROMTHE ATMOSPHERE4HECHANGESINECOSYSTEMPRODUCTIVITYPRO JECTEDFROMECOLOGICALMODELSREmECTCLIMATECHANGESAS WELL AS THE POTENTIAL INmUENCE OF CARBON DIOXIDE FERTIL IZATIONONNETPRIMARYPRODUCTION)NTHELAST20!ANAL YSIS *OYCE .00 OF TEMPERATE FORESTS IN THE 53 INCREASEDFROMPERCENTTOPERCENTDEPENDINGUPON THECLIMATESCENARIOUSEDTABLE-ORERECENTANALYSES USINGAREVISEDVERSIONOFTHE4ERRESTRIAL%COSYSTEM-ODEL 4%-CORROBORATETHESEEARLIERRESULTSWITHSOMEEXCEP TIONS!LARGEMODELINGSTUDYCOMPAREDTHERESPONSESOF SEVERALBIOGEOCHEMISTRYMODELSFORALLECOSYSTEMSWITHIN THECONTERMINOUS5NITED3TATESTOALTEREDCLIMATEANDELE VATEDCARBONDIOXIDE6%-!0MEMBERS!VERAGED ACROSSALLECOSYSTEMS.00RESPONSESINCREASEDFROM TOPERCENTTABLE4HE.00RESPONSESFROM4%- WERE HIGHER THAN THE RESPONSES FOR THE #ENTURY -ODEL 0ARTONETALANDTHE"IOME"'#MODEL2UN NING2UNNINGAND#OUGHLANTABLE4HE LARGEST .00 RESPONSE FOR BOTH 4%- AND #ENTURY AND PERCENT RESPECTIVELY WAS FOR THE 5+-/ SCE NARIOINCONTRASTTOTHERESPONSEOF"IOME"'#PER CENTINCREASE4HELOWEST.00INCREASEFORBOTH4%-AND #ENTURYWASREPORTEDFORTHE/35SCENARIOAGAININCON TRASTTOTHERESPONSEOF"IOME"'#!TTHEGLOBALSCALE THE.00RESPONSESTOCLIMATECHANGEANDELEVATEDCARBON DIOXIDEFORAREVISEDVERSIONOF4%-8IAOETALARE LOWERTHANTHE.00RESPONSESREPORTEDFORTHELAST20! ANALYSIS(OWEVERTHISRESULTREmECTSTHEGLOBALEXTENTOF THESEMEASURES 4HESEPROJECTEDRESPONSESTOALTEREDCLIMATEANDELE VATEDATMOSPHERICCARBONDIOXIDEINCORPORATETHEVARIA TION OF CLIMATE AND ECOSYSTEMS ACROSS THE 5NITED 3TATES OR THE GLOBE )N CONTRAST EXPERIMENTAL STUDIES EXPLORE THE UNDERLYING MECHANISMS FOR A PLANTS RESPONSE TO A CHANGE IN ATMOSPHERIC CARBON DIOXIDE IN A CONTROLLED ENVIRONMENT %LEVATED ATMOSPHERIC CARBON DIOXIDE HAS BEENSHOWNTOINCREASEPHOTOSYNTHESISENHANCERATESOF CARBONASSIMILATIONINCREASESTEMANDROOTBIOMASSAND INTERACTWITHOTHERPLANTNUTRIENTS#EULEMANSAND-OUS SEAU #URTIS AND 7ANG -C'UIRE ET AL B 3AXEETAL)NAREVIEWOFWOODYPLANTS#EULEMANS AND -OUSSEAU REPORTED THE MEAN BIOMASS INCRE MENTFROMELEVATED#/WASPERCENTFORCONIFEROUS TREESANDPERCENTFORDECIDUOUSTREES&ORCONIFEROUS TREESTHERANGEOFRESPONSESWASFROMPERCENTTO PERCENT&ORDECIDUOUSTREESTHERANGEWASFROMnTO PERCENT 3UMMARIZING STUDIES NOT INVOLVING STRESS COMPONENTS 3AXE ET AL REPORTED LARGER AVERAGE LONGTERMBIOMASSINCREMENTDIFFERENCESUNDERELEVATED #/FORCONIFERSOFPERCENTANDSMALLERAVERAGESFOR DECIDUOUSTREESPERCENT 4ABLE.ETPRIMARYPRODUCTIONRESPONSETOCLIMATECHANGEANDELEVATEDCARBONDIOXIDEFORDIFFERENTBIOGEOCHEMICALMODELS 4%-4ERRESTRIAL%COSYSTEM-ODEL#EN#ENTURY-ODEL""'#"IOME"'#ANDATDIFFERENTSPATIALEXTENTS #ONTERMINOUS5NITED3TATES !LLFORESTS 'LOBE !LLECOSYSTEMS %COSYSTEMS #LIMATESCENARIO 4%- 4%- 4%- #EN ""'# 4%- ')33 '&$, '&$,1 /35 '&$,2 5+-/ -)4,/ *OYCE .UNGESSERETAL 6%-!0MEMBERS 8IAOETAL ')33REFERSTOTHESCENARIOFROMTHE'ODDARD)NSTITUTEFOR3PACE3CIENCEMODEL'&$,AND'&$,1REFERTORESULTSFROMTHE'EOPHYSICAL$YNAM ICS,ABMODEL/35REFERSTOACLIMATEMODELDEVELOPEDBY3CHLESINGERANDOTHERSAT/REGON3TATE5NIVERSITY5+-/REFERSTOTHE5NITED+INGDOM -ETEOROLOGY/FlCEMODELAND-)4REFERSTOTHE-ASSACHUSETTS)NSTITUTEOF4ECHNOLOGYMODEL 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE 4HESE RESPONSES FROM EXPERIMENTAL STUDIES ARE NOT WITHOUTCONTROVERSY/FPARTICULARCONCERNISWHETHERTHE RESPONSEISSUSTAINEDOVERTHELIFESPANOFTHETREE-OST OFTHESESTUDIESAREDONEWITHSEEDLINGSORJUVENILETREES .ORBY ET AL CONCLUDED THAT A RESPONSE IN PRO DUCTIVITY WAS THE RESULT OF AN EARLY STIMULUS AND THAT NO FURTHER SUSTAINED RESPONSE WAS OBSERVED 'ORISSEN ET AL SUGGEST THAT AN INITIAL GROWTH STIMULATION MAYBECANCELEDBYLATERPHYSIOLOGICALORMORPHOLOGICAL ADAPTATIONS &OR 9ELLOWPOPULAR ,IRIODENDRON TULIPIFERA , WHOLEPLANT CARBON STORAGE DID NOT INCREASE EVEN THOUGHLEAFLEVELPHOTOSYNTHESISANDLOWERRATESOFFOLIAR RESPIRATIONIN#/ENRICHEDTREESWASOBSERVED.ORBYET AL!NUMBEROFSTUDIESHAVESUGGESTEDTHATTHERE MAYBEARESPONSESPECIlCITYAMONGTREEGENERA#EULE MANSAND-OUSSEAUTOANINCREASEINATMOSPHERIC #/ AS WELL AS WITHIN GENERA #EULEMANS ET AL $E,UCIAETALSUGGESTEDTHATALLOCATIONPATTERNSIN PONDEROSAPINESMAYOFFSETANYINCREASESINPHOTOSYNTHE SISRESULTINGINPOTENTIALDECLINESINPRODUCTIVITYUNDER ALTEREDCLIMATEANDELEVATEDATMOSPHERICCARBONDIOXIDE 2ESPONSESINNATURALSTANDSAREUNCERTAIN #URTIS AND 7ANG CONDUCTED A METAANALYSIS OF OVER REPORTS ON EXPERIMENTS ON THE EFFECT OF ELE VATED CARBON DIOXIDE ON WOODY PLANT MASS FORM AND PHYSIOLOGY4HESESTUDIESSHOWEDSUBSTANTIALVARIATIONIN PLANTRESPONSETOELEVATED#/RANGINGFROMINHIBITIONOF GROWTHTOOVERPERCENTENHANCEMENTRELATIVETOPLANTS GROWN IN AMBIENT CONDITIONS )RRESPECTIVE OF THE GROW ING CONDITIONS THEY FOUND THAT TOTAL BIOMASS INCREASED SIGNIlCANTLYATABOUTTWICEAMBIENTATMOSPHERICCARBON DIOXIDECONCENTRATIONSAVERAGINGAPERCENTINCREASE 3TRESS ALTERED THE RESPONSES ,OW NUTRIENT AVAILABILITY REDUCEDTHE#/RESPONSETOAPERCENTINCREASE,OW LIGHTINCREASEDTHERESPONSETOPERCENT4HEYFOUNDNO SHIFTSINBIOMASSALLOCATIONUNDERELEVATED#/"ELOW GROUND RESPONSES WERE SENSITIVE TO LENGTH OF THE STUDY ANDTHESTRESSESINDUCED 0AN ET AL EXAMINED THE MODELED RESPONSES OFTERRESTRIALECOSYSTEMSTOELEVATEDATMOSPHERICCARBON DIOXIDE 4HE FORESTED ECOSYSTEM .00 RESPONSE RANGED FROMTOPERCENTINCREASESTABLE7HILETHESE BIOGEOCHEMISTRYMODELSASSUMEOPTIMALRESPONSESSIM ILAR TO THOSE OBSERVED EXPERIMENTALLY EG n PER CENT THESE SPATIALLY EXTRAPOLATED RESPONSES TO ELEVATED CARBON DIOXIDE BY ECOSYSTEM ARE SUBSTANTIALLY LOWER 7HEN EXAMINED FOR