%COSYSTEM )NTRODUCTION #(!04%2 ,INDA

advertisement
#(!04%2
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
,INDA!*OYCE53$!&OREST3ERVICE2OCKY-OUNTAIN2ESEARCH3TATION
-ARTHA.UNGESSER-!4#/-
)NTRODUCTION
%ARLIER ANALYSES OF THE SUPPLY AND DEMAND OF TIMBER
ASSUMEDTHECONTINUATIONOFHISTORICALCLIMATEANDTHUS
DIDNOTEXPLICITLYINCORPORATEFACTORSSUCHASTEMPERATURE
ORPRECIPITATIONINTOTHEPROJECTIONSOFTIMBERGROWTH&OR
ESTS ARE ADAPTED TO LOCAL CLIMATES AND CHANGES IN THESE
CLIMATES ARE LIKELY TO IMPACT FUTURE FOREST GROWTH AND
TIMBER OUTPUTS 7ITHIN THE STRATEGIC PLANNING PROCESS OF
THE&OREST3ERVICE*OYCEETALTHEANALYSISOFECO
SYSTEMPRODUCTIVITYASINmUENCEDBYCLIMATICFACTORSHAS
BEEN IDENTIlED AS A CRITICAL QUESTION IN ORDER TO ADDRESS
THECHALLENGINGPROBLEMSASSOCIATEDWITHCLIMATECHANGE
s 7HAT ARE THE LIKELY EFFECTS OF INCREASING ATMOSPHERIC
CARBONDIOXIDEANDPROSPECTIVECLIMATECHANGEONECO
SYSTEM PRODUCTIVITY AS MEASURED BY CHANGES IN NET
PRIMARYPRODUCTIVITY
)NTHELAST20!!SSESSMENT53$!&OREST3ERVICE
THECONSEQUENCESOFELEVATEDCARBONDIOXIDEANDCLIMATE
CHANGEONNETPRIMARYPRODUCTIVITYOFFORESTSWEREEXAM
INED USING CLIMATE MODEL SCENARIOS AND AN ECOLOGICAL
MODEL 4%- *OYCE ET AL 4HESE RESULTS WERE THEN
USED TO EXAMINE THE IMPACT OF CLIMATE CHANGE ON THE
SUPPLYANDDEMANDFORTIMBERPRODUCTSONPRIVATETIM
BERLANDS IN THE 5NITED 3TATES SEE lG *OYCE AND
"IRDSEYTHISVOLUME)NTHISANALYSISMOSTOFTHEFOREST
PRODUCTIVITYCHANGESACROSSTHE5NITED3TATESWEREPOSI
TIVELEADINGTOINCREASESINTHETIMBERINVENTORIES7ITH
THIS CHANGE IN HARVESTABLE INVENTORIES TIMBER HARVESTS
ACROSS THE 5NITED 3TATES SHIFTED AS DEMAND IN VARIOUS
REGIONS ADJUSTED TO TAKE ADVANTAGE OF LOWER COST RAW
MATERIALS3INCETHESELAST20!ANALYSESNEWEXPERIMEN
TALDATAANDMODELINGANALYSESENHANCETHEPICTUREOFTHE
POTENTIAL IMPACT OF CARBON DIOXIDE AND CLIMATE CHANGE
ONFORESTS)NTHISCHAPTERWEPRESENTTHERESULTSOFSUCH
RESEARCH
.EWEXPERIMENTSONTHEIMPACTOFCARBONDIOXIDEON
VEGETATIONANDMETAANALYSESOFTHEACCUMULATEDRESEARCH
HAVEDEMONSTRATEDTHEIMPACTOFCARBONDIOXIDEONPLANT
PROCESSES)NTHISCHAPTERWECOMPARETHEIMPACTOFELE
VATEDCARBONDIOXIDEONECOSYSTEMPRODUCTIVITYASDETER
MINEDFROMRECENTEXPERIMENTALDATAWITHPRODUCTIVITY
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
RESULTSFROMTHELAST20!ANDOTHERECOLOGICALMODELANAL
YSES)NADDITIONWEEXPLOREWHETHERTHESEPOTENTIALPRO
DUCTIVITYSHIFTSAREWITHINTHERANGEOFPRODUCTIVITYSHIFTS
THATTIMBERMANAGEMENTTREATMENTSCOULDINDUCEINMAN
AGEDFORESTS7ECOMPARETHERESULTSOFECONOMICRESEARCH
ANALYZING THE POTENTIAL TO INCREASE TIMBER GROWTH AS AN
INCENTIVETOSTORECARBONWITHTHESEPROJECTEDCHANGESIN
PRODUCTIVITYFROMCLIMATECHANGE
)NTHELAST20!!SSESSMENTTWOOFTHEMODELINGASSUMP
TIONSWERECLIMATEANDVEGETATIONWEREINEQUILIBRIUM
ANDTHESPATIALSCALEOFTHEECOLOGICALMODELWASANADE
QUATE SCALE FOR NATIONAL LEVEL ANALYSES 2ECENT MODELING
STUDIESHAVEEXAMINEDTHESEASSUMPTIONS
)N MOST ECOLOGICAL MODELS EVEN NOW VEGETATION IS
REPRESENTED AS PRISTINE MATURE VEGETATION RATHER THAN
THEACTUALVEGETATIONOFMANYDIFFERENTAGECLASSESSUC
CESSIONAL STAGES INTRODUCED SPECIES AND MANAGEMENT
HISTORIES &URTHER FOR BOTH THE ECOLOGICAL AND THE ECO
NOMIC MODELS A BROAD RANGE OF HABITATS AND SPECIES
WEREAGGREGATEDINTOECOSYSTEMSORTIMBERMANAGEMENT
TYPES RESPECTIVELY FOR MODELING PURPOSES 4HE VEGETA
TION AGGREGATION SCHEMES AND THE NATURE OF THE SPATIAL
EXTRAPOLATION DIFFER BETWEEN THE ECOLOGICAL AND THE ECO
NOMIC MODELS 4HE IMPLICATIONS OF AN ECOLOGICAL VERSUS
ATIMBERMANAGEMENTCLASSIlCATIONSCHEMEAREDISCUSSED
INTHISCHAPTER
#OMPUTATIONAL PROBLEMS ARISE WHEN INTEGRATING OR
LINKING MODELS THAT OPERATE AT DIFFERENT TEMPORAL AND
SPATIALSCALES#OMPUTATIONALLIMITATIONSFORCEATRADEOFF
BETWEENTHESPATIALEXTENTOFTHEANALYSISOFTENDICTATED
BY POLICY CONSIDERATIONS AND THE GRAIN OF THE ANALYSIS
#LIMATE MODELS IN ORDER TO COMPUTE GLOBAL DYNAMICS
OPERATE AT LARGE SPATIAL SCALES GRID CELLS RANGE FROM DEGREES TO DEGREES RESULTING IN COARSE RESOLUTION OF
MOUNTAINOUS REGIONS AND SMALL SCALE CLIMATE FEATURES
-ESOSCALE CLIMATE MODELING NOW OFFERS A lNER DEPIC
TIONOFCLIMATEFEATURESANDTHEPOSSIBILITYTOINCORPORATE
THEEFFECTSOFVEGETATIONANDLANDUSEFEEDBACKSONTOTHE
ATMOSPHERIC PROCESSES BUT AT LARGE COMPUTATIONAL COST
'IVENTHISOPPORTUNITYITISIMPORTANTTOUNDERSTANDTHE
UTILITYOFGOINGTOAlNERSCALEINTHEECOLOGICALANALYSES
BECAUSETHECOMPUTATIONALINTENSITYINCREASESBYANORDER
OFMAGNITUDEWHENAKMSCALEISUSEDINSTEADOFTHE
TRADITIONAL DEGREE SCALE )N THIS CHAPTER WE REPORT
RESEARCHRESULTSQUANTIFYINGTHECLIMATECHANGERESPONSES
INECOSYSTEMPRODUCTIVITYATlNERVERSUSCOARSERSPATIAL
SCALES
*OYCEAND.UNGESSER
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
0LANTAND%COSYSTEM
0RODUCTIVITY
)MPACTOF%LEVATED#ARBON$IOXIDEON
0RODUCTIVITY
.ETPRIMARYPRODUCTION.00ISTHEPROCESSBYWHICH
THEVEGETATIONINANECOSYSTEMCAPTURESCARBONFROMTHE
ATMOSPHERE4HECHANGESINECOSYSTEMPRODUCTIVITYPRO
JECTEDFROMECOLOGICALMODELSREmECTCLIMATECHANGESAS
WELL AS THE POTENTIAL INmUENCE OF CARBON DIOXIDE FERTIL
IZATIONONNETPRIMARYPRODUCTION)NTHELAST20!ANAL
YSIS *OYCE .00 OF TEMPERATE FORESTS IN THE 53
INCREASEDFROMPERCENTTOPERCENTDEPENDINGUPON
THECLIMATESCENARIOUSEDTABLE-ORERECENTANALYSES
USINGAREVISEDVERSIONOFTHE4ERRESTRIAL%COSYSTEM-ODEL
4%-CORROBORATETHESEEARLIERRESULTSWITHSOMEEXCEP
TIONS!LARGEMODELINGSTUDYCOMPAREDTHERESPONSESOF
SEVERALBIOGEOCHEMISTRYMODELSFORALLECOSYSTEMSWITHIN
THECONTERMINOUS5NITED3TATESTOALTEREDCLIMATEANDELE
VATEDCARBONDIOXIDE6%-!0MEMBERS!VERAGED
ACROSSALLECOSYSTEMS.00RESPONSESINCREASEDFROM
TOPERCENTTABLE4HE.00RESPONSESFROM4%-
WERE HIGHER THAN THE RESPONSES FOR THE #ENTURY -ODEL
0ARTONETALANDTHE"IOME"'#MODEL2UN
NING2UNNINGAND#OUGHLANTABLE4HE
LARGEST .00 RESPONSE FOR BOTH 4%- AND #ENTURY AND PERCENT RESPECTIVELY WAS FOR THE 5+-/ SCE
NARIOINCONTRASTTOTHERESPONSEOF"IOME"'#PER
CENTINCREASE4HELOWEST.00INCREASEFORBOTH4%-AND
#ENTURYWASREPORTEDFORTHE/35SCENARIOAGAININCON
TRASTTOTHERESPONSEOF"IOME"'#!TTHEGLOBALSCALE
THE.00RESPONSESTOCLIMATECHANGEANDELEVATEDCARBON
DIOXIDEFORAREVISEDVERSIONOF4%-8IAOETALARE
LOWERTHANTHE.00RESPONSESREPORTEDFORTHELAST20!
ANALYSIS(OWEVERTHISRESULTREmECTSTHEGLOBALEXTENTOF
THESEMEASURES
4HESEPROJECTEDRESPONSESTOALTEREDCLIMATEANDELE
VATEDATMOSPHERICCARBONDIOXIDEINCORPORATETHEVARIA
TION OF CLIMATE AND ECOSYSTEMS ACROSS THE 5NITED 3TATES
OR THE GLOBE )N CONTRAST EXPERIMENTAL STUDIES EXPLORE
THE UNDERLYING MECHANISMS FOR A PLANTS RESPONSE TO A
CHANGE IN ATMOSPHERIC CARBON DIOXIDE IN A CONTROLLED
ENVIRONMENT %LEVATED ATMOSPHERIC CARBON DIOXIDE HAS
BEENSHOWNTOINCREASEPHOTOSYNTHESISENHANCERATESOF
CARBONASSIMILATIONINCREASESTEMANDROOTBIOMASSAND
INTERACTWITHOTHERPLANTNUTRIENTS#EULEMANSAND-OUS
SEAU #URTIS AND 7ANG -C'UIRE ET AL B
3AXEETAL)NAREVIEWOFWOODYPLANTS#EULEMANS
AND -OUSSEAU REPORTED THE MEAN BIOMASS INCRE
MENTFROMELEVATED#/WASPERCENTFORCONIFEROUS
TREESANDPERCENTFORDECIDUOUSTREES&ORCONIFEROUS
TREESTHERANGEOFRESPONSESWASFROMPERCENTTO
PERCENT&ORDECIDUOUSTREESTHERANGEWASFROMnTO
PERCENT 3UMMARIZING STUDIES NOT INVOLVING STRESS
COMPONENTS 3AXE ET AL REPORTED LARGER AVERAGE
LONGTERMBIOMASSINCREMENTDIFFERENCESUNDERELEVATED
#/FORCONIFERSOFPERCENTANDSMALLERAVERAGESFOR
DECIDUOUSTREESPERCENT
4ABLEˆ.ETPRIMARYPRODUCTIONRESPONSETOCLIMATECHANGEANDELEVATEDCARBONDIOXIDEFORDIFFERENTBIOGEOCHEMICALMODELS