UNDERLYING DIFFERENCES 0AN ET AL NOTEDTHATTHETHREEMODELSTENDTOAGREEINTHEIR PROJECTED ESTIMATES OF .00RESPONSE TO DOUBLED CARBON DIOXIDE ALONG PRECIPITATION GRADIENTS BUT DIFFER ALONG TEMPERATUREGRADIENTS!LTHOUGHTHEEXPERIMENTALLITERA TUREISEXPANDINGWITH#/IMPACTSTUDIESTHEREISLITTLE INFORMATION ON THE RELATIVE ECOSYSTEMLEVEL RESPONSE OF .00TOELEVATED#/ALONGCLIMATICGRADIENTS0ANETAL 4HESE BIOGEOCHEMISTRY MODELS SERVE AS DIFFERENT 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n *OYCEAND.UNGESSER 4ABLE.ETPRIMARYPRODUCTION.00RESPONSETODOU BLEDATMOSPHERIC#/VERSUSPPMVSIMULATEDBYTHE 6%-!0BIOGEOCHEMISTRYMODELS0ANETAL &ORESTTYPE ")/-%"'# #ENTURY 4%- "OREALCONIFER -ARITIMECONIFER #ONTINENTALCONIFER #OOLTEMPERATEMIXED 7ARMTEMPERATESUBTROPICMIXED 4EMPERATEDECIDUOUS 4EMPERATEMIXEDXEROMOPHIC 4EMPERATECONIFERXEROMOPHIC HYPOTHESESONHOWECOSYSTEMPROCESSESCONTROLTHE.00 RESPONSETOELEVATED#/ 7HEN EXPERIMENTAL STUDIES SINCE REPORTING CHANGESINBIOMASSAREGROUPEDBYFORESTTYPETHESPE CIES RESPONSE IS VARIABLE TABLE 4HIS VARIABILITY IS EXPLAINED IN SOME CASES BY THE DIFFERENT TREATMENTS /PTIMALCONDITIONSSUCHASHIGH.TENDTOIMPROVETHE BIOMASSRESPONSETOELEVATEDCARBONDIOXIDE 7ITHINFORESTTYPES.00RESPONSESTABLEFROMTHE LAST 20!ANALYSIS RANGED FROM A PERCENT DECLINE FOR TEMPERATE DECIDUOUS FOREST PRODUCTIVITY TO AN INCREASE OFPERCENTFORBOREALFORESTPRODUCTIVITY4HEEXPERI MENTALSTUDIESONWOODYSPECIESASSOCIATEDWITHBOREAL FORESTTYPESSHOWEDRESPONSESTOELEVATEDCARBONDIOXIDE OFTOPERCENTINCREASESTABLE2ESULTSFROMTHE MODELINGSTUDIESWHICHINCLUDEPOTENTIALCHANGESINCLI MATEASWELLASCARBONDIOXIDERANGEDFROMINCREASESOF TOPERCENTTABLE&ORTHETEMPERATEDECIDU OUS SPECIES THE EXPERIMENTAL RESULTS INCLUDED A DECLINE OFPERCENTTOANINCREASEOFPERCENT2ESULTSINTHE MODELEDSTUDIESFORTEMPERATEDECIDUOUSFORESTSRANGED FROM A DECLINE OF PERCENT TO AN INCREASE OF PER CENT&ORCONIFERSPECIESTHEEXPERIMENTALRESULTSRANGED FROMNOSIGNIlCANTINCREASESTOANINCREASEOFPERCENT 2ESPONSESFROMTHEMODELEDSTUDIESFORTEMPERATECONIF EROUSFORESTSRANGEDFROMAPERCENTINCREASETOA PERCENTINCREASEIN.004HESEPROJECTEDRESPONSESTOELE VATED#/ANDCLIMATEINTHELAST20!ANALYSESARELOWER THANTHEPOTENTIALRESPONSESINTHEEXPERIMENTALSTUDIES TABLEVERSUSTABLE 0ANETALDETECTEDTHEDIFFERENTECOSYSTEMLEVEL HYPOTHESES THAT THESE BIOGEOCHEMISTRY MODELS REmECT 4HESEAREASOFUNCERTAINTYIFEXAMINEDIDENTIFYOPPORTU NITIESTORElNEOURABILITYTOASSESSTHEIMPACTOFCLIMATE CHANGEONECOSYSTEMS s7HATROLEDOESTHEHYDROLOGICALCYCLEPLAYINCONTROL LING THE #/ RESPONSES OF LEAF AREA AND SOIL MOISTURE ALONGTEMPERATUREANDMOISTUREGRADIENTS *OYCEAND.UNGESSER %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE 4ABLE"IOMASSRESPONSEPERCENTBYWOODYSPECIESUNDERELEVATEDCARBONDIOXIDEINEXPERIMENTALSTUDIES 3PECIES 3IGNIlCANT .ONSIGNIlCANT "OREAL 0ICEAABIES 0ICEAGLAUCA 0ICEAMARIANA 0ICEAMARIANA 0ICEASITCHENSIS 0ICEASITCHENSIS 0ICEASITCHENSIS 0ICEASITCHENSIS !BOVEGROUNDBIOMASSFRESHWT 4OTALBIOMASS 4OTALBIOMASS 4OTALBIOMASS 4OTALBIOMASSIRRIGATIONFERTILIZATION 4OTALBIOMASSFERTILIZATION 4OTALBIOMASSNOIRRIGATIONORFERTILIZATION 4OTALBIOMASSIRRIGATION 4EMPERATECONIFEROUS 0INUSBANKSIANA 0INUSSILVESTRIS 0INUSSILVESTRIS 0INUSTAEDA 0INUSTAEDA 0INUSTAEDA 0INUSTAEDA 0INUSTAEDA 0INUSTAEDA 0INUSTAEDA 0INUSPONDEROSA 0INUSPONDEROSA 0INUSPONDEROSA 0INUSPONDEROSA 0INUSPONDEROSA 0INUSPONDEROSA 0INUSPONDEROSA 0SEUDOTSUGAMENZIESII 0SEUDOTSUGAMENZIESII 2OOTBIOMASS 4OTALBIOMASSHIGH. 4OTALROOTBIOMASS 4OTALROOTBIOMASS 4OTALROOTBIOMASS 4OTALROOTBIOMASS 4OTALBIOMASS 4OTALBIOMASSHIGHTEMP 4OTALBIOMASSHIGH. 4OTALROOTBIOMASS 4OTALROOTBIOMASS 4OTALROOTBIOMASS 4OTALBIOMASS 3HOOTBIOMASS 4OTALBIOMASSLOW. 4OTALBIOMASSLOWTEMP 4OTALBIOMASSLOW. 4OTALBIOMASSAGE 4OTALBIOMASSAGE .3 9AKIMCHUKAND (ODDINOTT )NEICHENETAL )NEICHENETAL 'RIFlNETAL 'RIFlNETAL +INGETAL +INGETAL +INGETAL +INGETAL 4ISSUEETAL $ELUCIAETAL $ELUCIAETAL 'RIFlNETAL 'RIFlNETAL +INGETAL +INGETAL +INGETAL 'ORISSENETAL 'ORISSENETAL 4EMPERATEDECIDUOUS 0RUNUSAVIUM 0RUNUSAVIUM 0RUNUSAVIUM 0RUNUSAVIUM 0RUNUSAVIUM8 PSEUDOCERASUS 1UERCUSROBUR 1UERCUSRUBRA 1UERCUSRUBRA !LNUSRUBRA !LNUSRUBRA 0OPULUSDELTOIDESX NIGRA 0OPULUSDELTOIDESX NIGRA 0OPULUSDELTOIDESX NIGRA 4OTALBIOMASSLOW. 4OTALBIOMASSDECLINE 4OTALBIOMASSHIGH. 4OTALBIOMASSHIGH. ,EAFSHOOTMONTH ,EAFSHOOTMONTH 4OTALBIOMASSLEAFMASS 4OTALBIOMASS 4OTALBIOMASS 4OTALBIOMASS 3TEMVOLUME 4OTALBRANCHBIOMASS 4OTALBIOMASSOFLEAVES ,EAFSHOOT ROOTMONTH 7ILKINSETAL 7ILKINSETAL 7ILKINSETAL 7ILKINSETAL !TKINSONETAL !TKINSONETAL ,INDROTHETAL -IAO (IBBSETAL (IBBSETAL #EULEMANSETAL #EULEMANSETAL #EULEMANSETAL 0ERCENT 2ESPONSE 3OURCE 0OLLEETAL 9AKIMCHUKAND (ODDINOTT ,ORDETAL 9AKIMCHUKAND (ODDINOTT 4OWNEND 4OWNEND 4OWNEND 4OWNEND CONTINUED 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE *OYCEAND.UNGESSER 4ABLECONTINUED 3PECIES 3IGNIlCANT .ONSIGNIlCANT 0ERCENT 2ESPONSE 0OPULUSDELTOIDESX NIGRA 0OPULUSDELTOIDESX NIGRA 0OPULUSTREMULOIDES 0OPULUSTRICHOCARPA XDELTOIDES 0OPULUSTRICHOCARPA XDELTOIDES 0OPULUSTRICHOCARPA XDELTOIDES !CERRUBRUM !CERSACCHARUM !CERSACCHARUM !CERSACCHARUM "ETULAALLEGHANIENSIS "ETULAALLEGHANIENSIS FAMILY' "ETULAALLEGHANIENSIS FAMILY7 "ETULAALLEGHANIENSIS FAMILY9 "ETULALENTA "ETULAPAPYRIFERA "ETULAPAPYRIFERA "ETULAPOPULIFOLIA ,IRIODENDRONTULIPIFERA ,IRIODENDRONTULIPIFERA 4OTALBIOMASS #URTISETAL 4OTALBIOMASS #URTISETAL 4OTALBIOMASSLEAFMASS 3TEMVOLUME 4OTALBRANCHBIOMASS 4OTALBIOMASSOFLEAVES 