4%-4ERRESTRIAL%COSYSTEM-ODEL#EN#ENTURY-ODEL""'#"IOME"'#ANDATDIFFERENTSPATIALEXTENTS
#ONTERMINOUS5NITED3TATES
!LLFORESTS
'LOBE
!LLECOSYSTEMS
%COSYSTEMS
#LIMATESCENARIO
4%-
4%-
4%-
#EN
""'#
4%-
')33
'&$,
'&$,1
/35
'&$,2
5+-/
-)4,/
*OYCE
.UNGESSERETAL
6%-!0MEMBERS
8IAOETAL
')33REFERSTOTHESCENARIOFROMTHE'ODDARD)NSTITUTEFOR3PACE3CIENCEMODEL'&$,AND'&$,1REFERTORESULTSFROMTHE'EOPHYSICAL$YNAM
ICS,ABMODEL/35REFERSTOACLIMATEMODELDEVELOPEDBY3CHLESINGERANDOTHERSAT/REGON3TATE5NIVERSITY5+-/REFERSTOTHE5NITED+INGDOM
-ETEOROLOGY/FlCEMODELAND-)4REFERSTOTHE-ASSACHUSETTS)NSTITUTEOF4ECHNOLOGYMODEL
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
4HESE RESPONSES FROM EXPERIMENTAL STUDIES ARE NOT
WITHOUTCONTROVERSY/FPARTICULARCONCERNISWHETHERTHE
RESPONSEISSUSTAINEDOVERTHELIFESPANOFTHETREE-OST
OFTHESESTUDIESAREDONEWITHSEEDLINGSORJUVENILETREES
.ORBY ET AL CONCLUDED THAT A RESPONSE IN PRO
DUCTIVITY WAS THE RESULT OF AN EARLY STIMULUS AND THAT
NO FURTHER SUSTAINED RESPONSE WAS OBSERVED 'ORISSEN
ET AL SUGGEST THAT AN INITIAL GROWTH STIMULATION
MAYBECANCELEDBYLATERPHYSIOLOGICALORMORPHOLOGICAL
ADAPTATIONS &OR 9ELLOWPOPULAR ,IRIODENDRON TULIPIFERA
, WHOLEPLANT CARBON STORAGE DID NOT INCREASE EVEN
THOUGHLEAFLEVELPHOTOSYNTHESISANDLOWERRATESOFFOLIAR
RESPIRATIONIN#/ENRICHEDTREESWASOBSERVED.ORBYET
AL!NUMBEROFSTUDIESHAVESUGGESTEDTHATTHERE
MAYBEARESPONSESPECIlCITYAMONGTREEGENERA#EULE
MANSAND-OUSSEAUTOANINCREASEINATMOSPHERIC
#/ AS WELL AS WITHIN GENERA #EULEMANS ET AL $E,UCIAETALSUGGESTEDTHATALLOCATIONPATTERNSIN
PONDEROSAPINESMAYOFFSETANYINCREASESINPHOTOSYNTHE
SISRESULTINGINPOTENTIALDECLINESINPRODUCTIVITYUNDER
ALTEREDCLIMATEANDELEVATEDATMOSPHERICCARBONDIOXIDE
2ESPONSESINNATURALSTANDSAREUNCERTAIN
#URTIS AND 7ANG CONDUCTED A METAANALYSIS
OF OVER REPORTS ON EXPERIMENTS ON THE EFFECT OF ELE
VATED CARBON DIOXIDE ON WOODY PLANT MASS FORM AND
PHYSIOLOGY4HESESTUDIESSHOWEDSUBSTANTIALVARIATIONIN
PLANTRESPONSETOELEVATED#/RANGINGFROMINHIBITIONOF
GROWTHTOOVERPERCENTENHANCEMENTRELATIVETOPLANTS
GROWN IN AMBIENT CONDITIONS )RRESPECTIVE OF THE GROW
ING CONDITIONS THEY FOUND THAT TOTAL BIOMASS INCREASED
SIGNIlCANTLYATABOUTTWICEAMBIENTATMOSPHERICCARBON
DIOXIDECONCENTRATIONSAVERAGINGAPERCENTINCREASE
3TRESS ALTERED THE RESPONSES ,OW NUTRIENT AVAILABILITY
REDUCEDTHE#/RESPONSETOAPERCENTINCREASE,OW
LIGHTINCREASEDTHERESPONSETOPERCENT4HEYFOUNDNO
SHIFTSINBIOMASSALLOCATIONUNDERELEVATED#/"ELOW
GROUND RESPONSES WERE SENSITIVE TO LENGTH OF THE STUDY
ANDTHESTRESSESINDUCED
0AN ET AL EXAMINED THE MODELED RESPONSES
OFTERRESTRIALECOSYSTEMSTOELEVATEDATMOSPHERICCARBON
DIOXIDE 4HE FORESTED ECOSYSTEM .00 RESPONSE RANGED
FROMTOPERCENTINCREASESTABLE7HILETHESE
BIOGEOCHEMISTRYMODELSASSUMEOPTIMALRESPONSESSIM
ILAR TO THOSE OBSERVED EXPERIMENTALLY EG n PER
CENT THESE SPATIALLY EXTRAPOLATED RESPONSES TO ELEVATED
CARBON DIOXIDE BY ECOSYSTEM ARE SUBSTANTIALLY LOWER
7HEN EXAMINED FOR UNDERLYING DIFFERENCES 0AN ET AL
NOTEDTHATTHETHREEMODELSTENDTOAGREEINTHEIR
PROJECTED ESTIMATES OF .00RESPONSE TO DOUBLED CARBON
DIOXIDE ALONG PRECIPITATION GRADIENTS BUT DIFFER ALONG
TEMPERATUREGRADIENTS!LTHOUGHTHEEXPERIMENTALLITERA
TUREISEXPANDINGWITH#/IMPACTSTUDIESTHEREISLITTLE
INFORMATION ON THE RELATIVE ECOSYSTEMLEVEL RESPONSE OF
.00TOELEVATED#/ALONGCLIMATICGRADIENTS0ANETAL
4HESE BIOGEOCHEMISTRY MODELS SERVE AS DIFFERENT
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
*OYCEAND.UNGESSER
4ABLEˆ.ETPRIMARYPRODUCTION.00RESPONSETODOU
BLEDATMOSPHERIC#/VERSUSPPMVSIMULATEDBYTHE
6%-!0BIOGEOCHEMISTRYMODELS0ANETAL
&ORESTTYPE
")/-%"'# #ENTURY 4%-
"OREALCONIFER
-ARITIMECONIFER
#ONTINENTALCONIFER
#OOLTEMPERATEMIXED
7ARMTEMPERATESUBTROPICMIXED
4EMPERATEDECIDUOUS
4EMPERATEMIXEDXEROMOPHIC
4EMPERATECONIFERXEROMOPHIC
HYPOTHESESONHOWECOSYSTEMPROCESSESCONTROLTHE.00
RESPONSETOELEVATED#/
7HEN EXPERIMENTAL STUDIES SINCE REPORTING
CHANGESINBIOMASSAREGROUPEDBYFORESTTYPETHESPE
CIES RESPONSE IS VARIABLE TABLE 4HIS VARIABILITY IS
EXPLAINED IN SOME CASES BY THE DIFFERENT TREATMENTS
/PTIMALCONDITIONSSUCHASHIGH.TENDTOIMPROVETHE
BIOMASSRESPONSETOELEVATEDCARBONDIOXIDE
7ITHINFORESTTYPES.00RESPONSESTABLEFROMTHE
LAST 20!ANALYSIS RANGED FROM A PERCENT DECLINE FOR
TEMPERATE DECIDUOUS FOREST PRODUCTIVITY TO AN INCREASE
OFPERCENTFORBOREALFORESTPRODUCTIVITY4HEEXPERI
MENTALSTUDIESONWOODYSPECIESASSOCIATEDWITHBOREAL
FORESTTYPESSHOWEDRESPONSESTOELEVATEDCARBONDIOXIDE
OFTOPERCENTINCREASESTABLE2ESULTSFROMTHE
MODELINGSTUDIESWHICHINCLUDEPOTENTIALCHANGESINCLI
MATEASWELLASCARBONDIOXIDERANGEDFROMINCREASESOF
TOPERCENTTABLE&ORTHETEMPERATEDECIDU
OUS SPECIES THE EXPERIMENTAL RESULTS INCLUDED A DECLINE
OFPERCENTTOANINCREASEOFPERCENT2ESULTSINTHE
MODELEDSTUDIESFORTEMPERATEDECIDUOUSFORESTSRANGED
FROM A DECLINE OF PERCENT TO AN INCREASE OF PER
CENT&ORCONIFERSPECIESTHEEXPERIMENTALRESULTSRANGED
FROMNOSIGNIlCANTINCREASESTOANINCREASEOFPERCENT
2ESPONSESFROMTHEMODELEDSTUDIESFORTEMPERATECONIF
EROUSFORESTSRANGEDFROMAPERCENTINCREASETOA
PERCENTINCREASEIN.004HESEPROJECTEDRESPONSESTOELE
VATED#/ANDCLIMATEINTHELAST20!ANALYSESARELOWER
THANTHEPOTENTIALRESPONSESINTHEEXPERIMENTALSTUDIES
TABLEVERSUSTABLE
0ANETALDETECTEDTHEDIFFERENTECOSYSTEMLEVEL
HYPOTHESES THAT THESE BIOGEOCHEMISTRY MODELS REmECT
4HESEAREASOFUNCERTAINTYIFEXAMINEDIDENTIFYOPPORTU
NITIESTORElNEOURABILITYTOASSESSTHEIMPACTOFCLIMATE
CHANGEONECOSYSTEMS
s7HATROLEDOESTHEHYDROLOGICALCYCLEPLAYINCONTROL
LING THE #/ RESPONSES OF LEAF AREA AND SOIL MOISTURE
ALONGTEMPERATUREANDMOISTUREGRADIENTS
*OYCEAND.UNGESSER
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
4ABLEˆ"IOMASSRESPONSEPERCENTBYWOODYSPECIESUNDERELEVATEDCARBONDIOXIDEINEXPERIMENTALSTUDIES
3PECIES
3IGNIlCANT
.ONSIGNIlCANT
"OREAL
0ICEAABIES
0ICEAGLAUCA
0ICEAMARIANA
0ICEAMARIANA
0ICEASITCHENSIS
0ICEASITCHENSIS
0ICEASITCHENSIS
0ICEASITCHENSIS
!BOVEGROUNDBIOMASSFRESHWT
4OTALBIOMASS
4OTALBIOMASS
4OTALBIOMASS
4OTALBIOMASSIRRIGATIONFERTILIZATION
4OTALBIOMASSFERTILIZATION
4OTALBIOMASSNOIRRIGATIONORFERTILIZATION
4OTALBIOMASSIRRIGATION
4EMPERATECONIFEROUS
0INUSBANKSIANA
0INUSSILVESTRIS
0INUSSILVESTRIS
0INUSTAEDA
0INUSTAEDA
0INUSTAEDA
0INUSTAEDA
0INUSTAEDA
0INUSTAEDA
0INUSTAEDA
0INUSPONDEROSA
0INUSPONDEROSA
0INUSPONDEROSA
0INUSPONDEROSA
0INUSPONDEROSA
0INUSPONDEROSA
0INUSPONDEROSA
0SEUDOTSUGAMENZIESII
0SEUDOTSUGAMENZIESII
2OOTBIOMASS
4OTALBIOMASSHIGH.
4OTALROOTBIOMASS
4OTALROOTBIOMASS
4OTALROOTBIOMASS
4OTALROOTBIOMASS
4OTALBIOMASS
4OTALBIOMASSHIGHTEMP
4OTALBIOMASSHIGH.
4OTALROOTBIOMASS
4OTALROOTBIOMASS
4OTALROOTBIOMASS
4OTALBIOMASS
3HOOTBIOMASS
4OTALBIOMASSLOW.
4OTALBIOMASSLOWTEMP
4OTALBIOMASSLOW.
4OTALBIOMASSAGE
4OTALBIOMASSAGE
.3
9AKIMCHUKAND
(ODDINOTT
)NEICHENETAL
)NEICHENETAL
'RIFlNETAL
'RIFlNETAL
+INGETAL
+INGETAL
+INGETAL
+INGETAL
4ISSUEETAL
$ELUCIAETAL
$ELUCIAETAL
'RIFlNETAL
'RIFlNETAL
+INGETAL
+INGETAL
+INGETAL
'ORISSENETAL
'ORISSENETAL
4EMPERATEDECIDUOUS
0RUNUSAVIUM
0RUNUSAVIUM
0RUNUSAVIUM
0RUNUSAVIUM
0RUNUSAVIUM8
PSEUDOCERASUS
1UERCUSROBUR
1UERCUSRUBRA
1UERCUSRUBRA
!LNUSRUBRA
!LNUSRUBRA
0OPULUSDELTOIDESX
NIGRA
0OPULUSDELTOIDESX
NIGRA
0OPULUSDELTOIDESX
NIGRA
4OTALBIOMASSLOW.
4OTALBIOMASSDECLINE
4OTALBIOMASSHIGH.
4OTALBIOMASSHIGH.