3TEMMASSROOTMASSLEAFMASS 4OTALBIOMASS 4OTALBIOMASS 3TEMMASSROOTMASSLEAFMASS 4OTALBIOMASSlNECOARSEROOTMASS 3TEMMASSROOTMASSLEAFMASS 3TEMMASSROOTMASSLEAFMASS 4APROOT 4OTALlNECOARSEROOTMASS 4OTALBIOMASS 4OTALBIOMASS 4OTALBIOMASS 4OTALBIOMASS n "RANCHESLEAVESBOLE 3OURCE ,INDROTHETAL #EULEMANSETAL #EULEMANSETAL #EULEMANSETAL "ERNTSONAND"AZZAZ ,INDROTHETAL .OBLEETAL .OBLEETAL 2OCHEFORTAND"AZZAZ 7AYNEAND"AZZAZ 7AYNEAND"AZZAZ 7AYNEAND"AZZAZ 2OCHEFORTAND"AZZAZ "ERNTSONAND"AZZAZ 2OCHEFORTAND"AZZAZ 2OCHEFORTAND"AZZAZ .ORBYETAL .ORBYETAL 4ABLE#OMPARISONOFPROJECTEDCHANGESINFORESTPRODUCTIVITYUNDERCLIMATECHANGEANDELEVATEDCARBONDIOXIDE 4%- &ORESTTYPE "OREAL "OREALFORESTWETLAND 4EMPERATECONIFER 4EMPERATEDECIDUOUS 4EMPERATEMIXED 4EMPERATEBROADLEAVEDEVERGREEN 4EMPERATEFORESTWETLAND !LLFORESTS 4%- '&$, '&$,1 ')33 /35 /35 '&$,1 n n *OYCE .UNGESSERETAL 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n *OYCEAND.UNGESSER s 7HAT ROLE DOES THE NITROGEN CYCLE PLAY IN THE #/ RESPONSESOFLEAFAREAANDLEAFNITROGENCONTENTALONG TEMPERATUREANDMOISTUREGRADIENTS s7HATISTHERELATIVEROLEOFCHANGESINNITROGENREQUIRE MENTS ALLOCATION TISSUE # TO . RATIOS AND RATES OF DECOMPOSITION IN DETERMINING #/ RESPONSES ALONG TEMPERATUREANDMOISTUREGRADIENTS s 7HAT ARE THE RELATIVE CONTRIBUTIONS AND IMPORTANCE OFINTERACTIONSBETWEENTHEHYDROLOGICALANDNUTRIENT CYCLESINCONTROLLING.00RESPONSESTOELEVATED#/ )MPORTANTLY0ANETALCONCLUDETHATFUTURESTUD IES SHOULD MEASURE THE mUXES AND THE POOLS OF CARBON NITROGEN AND WATER ! CLEAR PICTURE OF BOTH mUXES AND POOLSISIMPORTANTINIMPROVINGOURUNDERSTANDINGOFTHE INTERACTIONSAMONGPROCESSESTHATCONTROL#/RESPONSES OF ECOSYSTEMS /UR UNDERSTANDING OF THESE PROCESSES IS THE BASIS FOR THE DEVELOPMENT OF POLICIES ON CARBON SEQUESTRATIONOPTIONSINFORESTS #LIMATE6ERSUS-ANAGEMENT)NmUENCESIN 4IMBER0RODUCTIVITY 4HEPRODUCTIVITYSHIFTSINTHELAST20!CLIMATECHANGE ANALYSISWEREARESPONSETOINCREASEDATMOSPHERICCARBON DIOXIDE AND CHANGES IN TEMPERATURE AND PRECIPITATION 4HE TIME PERIOD WAS YEARS !RE THOSE PRODUCTIVITY SHIFTSSIMILARTOTHEBIOLOGICALPOTENTIALOFCURRENT53FOR ESTS/RARETHOSEPRODUCTIVITYSHIFTSSIMILARTOINCREASES SEEN UNDER ECONOMIC OPPORTUNITIES FOSTERED BY TIMBER MANAGEMENT OVER A SIMILAR TIME FRAME 6ASIEVICH AND !LIGUSEDFORESTINVENTORYDATATOASSESSTHEPOTEN TIAL TO INCREASE TIMBER GROWTH FOR CARBON STORAGE "IO LOGICAL OPPORTUNITIES WERE DElNED AS THE POTENTIAL NET ANNUALGROWTHOFTHEMOSTPRODUCTIVEPLOTSTOPOF MEASURED PLOTS FOR EACH SITE CLASS FOREST MANAGEMENT TYPE AND TREATMENT OPPORTUNITY ON TIMBERLAND SUITABLE FOR TREATMENT 4HIS ESTIMATE REPRESENTS ACTUAL MANAGE MENT BEING APPLIED TO CURRENT STANDS AND WAS THUS DEEMEDACHIEVABLE%CONOMICOPPORTUNITIESWEREDElNED ASINCREASESINGROWTHONTIMBERLANDTHATCOULDBETREATED ANDYIELDORMOREONTHEDIRECTCOSTSOFTREATMENT "ASEDON6ASIEVICHAND!LIGSANALYSISBIOLOGI CALOPPORTUNITIESEXISTTOINCREASETIMBERGROWTHBYABOUT BILLION CUBIC FEET OVER MILLION ACRES AN INCREASE OF PERCENT OVER THE CURRENT NET ANNUAL GROWTH OF BILLION CUBIC FEET 3EVERAL DECADES WOULD BE REQUIRED TO IMPLEMENT THE TREATMENTS TO ATTAIN THESE INCREASES &OR ECONOMIC OPPORTUNITIES 6ASIEVICH AND !LIG ESTI MATEDTHATNETANNUALGROWTHCOULDBEINCREASEDBY BILLIONCUBICFEETAPPROXIMATELYPERCENTOFTHECURRENT NET ANNUAL GROWTH #APITAL INVESTMENT COSTS WOULD BE BILLION 4HESE BIOLOGICAL AND ECONOMIC OPPORTUNI TIESWOULDTAKEDECADESTOIMPLEMENTWITHTHEFULLEFFECT NOTBEINGSEENUNTILNEARTHEENDOFTHESTCENTURY4HUS %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE TIMBERMANAGEMENTCOULDPOTENTIALLYENHANCEFORESTPRO DUCTIVITYTOALARGERDEGREETOPERCENTTHANISCUR RENTLYPROJECTEDFORTHEPRODUCTIVITYRESPONSESTOCHANGES INCARBONDIOXIDEORCLIMATETOPERCENTTABLE 0OTENTIAL6EGETATIONAND#URRENT 6EGETATION$ESCRIPTIONS )N THE LAST &OREST 3ERVICE CLIMATE CHANGE ASSESSMENT .00RESPONSEFROMTHEECOLOGICALMODEL4%-WASUSED TO ADJUST GROWTH IN THE FOREST SECTOR MODEL 4!-- !4,!3.!0!0lG*OYCEAND"IRDSEYTHISVOLUME 4HEVEGETATIONCLASSIlCATIONSYSTEMSOFTHESETWOMODELS DIFFERED#LASSIlCATIONSYSTEMSWITHINTHEFORESTRYSECTOR HAVE FOCUSED ON COMMERCIAL TIMBER WHILE CLASSIlCATION SYSTEMSWITHINBOTANYANDECOLOGYHAVEFOCUSEDONTHE DYNAMICS OF PRISTINE PLANT COMMUNITIES )MBEDDED IN THELAST20!CLIMATECHANGEASSESSMENTISTHECONVERSION OF.00RESPONSESFROMTHEECOSYSTEMCLASSIlCATIONUSED IN 4%- INTO THE FOREST MANAGEMENT TYPES USED IN THE 4!--!4,!3.!0!0MODEL 7ITHIN THE FOREST SECTOR MODEL YIELD TABLES TO PROJECT TIMBERGROWTHAREDERIVEDFROMINVENTORYPLOTDATACOL LECTEDOVERAPERIODOFSEVERALYEARSWITHINEACHOFTHESIX 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TYPES OF !SPENBIRCH AND ARE EAST AND WEST EXPRESSIONS OF THE !SPENBIRCH TYPE)N THE %AST THIS TYPE WAS INDEXED TO THE SERAL STAGES OF .ORTHERN HARDWOODS .ORTHERN HARDWOODSlRFORESTSAND.ORTHERNHARDWOODSSPRUCEFORESTANDIDENTIlEDASTEMPERATEDECIDUOUS6%-!0TYPE)NTHE7ESTTHISTYPEWASALSOCODEDASTEMPERATEDECIDUOUS6%-!0 TYPE!LL&)!WESTERNHARDWOODSTYPEWEREIDENTIlEDASTEMPERATEDECIDUOUS6%-!0TYPE #ALIFORNIAMIXEDEVERGREEN "LACKBELT ,IVEOAKSEAOATS #YPRESSSAVANNA -ANGROVE /AKHICKORYPINE 3OUTHERNMIXED 3ANDPINESCRUB 3UBTROPICALPINEFOREST +UCHLER 7ARMTEMPERATESUBTROPICAL MIXEDFOREST 6%-!0VEGETATION 4ABLECONTINUED %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE *OYCEAND.UNGESSER *OYCEAND.UNGESSER IF SILVICULTURAL MANAGEMENT FAVORED A SERAL SPECIES OR IF CLIMAX VEGETATION WERE NOT DOMINANT WITHIN PARTS OF THEREGIONORIFTHECOMMERCIALSPECIESHAPPENEDNOTTO BETHETREESPECIESNAMEDINTHECLIMAXTYPE&OREXAM PLESOFTWOODMIXAFORESTMANAGEMENTTYPEUSEDINTHE 0ACIlC.ORTHWESTWESTTIMBERSUPPLYREGIONWASAMIX OFTWOFORESTTYPESREDWOODANDLARCH.O0.6TYPEWAS REPRESENTATIVEOFREDWOODWITHINTHE0ACIlC.ORTHWEST WESTREGIONWESTSIDEOF/REGONAND7ASHINGTON4HE NEAREST 0.6 TYPE WAS THE REDWOOD TYPE IN THE 0ACIlC 3OUTHWEST REGION (ERE THE ASSUMPTION WAS MADE THAT THERESPONSETOCLIMATECHANGEWOULDBEMOREAPPROPRI ATELY DESCRIBED BY USING A SIMILAR ECOSYSTEM TYPE BUT OUTSIDE OF THE REGION RATHER THAN A DIFFERENT VEGETATION BUT INSIDE THE REGION )N ANOTHER EXAMPLE IN THE 0ACIlC .ORTHWESTWEST REGION RED ALDER WAS A FOREST MANAGE MENTTYPETHATDIDNOTHAVEACORRESPONDING0.6TYPE !CCORDINGTO%YREREDALDERISASUCCESSIONALTYPE REPLACEDBYTHE0ACIlC$OUGLASlRANDWESTERNHEMLOCK SITKA SPRUCE TYPES %YRE DID ASSIGN RED ALDER TO THEINVENTORYTYPEOFWESTERNHARDWOODSAND'ARRISON ET AL ASSIGNED THE 0.6 TYPE /REGON OAKWOOD WITHTHEINVENTORYTYPEOFWESTERNHARDWOODS(OWEVER THIS0.6TYPEOCCURSSOLELYALONGTHE/REGON#ALIFORNIA BORDER IN THE 0ACIlC .ORTHWESTWEST REGION AND RED ALDER IS COMMON ON BOTTOM LANDS SHELTERED COVES AND ON MOIST SLOPES OF THE #OAST AND #ASCADE RANGES %YRE 4HUSTHE0.6TYPEASSOCIATEDWITHTHECLIMAXVEG ETATION THAT TYPICALLY REPLACES RED ALDER THE CEDAR HEM LOCK$OUGLASlRTYPEWASUSEDTOMODIFYTHEYIELDTABLE FORREDALDER )N THE 0ACIlC .ORTHWESTEAST REGION LODGEPOLE PINE WAS A FOREST MANAGEMENT TYPE USED IN THE FOREST SECTOR MODEL(OWEVERTHE0.6TYPELODGEPOLEPINEDIDNOT DOMINATEANYOFTHEGRIDCELLSWITHINTHISREGION!CCORD INGTO%YRELODGEPOLEPINEWITHINTHISREGIONWAS ASSOCIATED WITH SUBALPINE lR %NGELMANN SPRUCE WHITE SPRUCEAND2OCKY-OUNTAIN$OUGLASlR4HE0.6TYPES WITH THESE ASSOCIATES lRHEMLOCK $OUGLAS lR WESTERN SPRUCElRWEREUSEDTODElNETHERESPONSEOFLODGEPOLE PINEWITHINTHISREGION 4HE ASPENBIRCH TYPE USED AS A FOREST MANAGEMENT TYPEINTHE.ORTHEASTREGIONISCONSIDEREDBY+àCHLERTO BEASERALVEGETATIONREPLACEDBY.ORTHEASTERNSPRUCElR %YRE CONSIDERED ASPENBIRCH TO BE A BOREAL HARD WOOD!SPENWILLBEREPLACEDBYTHE0.6TYPESOFNORTH ERN HARDWOODS OR SPRUCElR TYPES AND SUCCESSION TO THESE TYPES IS MORE RAPID THAN TO PINE %YRE 4HE VOLUME OF ASPENBIRCH WAS MODIlED BY THE 4%- .00 RESPONSEFROMANAGGREGATIONOF.ORTHEASTERNSPRUCElR NORTHERN HARDWOODS AND NORTHERN HARDWOODSSPRUCE 0.6TYPES )NTHE3OUTHERNREGIONPLANTEDPINEANDNATURALPINE WERETWOFORESTMANAGEMENTTYPESTABLE4HEREWERE NO CORRESPONDING 0.6 TYPES FOR THESE FOREST TYPES &OR %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE THELAST20!ANALYSISTHEOAKHICKORYPINEANDTHESOUTH ERN MIXED FOREST 0.6 TYPES WERE USED TO ASSESS THE IMPACT OF CLIMATE CHANGE ON TIMBER VOLUME OF THE TWO FORESTMANAGEMENTTYPES #LEARLYTHEREARELIMITATIONSWITHASSIGNINGFORESTMAN AGEMENTTYPESTOPOTENTIALNATURALVEGETATIONTYPES%CO LOGICALMODELSREPRESENTTHEMOSTCURRENTUNDERSTANDING OFHOWECOLOGICALPROCESSESOPERATEATTHEECOSYSTEMSCALE 4HE POTENTIAL DISTRIBUTION OF THESE ECOSYSTEMS IMPLIES A SIMILARITY OF ECOSYSTEM FUNCTION WITHIN THE RANGE OF EACH ECOSYSTEM TYPE AS WELL AS A GEOGRAPHIC PRESENCE UNALTEREDBYLANDUSE&ORESTMANAGEMENTMODELSHAVE FOCUSEDONTHEYIELDOFWOODFROMFORESTLAND4HEAGGRE GATIONOFINVENTORYPLOTSINTOAFORESTMANAGEMENTTYPE IMPLIES A SIMILARITY IN TIMBER PRODUCTION WITH THE GEO GRAPHICRANGEOFTHATFORESTMANAGEMENTTYPE&ORBOTH OFTHESECLASSIlCATIONSYSTEMSTHEECOSYSTEMDYNAMICSOR TIMBER PRODUCTION WITHIN A TYPE MAY BE QUITE VARIABLE #URRENT LAND USES HAVE ALTERED POTENTIAL DISTRIBUTIONS )NVENTORYDATAISMORELIKELYTOREPRESENTTHECURRENTDIS TRIBUTIONOFFORESTS 5SE OF VEGETATION TYPES PRESENTS PROBLEMS BY POTEN TIALLYIGNORINGDIFFERENCESBETWEENSPECIESANDNEWASSOCI ATIONSANDHOWBOTHMAYAFFECTECOSYSTEMSUNDERCLIMATE CHANGE 4HE ASSUMPTION THAT SPECIES ASSOCIATIONS WILL REMAINCONSTANTISACONSEQUENCEOFLUMPINGSPECIESSPE CIlCINFORMATIONINTOAhTYPEv+IRSCHBAUMAND&ISCHLIN 3PECIESASSOCIATIONSHAVEBEENVERYDIFFERENTINTHE PASTUNDERDIFFERENTCLIMATES$AVISREPORTSPALEO ECOLOGICAL EVIDENCE OF COMMUNITY TYPES NO LONGER PRES ENT SUCH AS THE SPRUCEOAK WOODLAND ASSOCIATION 4HE APPROACH OF USING FUNCTIONAL TYPES (ENDERSON3ELLERS AND-C'UFlE7OODWARDETALINLIEUOFSPE CIESINCLIMATECHANGEMODELINGISATTRACTIVEBECAUSETHIS APPROACH REDUCES THE COMPUTATIONAL COMPLEXITY ASSOCI ATEDWITHPROJECTINGEACHPLANTSPECIES(OWEVERTHEUSE OF FUNCTIONAL TYPES OFTEN ASSUMES THAT THESE GROUPINGS OF SPECIES WILL REMAIN TOGETHER AND RESPOND TO CLIMATE CHANGE AS A UNIT 4HE USE OF FUNCTIONAL TYPES RAISES THE ISSUEOFWHETHERhFUNCTIONALTYPESvPRESERVESPECIESDIF FERENCES3OLBRIG4HEVALUEOFTHESEFUNCTIONALTYPES IS THAT THEY GROUP SIMILAR PHYSIOLOGICAL AND ECOLOGICAL ROLES 3OLBRIG 6INTON AND "URKE BUT SPECIES BEHAVIORALDIFFERENCESMAYBEOVERLOOKED&OREXAMPLEIS THERATEOFREACHINGEQUILIBRIUMWITHCLIMATETHESAMEFOR ALLSPECIESWITHINAFUNCTIONALGROUP 3IMILARLY THE AGGREGATION OF DIVERSE TREE SPECIES INTO FORESTMANAGEMENTTYPESPRESENTSPROBLEMSBYIGNORING POTENTIALDIFFERENCESBETWEENCOMMERCIALTIMBERSPECIES ANDHOWCLIMATECHANGEMIGHTALTERWOODlBERPRODUC TIONFOREXAMPLEHOWVOLUMESMIGHTSHIFTWHENWOOD DEVELOPSANDTHEQUALITYOFWOODUNDERALTEREDCLIMATE %XPERIMENTALRESULTSSUGGESTTHATTHERESPONSESTOCLIMATE CHANGEMIGHTBEGENERAIFNOTSPECIESSPECIlCSEEEARLIER DISCUSSION 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE 7ITHIN THE LAST 20! ANALYSIS THE IMPACT OF CLIMATE