,EAFSHOOTMONTH
,EAFSHOOTMONTH
4OTALBIOMASSLEAFMASS
4OTALBIOMASS
4OTALBIOMASS
4OTALBIOMASS
3TEMVOLUME
4OTALBRANCHBIOMASS
4OTALBIOMASSOFLEAVES
,EAFSHOOT
ROOTMONTH
7ILKINSETAL
7ILKINSETAL
7ILKINSETAL
7ILKINSETAL
!TKINSONETAL
!TKINSONETAL
,INDROTHETAL
-IAO
(IBBSETAL
(IBBSETAL
#EULEMANSETAL
#EULEMANSETAL
#EULEMANSETAL
0ERCENT
2ESPONSE
3OURCE
0OLLEETAL
9AKIMCHUKAND
(ODDINOTT
,ORDETAL
9AKIMCHUKAND
(ODDINOTT
4OWNEND
4OWNEND
4OWNEND
4OWNEND
CONTINUED
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
*OYCEAND.UNGESSER
4ABLECONTINUED
3PECIES
3IGNIlCANT
.ONSIGNIlCANT
0ERCENT
2ESPONSE
0OPULUSDELTOIDESX
NIGRA
0OPULUSDELTOIDESX
NIGRA
0OPULUSTREMULOIDES
0OPULUSTRICHOCARPA
XDELTOIDES
0OPULUSTRICHOCARPA
XDELTOIDES
0OPULUSTRICHOCARPA
XDELTOIDES
!CERRUBRUM
!CERSACCHARUM
!CERSACCHARUM
!CERSACCHARUM
"ETULAALLEGHANIENSIS
"ETULAALLEGHANIENSIS
FAMILY'
"ETULAALLEGHANIENSIS
FAMILY7
"ETULAALLEGHANIENSIS
FAMILY9
"ETULALENTA
"ETULAPAPYRIFERA
"ETULAPAPYRIFERA
"ETULAPOPULIFOLIA
,IRIODENDRONTULIPIFERA
,IRIODENDRONTULIPIFERA
4OTALBIOMASS
#URTISETAL
4OTALBIOMASS
#URTISETAL
4OTALBIOMASSLEAFMASS
3TEMVOLUME
4OTALBRANCHBIOMASS
4OTALBIOMASSOFLEAVES
3TEMMASSROOTMASSLEAFMASS
4OTALBIOMASS
4OTALBIOMASS
3TEMMASSROOTMASSLEAFMASS
4OTALBIOMASSlNECOARSEROOTMASS
3TEMMASSROOTMASSLEAFMASS
3TEMMASSROOTMASSLEAFMASS
4APROOT
4OTALlNECOARSEROOTMASS 4OTALBIOMASS
4OTALBIOMASS
4OTALBIOMASS
4OTALBIOMASS
n
"RANCHESLEAVESBOLE
3OURCE
,INDROTHETAL
#EULEMANSETAL
#EULEMANSETAL
#EULEMANSETAL
"ERNTSONAND"AZZAZ
,INDROTHETAL
.OBLEETAL
.OBLEETAL
2OCHEFORTAND"AZZAZ
7AYNEAND"AZZAZ
7AYNEAND"AZZAZ
7AYNEAND"AZZAZ
2OCHEFORTAND"AZZAZ
"ERNTSONAND"AZZAZ
2OCHEFORTAND"AZZAZ
2OCHEFORTAND"AZZAZ
.ORBYETAL
.ORBYETAL
4ABLEˆ#OMPARISONOFPROJECTEDCHANGESINFORESTPRODUCTIVITYUNDERCLIMATECHANGEANDELEVATEDCARBONDIOXIDE
4%-
&ORESTTYPE
"OREAL
"OREALFORESTWETLAND
4EMPERATECONIFER
4EMPERATEDECIDUOUS
4EMPERATEMIXED
4EMPERATEBROADLEAVEDEVERGREEN
4EMPERATEFORESTWETLAND
!LLFORESTS
4%-
'&$,
'&$,1
')33
/35
/35
'&$,1
n
n
*OYCE
.UNGESSERETAL
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
*OYCEAND.UNGESSER
s 7HAT ROLE DOES THE NITROGEN CYCLE PLAY IN THE #/
RESPONSESOFLEAFAREAANDLEAFNITROGENCONTENTALONG
TEMPERATUREANDMOISTUREGRADIENTS
s7HATISTHERELATIVEROLEOFCHANGESINNITROGENREQUIRE
MENTS ALLOCATION TISSUE # TO . RATIOS AND RATES OF
DECOMPOSITION IN DETERMINING #/ RESPONSES ALONG
TEMPERATUREANDMOISTUREGRADIENTS
s 7HAT ARE THE RELATIVE CONTRIBUTIONS AND IMPORTANCE
OFINTERACTIONSBETWEENTHEHYDROLOGICALANDNUTRIENT
CYCLESINCONTROLLING.00RESPONSESTOELEVATED#/
)MPORTANTLY0ANETALCONCLUDETHATFUTURESTUD
IES SHOULD MEASURE THE mUXES AND THE POOLS OF CARBON
NITROGEN AND WATER ! CLEAR PICTURE OF BOTH mUXES AND
POOLSISIMPORTANTINIMPROVINGOURUNDERSTANDINGOFTHE
INTERACTIONSAMONGPROCESSESTHATCONTROL#/RESPONSES
OF ECOSYSTEMS /UR UNDERSTANDING OF THESE PROCESSES
IS THE BASIS FOR THE DEVELOPMENT OF POLICIES ON CARBON
SEQUESTRATIONOPTIONSINFORESTS
#LIMATE6ERSUS-ANAGEMENT)NmUENCESIN
4IMBER0RODUCTIVITY
4HEPRODUCTIVITYSHIFTSINTHELAST20!CLIMATECHANGE
ANALYSISWEREARESPONSETOINCREASEDATMOSPHERICCARBON
DIOXIDE AND CHANGES IN TEMPERATURE AND PRECIPITATION
4HE TIME PERIOD WAS YEARS !RE THOSE PRODUCTIVITY
SHIFTSSIMILARTOTHEBIOLOGICALPOTENTIALOFCURRENT53FOR
ESTS/RARETHOSEPRODUCTIVITYSHIFTSSIMILARTOINCREASES
SEEN UNDER ECONOMIC OPPORTUNITIES FOSTERED BY TIMBER
MANAGEMENT OVER A SIMILAR TIME FRAME 6ASIEVICH AND
!LIGUSEDFORESTINVENTORYDATATOASSESSTHEPOTEN
TIAL TO INCREASE TIMBER GROWTH FOR CARBON STORAGE "IO
LOGICAL OPPORTUNITIES WERE DElNED AS THE POTENTIAL NET
ANNUALGROWTHOFTHEMOSTPRODUCTIVEPLOTSTOPOF
MEASURED PLOTS FOR EACH SITE CLASS FOREST MANAGEMENT
TYPE AND TREATMENT OPPORTUNITY ON TIMBERLAND SUITABLE