CHANGEON.00WASASSESSEDUSINGTHEHISTORICALRANGEOF TEMPERATEFORESTSlGNOTTHECURRENTDISTRIBUTIONAS AFFECTEDBYLANDUSE4HUSTHEECOLOGICALVARIABILITYANA LYZEDUNDERCLIMATECHANGEREPRESENTEDAGREATERECOLOGI CALAMPLITUDETHANFORESTSCURRENTLYREPRESENT4HESPATIAL DISTRIBUTION OF EXISTING FORESTS USING THE FOREST MANAGE MENT TYPE CLASSIlCATION HAS BEEN RECENTLY MAPPED 7E USEDTHISSPATIALDISTRIBUTIONTODEVELOPAMAPOFVEGETA TION CLASSIlED BY THE SAME SYSTEM USED BY THE 6%-!0 MEMBERS4HEFORESTMANAGEMENTTYPEINFORMATIONWAS AVAILABLE AT THE KM SCALE 7E IDENTIlED A +àCHLER 0.6 TYPE FOR EACH GRID CELL ON THE FOREST MANAGEMENT MAPBASEDON'ARRISONETALTABLE/NCETHE +àCHLERTYPEWASASSOCIATEDWITHEACHGRIDCELLWETHEN USEDTHECLASSIlCATIONGIVENIN6%-!0MEMBERS TOLINKTHECELLTOA66%'TYPETABLE4HEMAPWAS THEN RESAMPLED TO THE KM SCALE lG 4HE lNER SCALEOFTHISMAPALLOWSSMALLERISOLATEDPATCHESOFFOREST TOREMAINONTHEMAPPARTICULARLYINTHE3OUTHWESTAND INTHE'REAT0LAINS(OWEVERTHISDISTRIBUTIONCONTRASTS WITH THE DISTRIBUTION USED IN THE LAST 20!ANALYSIS lG INTHEDRASTICALLYREDUCEDAREAOFFORESTSINTHEEAST ERN PART OF THE 5NITED 3TATES AND IN THE PATCHINESS OF THE FORESTS ACROSS THE 5NITED 3TATES 4HE AREA OF FORESTS IN THE -IDWEST REGION AND THE -ISSISSIPPI 2IVER VALLEY DECLINESWHENTHELANDUSEINAGRICULTUREISREMOVED4HE HOMOGENEITYOFVEGETATIONTYPESISLESSENEDINlGURE PARTICULARLY FOR THE .EW %NGLAND STATES WHERE BOREAL TEMPERATE CONIFEROUS AND COOL TEMPERATE MIXED FOREST TYPESINTERMINGLEINCONTRASTTOTHEUNIFORMITYINlGURE 4HIS REEXAMINATION OF FOREST TYPES lG COULD BEUSEDASTHEBASISFORANANALYSISOFTHEIMPACTOFCLI MATECHANGEONFORESTPRODUCTIVITY)TWOULDLIKELYREmECT THE POTENTIAL SHIFTS IN FOREST PRODUCTIVITY MORE CLOSELY BECAUSECLIMATESHIFTSINREGIONSOFEXISTINGFORESTSWOULD BEUSEDASCLIMATEINPUTTOAMODELSUCHAS4%- 0ROJECTING%COSYSTEM 0RODUCTIVITYAT$IFFERENT 3PATIAL3CALES #LIMATETOPOGRAPHYVEGETATIONANDSOILSINPUTDATA FORECOLOGICALMODELSUSEDINLARGESCALEINTEGRATEDASSESS MENTSARETYPICALLYGRIDDEDATTHEDEGREELONGITUDEBY DEGREELATITUDESCALE#RAMERETAL(EIMANNET AL+ICKLIGHTERETAL-ELILLOETAL6%-!0 MEMBERS 5SE OF GRIDDED INPUT DATA IMPLICITLY ASSUMESTHATTHEMEANORDOMINANTSURFACEFEATURESREP RESENTTHEENTIREGRIDCELL)NHERENTINTHISASSUMPTIONIS 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n *OYCEAND.UNGESSER THEUNCERTAINTYWITHWHICHTHISGRIDDEDVALUEREPRESENTS THE HETEROGENEITY OF THE ACTUAL COARSE GRID CELL FEATURES AND THE REPRESENTATIVENESS OF THIS GRIDDED VALUE WHEN USED IN THE ECOLOGICAL MODELS TO DESCRIBE THE BIOLOGICAL PROCESSESOPERATINGWITHINTHEGRIDCELL7HILEOPPORTU NITIESEXISTTOMOVETHEANALYSISOFTHEIMPACTOFCLIMATE CHANGE ON FORESTS TO A lNER GRID SCALE THIS REDUCTION IN GRIDSIZEWOULDINCREASETHEPROCESSINGTIMEBYANORDER OFMAGNITUDE(ENCEITISIMPORTANTTOASSESSTHEUNCER TAINTYTHATAGGREGATIONOFCLIMATEDATACONTRIBUTESTOTHE ESTIMATION OF FOREST PRODUCTIVITY UNDER CLIMATE CHANGE .UNGESSER ET AL USED THE 4ERRESTRIAL %COSYSTEM -ODEL4%-VERSION-C'UIREETALATOEVALUATE THEUTILITYOFMOVINGTHECLIMATECHANGEIMPACTANALYSIS FROMTHEDEGREESCALEUSEDINTHELAST20!ANALYSISTO THEKMSCALE 4HEEFFECTOFAGGREGATIONONTHEESTIMATIONOFPRODUCTIV ITYHASBEENSTUDIED.ETPRIMARYPRODUCTION.00ESTI MATESDIFFEREDBYPERCENTWHENCOARSEGRAINVERSUSlNE GRAINSOILSDATAWEREUSEDASINPUTDATAFORTHE0N%TMODEL ,ATHROPETAL0IERCEAND2UNNINGOBTAINED OVERESTIMATESOFUPTOPERCENTIN.00FROMTHE&/2%34 "'#MODELWHENSUBGRIDVARIATIONSINCLIMATETOPOGRA PHYSOILSANDVEGETATIONWEREAVERAGEDACROSSASERIESOF GRAINSIZESFROMKMTODEGREE-OSTOFTHISERRORWAS PRODUCEDBYAVERAGETEMPERATUREWHILEAVERAGETOPOGRA PHYSOILSANDVEGETATIONTYPESALSOCONTRIBUTED .UNGESSER ET AL EXAMINED THE IMPACT OF TWO DIFFERENTSPATIALRESOLUTIONSONTHESIMULATEDFORESTECO SYSTEMRESPONSESFORABASELINECLIMATEANDTWOCLIMATE CHANGESCENARIOS4HE4%-MODELUSESSPATIALLYRESOLVED INFORMATION ON CLIMATE MONTHLY PRECIPITATION MONTHLY MEANAIRTEMPERATUREANDCLOUDINESSSOILTEXTUREPER CENTSANDSILTANDCLAYVEGETATIONTYPEANDELEVATION 4HElNERESOLUTIONGRIDCELLSWEREKMINSIZEAND OFTHESECELLSWERENESTEDWITHINACOARSERESOLUTIONGRID CELLOFAPPROXIMATELYAHALFDEGREEINSIZEKM4HE KMsKMRASTERDATAFORCLIMATEMONTHLYPRECIPITA TIONMONTHLYMEANAIRTEMPERATUREWEREOBTAINEDFROM .EILSON PERSONAL COMMUNICATION AS DESCRIBED IN $ALY ETAL-ARKSAND.EILSON&ORTHE KMsKMINPUTDATATHElNESCALEINPUTDATASETSFOR CONTINUOUS VARIABLES WERE AVERAGED TO THE KM s KMRESOLUTION!VERAGINGTOTHEKMSCALERESULTSINTHE SMOOTHING OUT OF PRECIPITATION AND TEMPERATURE VALUES ALONGTHEGRADIENTSOFCHANGEANDTHELOSSOFlNERDETAIL INSOMEAREASlGAND!REASOFlNESCALEPATCHI NESS OF PRECIPITATION IN THE WESTERN MOUNTAINS AND THE SOUTHERN COASTAL PLAINS ARE SMOOTHED OUT AT THE KM GRID SCALE lG !NNUAL AVERAGE TEMPERATURE VALUES AREINmUENCEDINAREASWHERETHEREISSUBSTANTIALTEMPER ATUREVARIABILITYSUCHASAROUNDTHEMOUNTAINOUSAREASIN THE7ESTlG 4HEHISTORICALRANGEOFTEMPERATEFORESTSWASTHESPA TIAL EXTENT OF THE .UNGESSER STUDY +àCHLER VEGETATION Ecosystem Productivity and the Impact of Climate Change Figure 3.1—Forest type distribution used in the last RPA analysis (Joyce et al. 1995). Joyce and Nungesser 56 USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. Figure 3.2—VEMAP vegetation classification associated with the current distribution of forests based on USDA Forest Service inventory plots. Ecosystem Productivity and the Impact of Climate Change USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. Joyce and Nungesser 57 Joyce and Nungesser Ecosystem Productivity and the Impact of Climate Change Figure 3.3—Baseline annual precipitation at 50 km scale and 10 km scales. 58 USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. Ecosystem Productivity and the Impact of Climate Change Joyce and Nungesser Figure 3.4—Baseline annual average temperature at 50 km and 10 km scales. USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. 59 *OYCEAND.UNGESSER %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE WASDIGITIZEDFROMTHEMAP+àCHLERATKM RESOLUTION 3TEVE (ODGE PERSONAL COMMUNICATION AND WASRECLASSIlEDTO4%-VEGETATIONTYPES&ORTHEKM sKMINPUTDATATHElNESCALEINPUTDATASETFORVEG ETATIONWEREAGGREGATEDTOTHEKMsKMRESOLUTION THROUGHTHEUSEOFTHEMAJORITYRULEPROPORTIONALAGGRE GATION METHOD OF #OSTANZA AND -AXWELL 7HILE THE COARSE RESOLUTION GRIDS INCLUDE SIX VEGETATION TYPES lGASEVENTHTYPETEMPERATEEVERGREENBROADLEAF APPEAREDINONLYTHREEGRIDCELLSATTHElNERESOLUTION!LL FORESTTYPESTHATCOMPRISEDGREATERTHANPERCENTOFTHE TOTALAREAWITHINTHEKMGRIDSRETAINEDATLEASTPER CENTOFTHEIRTOTALAREAWHENAGGREGATEDTOTHEKMGRID SIZE4HERARERFORESTTYPESBOREALWETBOREALTEMPERATE FOREST WETLAND AND TEMPERATE BROADLEAF EVERGREEN LOST FROM TO PERCENT OF THEIR AREA UNDER THE PROPOR TIONAL AGGREGATION RULE 2ARE TYPES DISPERSED ACROSS THE LANDSCAPEWEREMOSTLIKELYTOBELOSTINTHEAGGREGATION &OREXAMPLETHEBOREALGRIDCELLSIN.EW%NGLANDDISAP PEAR AT THE KM SCALE lG 3IMILARLY SOME OF THE FORESTEDBOREALWETLANDFORESTSINTHENORTHERN-IDWEST ARELOSTATTHEKMSCALE !NAGGREGATIONERRORWASCOMPUTEDASTHEDIFFERENCE OFTHEKMESTIMATEOF.00ANDTHEKMESTIMATEOF .00DIVIDEDBYTHEKMESTIMATEOF.004HISAGGREGA TION ERROR WAS COMPUTED FOR THE BASELINE RUNS BASELINE CLIMATEAND#/LEVELSOFMEQL 4ABLE.ETPRIMARYPRODUCTION.00FORBASELINECLIMATE 6ALUESARENETPRIMARYPRODUCTIONING#MREPORTEDVALUESARE MEANSANDSTANDARDDEVIATIONSAREINPARENTHESESFOLLOWEDBY RANGES4HEhNvISTHENUMBEROFCOARSERESOLUTIONGRIDCELLSIN EACHFORESTTYPE &OREST4YPEN 2ESOLUTION .00 "OREAL &INE #OARSE n n &ORESTEDBOREAL WETLAND &INE #OARSE n n -IXEDTEMPERATE &INE #OARSE n n #ONIFERS &INE #OARSE n n $ECIDUOUS &INE #OARSE n n 4EMPERATEFORESTED WETLAND &INE #OARSE n n !,,&/2%343 &INE #OARSE n n INBOREALANDFORESTEDBOREALWETLANDFORESTSWITHPOSI TIVEVALUESOFANDPERCENTRESPECTIVELY%STIMATES EB = 100 ∗ NPP10 − NPP50 / NPP10 OF .00 FOR BOREAL FOREST AT THE KM GRID SCALE WERE LESSTHANTHEKMESTIMATEBYAPPROXIMATELYG#M d = 100 ∗ NPP ( WHERE %" IS THE RELATIVE AGGREGATION ERROR AND GQ50 IS TABLE GQ50 − NPP50 ) / NPP50 THE AVERAGE OF THE MEAN ANNUAL .00IN G#M YR ESTI 4HE CLIMATE CHANGE SCENARIOS TEMPERATURE PRECIPI MATESFORTHElNEGRIDCELLSAND.00ISTHEESTIMATE TATION AT THE KM SCALE WERE BASED ON TWO '#-S OFTHE.00ATTHECOARSEGRIDSCALE USED IN THE LAST 20! ANALYSIS THE 'EOPHYSICAL &LUID 4HE .00RESULTS FOR THE KM REmECT lNER SCALE PAT $YNAMICS ,ABORATORY 1mUX '&$,1 -ANABE AND TERNSTHANTHEKMRESULTSlGBUTTHESEPATTERNS 7ETHERALD AND /REGON 3TATE 5NIVERSITY /35 ARENOTSUFlCIENTTOGENERATELARGEAGGREGATIONERRORSAT MODELS3CHLESINGERAND:HAO5SINGTHESAMEPRO THENATIONALFORESTTYPEORGRIDCELLSCALE4HE.00ESTI TOCOLASFORBASELINECLIMATElNERESOLUTIONCLIMATEINPUT MATEFORALLFORESTSATTHENATIONALEXTENTDIFFEREDBYLESS DATAWEREAVERAGEDWITHINACOARSEGRIDTOSERVEASCOARSE THAN G#M BETWEEN THE lNE AND COARSE RESOLUTION RESOLUTION INPUTS TO 4%- 6ALUES FOR FOREST TYPE ELEVA TIONANDSOILSREMAINUNCHANGEDFROMTHEBASELINESIM SCALESVERSUSG#MTABLE!GGREGATION ULATION 4HE CLIMATE CHANGE SCENARIOS INCLUDED A #/ ERRORBASEDONTHESEGRIDCELLSISVERYSMALLANDNEGA TIVEn%STIMATESOF.00ATTHEKMGRIDSCALEDIF CONCENTRATIONOFPPMV!GGREGATIONERRORIN.00FOR FEREDFROMTHECORRESPONDINGAVERAGEFORTHEKMGRID THE TWO CLIMATE CHANGE SCENARIOS WAS COMPUTED IN THE CELLS BY LESS THAN PERCENT ACROSS MOST OF THE HISTORIC SAMEMANNERASTHEBASELINEAGGREGATIONERROR RANGEOFTEMPERATEFORESTS4HESMALLESTAGGREGATIONERROR 'RIDLEVELRESPONSEOFNETPRIMARYPRODUCTIVITYTOTHE WASFOUNDGENERALLYTHROUGHOUTTHE%ASTAND3OUTHEAST CLIMATECHANGESCENARIOSWERECALCULATEDAS ASWELLASINTHEWESTERNMOUNTAINS2ARELYWEREAGGRE d OS10 = 100 ∗ NPPOS10 − NPP10 / NPP10 , GATION ERROR DIFFERENCES GREATER THAN PERCENT AT THE INDIVIDUALGRIDCELLLEVEL4HESELARGERAGGREGATIONERRORS d OS50 = 100 ∗ ( NPPOS50 − NPP50 ) / NPP50 , OCCURREDAROUNDTHE'REAT,AKESINNORTHERN.EW%NG LAND AND IN THE 2OCKY -OUNTAINS "Y FOREST TYPE THE d GQ10 = 100 ∗ NPPGQ10 − NPP10 / NPP10 , and AGGREGATION ERRORS WERE STILL SMALL LESS THAN PERCENT %STIMATESOF.00DIFFEREDBYLESSTHANG#MINMOST d GQ50 = 100 ∗ ( NPPGQ50 − NPP50 ) / NPP50 CASES TABLE 4HE LARGEST AGGREGATION ERROR OCCURRED ( ) ( ) ( ) 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n Ecosystem Productivity and the Impact of Climate Change Joyce and Nungesser Figure 3.5—Vegetation types mapped at 50 km and 10 km scales. USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. 61 Joyce and Nungesser Ecosystem Productivity and the Impact of Climate Change Figure 3.6—Baseline net primary productivity (NPP) at 50 km and 10 km scales. 