FOR TREATMENT 4HIS ESTIMATE REPRESENTS ACTUAL MANAGE
MENT BEING APPLIED TO CURRENT STANDS AND WAS THUS
DEEMEDACHIEVABLE%CONOMICOPPORTUNITIESWEREDElNED
ASINCREASESINGROWTHONTIMBERLANDTHATCOULDBETREATED
ANDYIELDORMOREONTHEDIRECTCOSTSOFTREATMENT
"ASEDON6ASIEVICHAND!LIGSANALYSISBIOLOGI
CALOPPORTUNITIESEXISTTOINCREASETIMBERGROWTHBYABOUT
BILLION CUBIC FEET OVER MILLION ACRES AN INCREASE
OF PERCENT OVER THE CURRENT NET ANNUAL GROWTH OF BILLION CUBIC FEET 3EVERAL DECADES WOULD BE REQUIRED TO
IMPLEMENT THE TREATMENTS TO ATTAIN THESE INCREASES &OR
ECONOMIC OPPORTUNITIES 6ASIEVICH AND !LIG ESTI
MATEDTHATNETANNUALGROWTHCOULDBEINCREASEDBY
BILLIONCUBICFEETAPPROXIMATELYPERCENTOFTHECURRENT
NET ANNUAL GROWTH #APITAL INVESTMENT COSTS WOULD BE
BILLION 4HESE BIOLOGICAL AND ECONOMIC OPPORTUNI
TIESWOULDTAKEDECADESTOIMPLEMENTWITHTHEFULLEFFECT
NOTBEINGSEENUNTILNEARTHEENDOFTHESTCENTURY4HUS
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
TIMBERMANAGEMENTCOULDPOTENTIALLYENHANCEFORESTPRO
DUCTIVITYTOALARGERDEGREETOPERCENTTHANISCUR
RENTLYPROJECTEDFORTHEPRODUCTIVITYRESPONSESTOCHANGES
INCARBONDIOXIDEORCLIMATETOPERCENTTABLE
0OTENTIAL6EGETATIONAND#URRENT
6EGETATION$ESCRIPTIONS
)N THE LAST &OREST 3ERVICE CLIMATE CHANGE ASSESSMENT
.00RESPONSEFROMTHEECOLOGICALMODEL4%-WASUSED
TO ADJUST GROWTH IN THE FOREST SECTOR MODEL 4!--
!4,!3.!0!0lG*OYCEAND"IRDSEYTHISVOLUME
4HEVEGETATIONCLASSIlCATIONSYSTEMSOFTHESETWOMODELS
DIFFERED#LASSIlCATIONSYSTEMSWITHINTHEFORESTRYSECTOR
HAVE FOCUSED ON COMMERCIAL TIMBER WHILE CLASSIlCATION
SYSTEMSWITHINBOTANYANDECOLOGYHAVEFOCUSEDONTHE
DYNAMICS OF PRISTINE PLANT COMMUNITIES )MBEDDED IN
THELAST20!CLIMATECHANGEASSESSMENTISTHECONVERSION
OF.00RESPONSESFROMTHEECOSYSTEMCLASSIlCATIONUSED
IN 4%- INTO THE FOREST MANAGEMENT TYPES USED IN THE
4!--!4,!3.!0!0MODEL
7ITHIN THE FOREST SECTOR MODEL YIELD TABLES TO PROJECT
TIMBERGROWTHAREDERIVEDFROMINVENTORYPLOTDATACOL
LECTEDOVERAPERIODOFSEVERALYEARSWITHINEACHOFTHESIX
FORESTINVENTORYREGIONSINTHE5NITED3TATES/NEOFTHE
STRATIlCATIONSFORTHESEYIELDTABLESISTIMBERTYPESTABLE
4HE &OREST 3ERVICE INVENTORY CLASSIlES FOREST LAND
BYFORESTTYPESINWHICHTHENAMEDSPECIESEITHERSINGLY
ORINCOMBINATIONCOMPRISEAPLURALITYOFLIVETREESTOCK
ING 4HE INVENTORY TYPES ARE BASED ON A STANDARD SET OF
LOCAL FOREST TYPES IN THE &OREST 3ERVICE (ANDBOOK ORGA
NIZEDINTOBROADERFORESTTYPEGROUPSTOFACILITATEREPORT
ING4HEREISSOMEAGGREGATIONOFTHEINVENTORYFORESTTYPES
INTO THE FOREST MANAGEMENT TYPES USED IN THE 4!--
!4,!3.!0!0MODEL4HENAMEDSPECIESTYPICALLYREFERS
TOACOMMERCIALTREESPECIESFOREXAMPLE$OUGLASlRORTO
ACLASSOFlBERSUCHASSOFTWOODMIX4HE4!--!4,!3
.!0!0 MODEL WAS NOT DEVELOPED TO MODEL GEOGRAPH
ICALLY RESOLVED DATA HENCE THE YIELD TABLES REPRESENTED
LARGER GEOGRAPHIC REGIONS TYPICALLY A MULTISTATE GROUP
INGOFOWNERSHIPFORESTTYPEAGECLASSES
&ORTHE4%-MODELSIMILARTOOTHERBIOGEOCHEMISTRY
MODELSDATAFROMINTENSIVELYSTUDIEDECOSYSTEMSREPRE
SENTATIVEOFPARTICULARVEGETATIONTYPESAREUSEDFORCAL
IBRATION OF MODEL BEHAVIOR 4HESE MODELS ARE SPATIALLY
EXTRAPOLATED BY USING VEGETATION MAPS &OR THE 4%-
MODELTHE5NITED3TATESHASBEENGRIDDEDINTODEGREE
BYDEGREEGRIDCELLS4HE+àCHLERCLASSIlCATIONSYSTEM
+àCHLER HAS BEEN USED TO ASSIGN THE DOM
INANT POTENTIAL NATURAL VEGETATION 0.6 TYPE TO EACH
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
*OYCEAND.UNGESSER
4ABLEˆ4IMBERTYPESUSEDINTHE4!--!TLASFORESTSECTORMODEL
0ACIlC.ORTHWEST
7EST
%AST
0ACIlC3OUTHWEST
.ORTH
3OUTH
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
$OUGLASlR
$OUGLASlRMIXED
$OUGLASlRLARCH
0UREHEMLOCK
&IRSPRUCE
4RUElR
0INES
,ODGEPOLEPINE
0ONDEROSAPINE
3OFTWOODMIX
*ACKPINE
2EDPINE
7HITEPINE
7HITEREDJACKPINE
3PRUCElR
2EDALDER
2EDWOOD