62 USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE WHERE D/3 D/3 D'1 AND D'1 ARE .00 RESPONSES FOREACHlNED?ANDCOARSED?RESOLUTIONGRIDTOTHE /35ANDTHE'&$,1CLIMATESCENARIOS4HISMETRICISTHE DELTATHATISPASSEDTOTHEFORESTSECTORMODELANDUSEDTO MODIFYTIMBERVOLUMEINCREASESINTHEINVENTORYMODEL .ETPRIMARYPRODUCTIVITYFORALLFORESTSINCREASEDUNDER BOTHCLIMATECHANGESCENARIOSATBOTHRESOLUTIONS5NDER THE/35CLIMATE.00OFALLFORESTSINCREASEDAPPROXIMATELY PERCENT ABOVE THE BASELINE .00 RESPONSE WHEREAS UNDER THE '&$,1 CLIMATE .00 OF ALL FORESTS INCREASED LESSTHANPERCENTlG7ITHINEACHCLIMATECHANGE SCENARIOTHERESPONSETOCLIMATECHANGEATTHECOARSERES OLUTIONDIFFEREDBYLESSTHANPERCENTFROMTHERESPONSE ATTHElNERRESOLUTION!GGREGATIONERRORFORALLFORESTSFOR THE/35CLIMATEnPERCENTWASSIMILARTOTHEERRORFOR THE'&$,1CLIMATEnPERCENTANDBOTHRESULTSWERE SIMILARTOTHEAGGREGATIONERROROFTHEBASELINECLIMATEn PERCENT 4HE SPATIAL PATTERNS OF THESE AGGREGATION ERRORS WERE SIMILAR BETWEEN THE TWO CLIMATE SCENARIOS AND THE BASELINEACROSSTHEHISTORICRANGEOFTEMPERATEFORESTS 7ITHINFORESTTYPES.00INCREASEDFROMTOPER CENT VARYING BY CLIMATE SCENARIO 4HE .00RESPONSE FOR BOREAL FORESTS AND FORESTED BOREAL WETLANDS WAS TO PERCENT GREATER UNDER THE '&$,1 CLIMATE THAN UNDER THE/35CLIMATE(OWEVERFORMIXEDTEMPERATEDECID UOUS AND TEMPERATE FORESTED WETLAND .00 UNDER THE /35 CLIMATE WAS TO PERCENT GREATER THAN UNDER THE'&$,1CLIMATE7ITHINEACHCLIMATESCENARIOFOREST .00RESPONSESATTHECOARSESCALEDIFFEREDBYLESSTHAN PERCENTFROMTHE.00RESPONSETOCLIMATECHANGEATTHE lNERRESOLUTIONFORALLFORESTTYPESEXCEPTCONIFERWHICH DIFFERED BY LESS THAN PERCENT 4HE ERROR IN ALL THREE CLIMATE SCENARIOS WAS HIGHEST IN THE BOREAL FORESTS AND FORESTED BOREAL WETLANDS AND PERCENT RESPEC TIVELY 4HE ABSOLUTE VALUE OF THE AGGREGATION ERROR FOR THE OTHER FORESTS WAS LESS THAN PERCENT 'EOGRAPHI CALLYAGGREGATIONERRORFORBOTH/35AND'&$,1ISCON CENTRATED IN THE SAME AREAS AS THAT OF BASELINE CLIMATE AROUND THE 'REAT ,AKES IN .EW %NGLAND AND IN THE 2OCKY -OUNTAINS 4HE SMALLEST AGGREGATION ERROR LESS THAN PERCENT WAS FOUND IN THE 3OUTH AND THE -ID !TLANTICANAREATHATHADTHEGREATESTDIFFERENCESINTHE .00RESPONSETOCLIMATECHANGEnTOPERCENT 2ELATIVETOTHEBASELINETHEPERCENTINCREASESORDECREASES IN .00ARE SIMILAR ACROSS THE lNE AND COARSE RESOLUTIONS WITHIN A CLIMATE CHANGE SCENARIO BUT DIFFER SIGNIlCANTLY ACROSS SCENARIOS lG 0ERCENT INCREASES IN .00 WERE SIMILARUNDERBOTHCLIMATESCENARIOSINTHE7ESTBUTTHE RESPONSESINTHE3OUTHAND-ID!TLANTICFORESTSWEREDRA MATICALLYDIFFERENT&ORTHE'&$,1CLIMATETHESOUTHERN ANDMID!TLANTICFOREST.00DECLINEDUPTOPERCENTREL ATIVETOTHEBASELINE.00WHEREASUNDERTHE/35CLIMATE THESEFORESTSINCREASEDIN.00FROMTOPERCENT 4HE KM GRIDCELL RESOLUTION IS MOST RELEVANT TO STANDLEVEL FOREST MANAGERS !GGREGATION ERROR AT THIS 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n *OYCEAND.UNGESSER SCALE IS LESS THAN PERCENT TWO STANDARD DEVIATIONS WHICHISWITHINTHEPERCENTPRECISIONTHATSTANDLEVEL .00ISMEASURED/URANALYSESINDICATETHATAGGREGATION ERROR IS THE LARGEST IN TRANSITION REGIONS AND IN REGIONS WITHSUBSTANTIALVARIABILITYINAIRTEMPERATURE!GGREGA TIONERRORISPRIMARILYASSOCIATEDWITHTHEREPRESENTATION OF A MOSAIC OF FOREST TYPES WITH A SINGLE FOREST TYPE AT KM RESOLUTION 4HIS SOURCE OF AGGREGATION ERROR CAN EASILY BE MINIMIZED BY MAKING .00ESTIMATES FOR EACH FORESTTYPEWITHINAKMGRIDCELLANDAGGREGATINGESTI MATESBASEDONTHEPROPORTIONOFEACHFORESTTYPEWITHIN THEGRIDCELL4HISAPPROACHHASBEENUSEDBY"ONAN AS A MEANS OF REPRESENTING VEGETATIVE HETEROGENEITY FOR ESTIMATINGCARBONWATERANDENERGYEXCHANGEINTHESUR FACEBOUNDARYLAYEROFGENERALCIRCULATIONMODELS0IERCE AND2UNNINGALSOFOUNDTHATAVERAGINGTEMPERA TURESUBSTANTIALLYINmUENCEDAGGREGATIONERRORINREGIONS OFCOMPLEXTERRAIN)TMAYBEPOSSIBLETOACHIEVECOMPUTA TIONALEFlCIENCYATKMRESOLUTIONBYAGGREGATINGTEM PERATUREFORALIMITEDNUMBEROFELEVATIONBANDSMAKING .00ESTIMATESFOREACHELEVATIONBANDANDAGGREGATING ESTIMATESBASEDONTHEPROPORTIONOFEACHELEVATIONBAND WITHINTHEGRIDCELL.UNGESSERETAL 4HERESOLUTIONOFFORESTTYPESISMOSTRELEVANTTOCOUN TRYSPECIlC ECONOMIC ASSESSMENTS OF THE IMPACTS OF CLI MATE CHANGE ON TIMBER RESOURCES &OR EXAMPLE RELATIVE CLIMATICRESPONSESOF.00FORDIFFERENTFORESTTYPESINDIF FERENTREGIONSWEREUSEDASINPUTSFORTHELAST20!*OYCE ET AL %XCEPT FOR BOREAL FORESTS AND FORESTED BOREAL WETLANDSTHEMEANAGGREGATIONERROROFBASELINE.00ESTI MATESISLESSTHANPERCENTFOREACHFORESTTYPE)NCONTRAST AGGREGATION ERROR FOR BOREAL FORESTS AND FORESTED BOREAL WETLANDSISAPPROXIMATELYPERCENT4HISLEVELOFERRORIS CAUSEDPRIMARILYBYTHEOVERREPRESENTATIONOFBOREALAND BOREALWETLANDFORESTSANDUNDERREPRESENTATIONOFMORE HIGHLYPRODUCTIVEFORESTSINTHEKMSIMULATION )NCOMPARISONTOTHEBASELINESIMULATIONSMEANAGGRE GATIONERRORFORTHEABSOLUTEESTIMATESOFEACHFORESTTYPE INTHECLIMATECHANGESIMULATIONSISSIMILAREXCEPTFORTEM PERATE CONIFER FOREST n PERCENT FOR /35 AND n PER CENTFOR'&$,VSnPERCENTFORBASELINECLIMATE3IMILAR TOAGGREGATIONERRORTHERELATIVERESPONSESOF.00ATEACH RESOLUTIONARESIMILAREXCEPTFORCONIFERFORESTSAND PERCENT LOWER RESPONSE FOR THE lNE RESOLUTION /35 AND '&$,SIMULATIONS 4HE NEGATIVE AGGREGATION ERRORS AND LOWER RESPONSES FOR TEMPERATE CONIFER FORESTS ARE ASSOCI ATED WITH THE AVERAGING OF TEMPERATURE IN THE COMPLEX TERRAIN IN THE NORTHERN 2OCKY -OUNTAINS AND IN WESTERN 7ASHINGTON /REGON AND #ALIFORNIA "ECAUSE DIFFERENCES BETWEENTHERESPONSESOF.00ATDIFFERENTRESOLUTIONSARE SMALLCOMPAREDWITHTHERESPONSESTODIFFERENTCLIMATESCE NARIOSTHEYCOULDBEIGNOREDINIMPACTASSESSMENTSTHAT EVALUATESENSITIVITIESTODIFFERENTCLIMATECHANGESCENARIOS 4HENATIONALRESOLUTIONISMOSTRELEVANTTOGLOBALECO NOMIC ASSESSMENTS OF THE IMPACTS OF CLIMATE CHANGE ON Joyce and Nungesser Ecosystem Productivity and the Impact of Climate Change Figure 3.7—Net primary productivity at 50 km and 10 km scales from the GFDL-Q climate scenario. 