(ARDWOODMIX
(ARDWOOD
-IXEDCONIFER
3WAMPCONIFER
/AKHICKORY
,OWLANDHARDWOODS
-APLEBEECH
,OBLOLLY324OAKGUM
/AKPINE
%LMASHREDMAPLE
-APLEBEECHBIRCH
!SPENBIRCH
0LANTEDPINE
.ATURALPINE
5PLANDHARDWOOD
,OWLANDHARDWOOD
2OCKY-OUNTAIN
.ORTHCENTRAL
.ORTH
8
8
8
8
8
8
3OUTH
8
GRID CELL -C'UIRE ET AL AND THESE TYPES WERE
THEN AGGREGATED INTO A SMALLER SET OF ECOSYSTEM TYPES
INCLUDINGFORESTTYPESBOREALFORESTBOREALFORESTWET
LAND TEMPERATE CONIFEROUS FOREST TEMPERATE DECIDUOUS
FORESTTEMPERATEMIXEDFORESTTEMPERATEBROADLEAFEVER
GREENFORESTANDTEMPERATEFORESTWETLAND4HISSPATIALLY
EXPLICIT VEGETATION INFORMATION IS THEN USED TO EXTRAPO
LATETHEECOLOGICALMODELTOTHELARGERSPATIALSCALEOFTHE
5NITED3TATES)NTHELAST20!ANALYSIS*OYCE.00
RESPONSESTOCLIMATECHANGEWERECOMPUTEDFOREACH
DEGREEGRIDCELLINTHE5NITED3TATES4HUS.00RESPONSE
DATAIN4%-WASRESOLVEDATAlNERGEOGRAPHICSCALETHAN
REGIONAL VOLUME CHANGES IN THE 4!--!4,!3.!0!0
MODEL
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
8
8
8
8
8
8
8
8
8
8
8
8
8
8
8
!METHODWASNECESSARYTORECLASSIFYINFORMATIONFROM
4%-SOTHATTHISINFORMATIONCOULDBEUSEDATTHESCALEOF
THEFORESTPOLICYMODEL!STHEECOLOGICALANDTHETIMBER
CLASSIlCATIONSYSTEMSHADBOTHBEENRELATEDTOTHE0.6
CLASSIlCATION -C'UIRE ET AL 'ARRISON ET AL %YRETHISCLASSIlCATIONWASUSEDTOLINK4%-ECO
SYSTEMTYPESWITHTIMBERMANAGEMENTTYPESTABLE
4HE CONVERSION PROCESS INVOLVED ASSOCIATING THE 4%-
ECOSYSTEM TYPE AND THE 0.6 TYPE WITHIN EACH GRID
CELL-C'UIREETALTOTHEFORESTTYPECORRESPONDING
TOTHAT0.6TYPEASDElNEDBY'ARRISONETAL
3EVERALASSUMPTIONSWEREMADEIFNOGRIDCELLWITHIN
AREGIONWASDOMINATEDBYTHE0.6TYPEASSOCIATEDWITH
THE FOREST MANAGEMENT TYPE 4HIS SITUATION MIGHT ARISE
+UCHLER
7ESTERNSPRUCElR
3OUTHWESTERNSPRUCElR
'REAT,AKESSPRUCElR
.ORTHEASTERNSPRUCElR
3OUTHEASTERNSPRUCElR
3PRUCECEDARHEMLOCK
#EDARHEMLOCK$OUGLASlR
3ILVERlR$OUGLASlR
&IRHEMLOCK
-IXEDCONIFER
2EDWOOD
2EDlR
0INECYPRESS
,ODGEPOLEPINESUBALPINE
0ONDEROSASHRUBNONE
7ESTERNPONDEROSA
$OUGLASlR
#EDARHEMLOCKPINE
'RANDlR$OUGLASlR
%ASTERNPONDEROSAPINE
"LACK(ILLSPINE
0INE$OUGLASlR
!RIZONAPINE
3PRUCElR$OUGLASlR
'REAT,AKESPINE
-OSAICOFCEDARHEMLOCK$OUGLAS
lRFORESTAND/REGONOAKWOODS
.ORTHERNHARDWOODS
.ORTHERNHARDWOODSlR
.ORTHERNHARDWOODSSPRUCE
-OSAICOF!PPALACHIANOAKAND
.ORTHERNHARDWOODS
.ORTHEASTERNOAKPINE
3ERALNORTHERNHARDWOODS
3ERALNORTHERNHARDWOODSlR
.ORTHERNHARDWOODSSPRUCE
6%-!0VEGETATION
"OREALCONIFEROUS
4EMPERATEMARITIME
CONIFEROUSFOREST
4EMPERATECONTINENTAL
CONIFEROUSFOREST
#OOLTEMPERATE
MIXEDFOREST
,OBLOLLYSHORTLEAF
!SPENBIRCH%!34
!SPENBIRCH%!34
!SPENBIRCH%!34
$OUGLASlR
-APLEBEECHBIRCH
-APLEBEECHBIRCH
-APLEBEECHBIRCH
&IRSPRUCE
0ONDEROSAPINE
,ODGEPOLEPINE
0ONDEROSAPINE
0ONDEROSAPINE
$OUGLASlR
7ESTERNWHITEPINE
,ARCH
0ONDEROSAPINE
0ONDEROSAPINE
0ONDEROSAPINE
&IRSPRUCE
&IRSPRUCE
7HITEREDJACKPINE
(EMLOCK3ITKASPRUCE
$OUGLASlR
&IRSPRUCE
&IRSPRUCE
0ONDEROSAPINE
2EDWOOD
&IRSPRUCE
&IRSPRUCE
3PRUCElR
3PRUCElR
3PRUCElR
&)!
CONTINENTALTEMPERATECONIFEROUS
FORESTTEMPERATEDECIDUOUSFOREST
#ONTINENTALTEMPERATECONIFEROUSFOREST
-ARITIMETEMPERATECONIFEROUSFOREST
"OREALCONIFEROUSFOREST
4%-
CONTINUED
4ABLEˆ4RANSLATIONOFTHE&OREST#LASSIlCATIONUSEDINTHE&OREST3ERVICE&)!INVENTORYUSING0OTENTIAL6EGETATIONOF+àCHLERTOTHE6%-!0CLASSIlCATION
*OYCEAND.UNGESSER
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
/REGONOAKWOODS
.ORTHERNmOODPLAIN
-APLEBASSWOOD
/AKHICKORY
%LMASH
"EECHMAPLE
-IXEDMESOPHYTIC
!PPALACHIANOAK
4EMPERATEDECIDUOUSFOREST
7ESTERNHARDWOODS
%LMASHCOTTONWOOD
-APLEBEECHBIRCH
/AKHICKORY
%LMASHCOTTONWOOD
-APLEBEECHBIRCH
-APLEBEECHBIRCH
/AKHICKORY
$OUGLASlR
/AKHICKORY
/AKGUMCYPRESS
/AKGUMCYPRESS
/AKGUMCYPRESS
,OBLOLLYSHORTLEAFPINE
ANDOAKPINE
,ONGLEAFSLASHPINE
ANDOAKPINE
,OBLOLLYSHORTLEAF
,ONGLEAFSLASH
&)!