64 USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. Joyce and Nungesser Figure 3.8—Forest responses to two climate change scenarios (OSU, GFDL-Q) at 10 km (fine grid) and 50 km (coarse grid) scales. Ecosystem Productivity and the Impact of Climate Change USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000. 65 *OYCEAND.UNGESSER TIMBERRESOURCES4HERELATIVECLIMATICRESPONSESOF.00 OF HARDWOODS AND SOFTWOODS FOR DIFFERENT REGIONS WERE USED AS INPUTS FOR A NATIONAL ASSESSMENT 0EREZ'ARCIA ET AL &OR THE CONTERMINOUS 5NITED 3TATES AGGRE GATION ERROR WAS PERCENT IN THE BASELINE SIMULATION PERCENTINTHE/35SIMULATIONANDPERCENTINTHE '&$,SIMULATION2ESPONSESOF.00ACROSSTHETWOSPA TIALSCALESWEREANDPERCENTLOWERFORTHE/35AND '&$,SIMULATIONS"ECAUSETHESEDIFFERENCESAREMINUS CULE IN COMPARISON TO THE NATIONAL .00RESPONSES THEY CANBEIGNOREDINIMPACTASSESSMENTSTHATEVALUATESENSI TIVITIESTODIFFERENTCLIMATECHANGESCENARIOS 4HESE RESULTS INDICATE THAT .00 RESPONSES OF 4%- TO PROJECTED CLIMATE CHANGE ARE INSENSITIVE TO THE RESO LUTION OF INPUTS BUT THAT AGGREGATION ERROR OF ABSOLUTE .00ESTIMATESISSENSITIVETOTHERESOLUTIONOFINPUTSFOR SOME SITUATIONS %XCEPT FOR TRANSITION AREAS AND REGIONS WITH SUBSTANTIAL TEMPERATURE VARIABILITY THESE SIMULA TIONSINDICATETHATTHEUSEOFORESOLUTIONPROVIDESAN ACCEPTABLELEVELOFAGGREGATIONERRORATTHETHREESCALESOF ANALYSISINTHISSTUDY )TISIMPORTANTTORECOGNIZETHATTHECONCLUSIONSINTHIS STUDYAREBASEDONTWORESOLUTIONSANDONEBIOGEOCHEM ISTRY MODEL 0IERCE AND 2UNNING USED A DIFFERENT BIOGEOCHEMISTRYMODELTOSIMULATE.00FORVARIOUSRES OLUTIONS RANGING FROM KM TO KM IN A REGION OF COMPLEX TOPOGRAPHY !T THE COARSEST SCALE THEY FOUND COARSERESOLUTION .00 WAS OVERESTIMATED BY UP TO PERCENTRELATIVETO.00ESTIMATESATTHElNESTRESOLUTION 4HERESULTSOF.UNGESSERETALQUALITATIVELYAGREE WITHTHOSEOF0IERCEAND2UNNING "ECAUSEMOSTLARGESCALEBIOGEOCHEMISTRYMODELSARE PARAMETERIZED WITH STANDLEVEL DATA A SYSTEMATIC ANAL YSIS OF AGGREGATION ERROR WITH SEVERAL BIOGEOCHEMISTRY MODELS ACROSS A RANGE OF SPATIAL RESOLUTIONS FROM STAND TO EG M TO KM TO KM TO KM SHOULDBEUNDERTAKENINDIFFERENTFORESTREGIONSTODETER MINE WHETHER OUR CONCLUSIONS AND THOSE OF 0IERCE AND 2UNNINGAREROBUST &INALLY IT IS IMPORTANT TO VERIFY THE CONCLUSION ABOUT THE INSENSITIVITY OF .00 RESPONSES TO THE RESOLUTION OF INPUTSWITHOTHERBIOGEOCHEMISTRYMODELS"YCLARIFYING THE SCALING ISSUES ASSOCIATED WITH .00 ESTIMATES AND RESPONSESTHESESUGGESTEDSTUDIESWOULDIMPROVEIMPACT ASSESSMENTSTHATRELYONTHEESTIMATESOFLARGESCALEECO LOGICALMODELS #ONCLUSIONS !NALYSES OF THE IMPACT OF CLIMATE CHANGE ON FOREST PRODUCTIVITYBASEDONEXPERIMENTALRESEARCHANDMODEL %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE INGWOULDSUGGESTTHATFORESTPRODUCTIVITYMAYINCREASE UNDER ELEVATED CARBON DIOXIDE BUT THAT THE LOCAL CON DITIONS OF MOISTURE STRESS AND NUTRIENT AVAILABILITY WILL STRONGLYTEMPERANYRESPONSE0ROJECTEDINCREASESINPRO DUCTIVITY FROM CARBON DIOXIDE FERTILIZATION APPEAR TO BE WITHINTHESAMEMAGNITUDEASPOTENTIALINCREASESINPRO DUCTIVITY FROM TIMBER MANAGEMENT TREATMENTS 2ElNE MENTS IN THE ANALYSIS SUCH AS ANALYZING THE IMPACT AT A lNERSCALEDONOTAPPEARTOALTERTHERESULTSFROMTHELAST 20!ANALYSIS)NCORPORATINGLANDUSECHANGESAPPEARSTO BEACRITICALNEXTSTEPINTHEANALYSISOFTHEIMPACTOFCLI MATECHANGEONFORESTPRODUCTIVITY !CKNOWLEDGMENTS 7EWOULDLIKETORECOGNIZE$AVE-C'UIREFORHISCON TRIBUTIONS TO THE AGGREGATION RESEARCH DESCRIBED IN THIS CHAPTER3COTT0OWELLAND!LICIA,IZARRAGAPROVIDED')3 SUPPORTAND-ONICA%NGELPROVIDEDPROGRAMMINGSUP PORT+AREN&ASSNACHTASSISTEDWITHTHELITERATUREREVIEW 7E GREATLY APPRECIATE THE EDITORIAL REVIEW BY 2OBERT (AMREANDTHESTATISTICALREVIEWOF2UDY+ING7EALSO THANK$OMINIQUE"ACHELETFORHERHELPFULREVIEW 4HISWORKWASSUPPORTEDBYTHE53$!&OREST3ERVICE 2ESOURCES0ROGRAMAND!SSESSMENTSTAFFANDTHE.ORTH EASTERN'LOBAL#HANGE0ROGRAM ,ITERATURE#ITED !TKINSON#*4AYLOR*-7ILKINS$;ETAL=%FFECTSOFELEVATED #/ ON CHLOROPLAST COMPONENTS GAS EXCHANGE AND GROWTH OF OAK ANDCHERRY4REE0HYSIOLOGYn "ERNTSON'-"AZZAZ&!4HEALLOMETRYOFROOTPRODUCTIONAND LOSS IN SEEDLINGS OF !CER RUBRUM !CERACEAE AND "ETULA PAPYRIFERA "ETULACEAEIMPLICATIONSFORROOTDYNAMICSINELEVATED#/!MERI CAN*OURNALOF"OTANYn "IRDSEY2ICHARD!0LANTINGA!NDREW*(EATH,INDA30ASTAND PROSPECTIVECARBONSTORAGEIN5NITED3TATESFORESTS&OREST%COLOGY AND-ANAGEMENTn "IRDSEY2!(EATH,3#ARBONCHANGESIN53&ORESTS)N*OYCE ,!ED#LIMATECHANGEANDTHEPRODUCTIVITYOF!MERICASFORESTS 'EN4ECH2EP2-&ORT#OLLINS#/53$EPARTMENTOF!GRI CULTURE &OREST 3ERVICE 2OCKY -OUNTAIN &OREST AND 2ANGE %XPERI MENT3TATIONn "ONAN'",ANDATMOSPHERE#/EXCHANGESIMULATEDBYALAND SURFACEPROCESSMODELCOUPLEDTOANATMOSPHERICGENERALCIRCULATION MODEL*OURNALOF'EOPHYSICAL2ESEARCHn #EULEMANS23HAO"9*IANG8.;ETAL=&IRSTANDSECOND YEARABOVEGROUNDGROWTHANDPRODUCTIVITYOFTWO0OPULUSHYBRIDS GROWNATAMBIENTANDELEVATED#/4REE0HYSIOLOGYn 53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n %COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE #EULEMANS 2 -OUSSEAU - 4ANSLEY 2EVIEW .O %FFECTS OF ELEVATEDATMOSPHERIC#/ONWOODYPLANTS.EW0HYTOLOGIST n #OSTANZA 2 -AXWELL 4 2ESOLUTION AND PREDICTABILITY AN APPROACHTOTHESCALINGPROBLEM,ANDSCAPE%COLOGYn #RAMER 7 +ICKLIGHTER $7 "ONDEAU ! 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