4EMPERATEDECIDUOUSFOREST
CONTINENTALTEMPERATECONIFEROUS
FORESTTEMPERATEDECIDUOUS
FORESTTEMPERATEBROADLEAVED
EVERGREENFOREST
4%-
+àCHLERNUMBERINGSCHEME4HElRSTNUMBERINEACHBOXISTHENUMBERREPORTEDINTHE+àCHLERPUBLICATION4HENUMBERINAFTERTHETYPENAMEISTHENUMBERREPORTEDONTHE
MAPWHICHWASALSOTHENUMBERUSEDINTHE&OREST3ERVICEPUBLICATIONMAPPING&2%3%COSYSTEMSANDTHE+àCHLER3YSTEM
)N6%-!0VEGETATIONTYPESREPRESENTEDBYMAPSYMBOLSNEVERDOMINATEDATTHELONGITUDEsLATITUDEGRIDCELLREPRESENTATION4YPES
-ESQUITEBOSQUES3EAOATSPRAIRIEAND.ORTHERNCORDGRASSPRAIRIEARENONFORESTTYPES&ORTHESETYPES)USEDATABLEPROVIDEDBY$AVE+ICKLIGHTERTHATHADIDENTIlEDTHE
4%-6%-!0TYPEASSOCIATEDWITHTHESE+àCHLERTYPES
&)! TYPES OF !SPENBIRCH AND ARE EAST AND WEST EXPRESSIONS OF THE !SPENBIRCH TYPE)N THE %AST THIS TYPE WAS INDEXED TO THE SERAL STAGES OF .ORTHERN HARDWOODS .ORTHERN
HARDWOODSlRFORESTSAND.ORTHERNHARDWOODSSPRUCEFORESTANDIDENTIlEDASTEMPERATEDECIDUOUS6%-!0TYPE)NTHE7ESTTHISTYPEWASALSOCODEDASTEMPERATEDECIDUOUS6%-!0
TYPE!LL&)!WESTERNHARDWOODSTYPEWEREIDENTIlEDASTEMPERATEDECIDUOUS6%-!0TYPE
#ALIFORNIAMIXEDEVERGREEN
"LACKBELT
,IVEOAKSEAOATS
#YPRESSSAVANNA
-ANGROVE
/AKHICKORYPINE
3OUTHERNMIXED
3ANDPINESCRUB
3UBTROPICALPINEFOREST
+UCHLER
7ARMTEMPERATESUBTROPICAL
MIXEDFOREST
6%-!0VEGETATION
4ABLECONTINUED
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
*OYCEAND.UNGESSER
*OYCEAND.UNGESSER
IF SILVICULTURAL MANAGEMENT FAVORED A SERAL SPECIES OR
IF CLIMAX VEGETATION WERE NOT DOMINANT WITHIN PARTS OF
THEREGIONORIFTHECOMMERCIALSPECIESHAPPENEDNOTTO
BETHETREESPECIESNAMEDINTHECLIMAXTYPE&OREXAM
PLESOFTWOODMIXAFORESTMANAGEMENTTYPEUSEDINTHE
0ACIlC.ORTHWESTWESTTIMBERSUPPLYREGIONWASAMIX
OFTWOFORESTTYPESREDWOODANDLARCH.O0.6TYPEWAS
REPRESENTATIVEOFREDWOODWITHINTHE0ACIlC.ORTHWEST
WESTREGIONWESTSIDEOF/REGONAND7ASHINGTON4HE
NEAREST 0.6 TYPE WAS THE REDWOOD TYPE IN THE 0ACIlC
3OUTHWEST REGION (ERE THE ASSUMPTION WAS MADE THAT
THERESPONSETOCLIMATECHANGEWOULDBEMOREAPPROPRI
ATELY DESCRIBED BY USING A SIMILAR ECOSYSTEM TYPE BUT
OUTSIDE OF THE REGION RATHER THAN A DIFFERENT VEGETATION
BUT INSIDE THE REGION )N ANOTHER EXAMPLE IN THE 0ACIlC
.ORTHWESTWEST REGION RED ALDER WAS A FOREST MANAGE
MENTTYPETHATDIDNOTHAVEACORRESPONDING0.6TYPE
!CCORDINGTO%YREREDALDERISASUCCESSIONALTYPE
REPLACEDBYTHE0ACIlC$OUGLASlRANDWESTERNHEMLOCK
SITKA SPRUCE TYPES %YRE DID ASSIGN RED ALDER TO
THEINVENTORYTYPEOFWESTERNHARDWOODSAND'ARRISON
ET AL ASSIGNED THE 0.6 TYPE /REGON OAKWOOD
WITHTHEINVENTORYTYPEOFWESTERNHARDWOODS(OWEVER
THIS0.6TYPEOCCURSSOLELYALONGTHE/REGON#ALIFORNIA
BORDER IN THE 0ACIlC .ORTHWESTWEST REGION AND RED
ALDER IS COMMON ON BOTTOM LANDS SHELTERED COVES AND
ON MOIST SLOPES OF THE #OAST AND #ASCADE RANGES %YRE
4HUSTHE0.6TYPEASSOCIATEDWITHTHECLIMAXVEG
ETATION THAT TYPICALLY REPLACES RED ALDER THE CEDAR HEM
LOCK$OUGLASlRTYPEWASUSEDTOMODIFYTHEYIELDTABLE
FORREDALDER
)N THE 0ACIlC .ORTHWESTEAST REGION LODGEPOLE PINE
WAS A FOREST MANAGEMENT TYPE USED IN THE FOREST SECTOR
MODEL(OWEVERTHE0.6TYPELODGEPOLEPINEDIDNOT
DOMINATEANYOFTHEGRIDCELLSWITHINTHISREGION!CCORD
INGTO%YRELODGEPOLEPINEWITHINTHISREGIONWAS
ASSOCIATED WITH SUBALPINE lR %NGELMANN SPRUCE WHITE
SPRUCEAND2OCKY-OUNTAIN$OUGLASlR4HE0.6TYPES
WITH THESE ASSOCIATES lRHEMLOCK $OUGLAS lR WESTERN
SPRUCElRWEREUSEDTODElNETHERESPONSEOFLODGEPOLE
PINEWITHINTHISREGION
4HE ASPENBIRCH TYPE USED AS A FOREST MANAGEMENT
TYPEINTHE.ORTHEASTREGIONISCONSIDEREDBY+àCHLERTO
BEASERALVEGETATIONREPLACEDBY.ORTHEASTERNSPRUCElR
%YRE CONSIDERED ASPENBIRCH TO BE A BOREAL HARD
WOOD!SPENWILLBEREPLACEDBYTHE0.6TYPESOFNORTH
ERN HARDWOODS OR SPRUCElR TYPES AND SUCCESSION TO
THESE TYPES IS MORE RAPID THAN TO PINE %YRE 4HE
VOLUME OF ASPENBIRCH WAS MODIlED BY THE 4%- .00
RESPONSEFROMANAGGREGATIONOF.ORTHEASTERNSPRUCElR
NORTHERN HARDWOODS AND NORTHERN HARDWOODSSPRUCE
0.6TYPES
)NTHE3OUTHERNREGIONPLANTEDPINEANDNATURALPINE
WERETWOFORESTMANAGEMENTTYPESTABLE4HEREWERE
NO CORRESPONDING 0.6 TYPES FOR THESE FOREST TYPES &OR
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
THELAST20!ANALYSISTHEOAKHICKORYPINEANDTHESOUTH
ERN MIXED FOREST 0.6 TYPES WERE USED TO ASSESS THE
IMPACT OF CLIMATE CHANGE ON TIMBER VOLUME OF THE TWO
FORESTMANAGEMENTTYPES
#LEARLYTHEREARELIMITATIONSWITHASSIGNINGFORESTMAN
AGEMENTTYPESTOPOTENTIALNATURALVEGETATIONTYPES%CO
LOGICALMODELSREPRESENTTHEMOSTCURRENTUNDERSTANDING
OFHOWECOLOGICALPROCESSESOPERATEATTHEECOSYSTEMSCALE
4HE POTENTIAL DISTRIBUTION OF THESE ECOSYSTEMS IMPLIES
A SIMILARITY OF ECOSYSTEM FUNCTION WITHIN THE RANGE OF
EACH ECOSYSTEM TYPE AS WELL AS A GEOGRAPHIC PRESENCE
UNALTEREDBYLANDUSE&ORESTMANAGEMENTMODELSHAVE
FOCUSEDONTHEYIELDOFWOODFROMFORESTLAND4HEAGGRE
GATIONOFINVENTORYPLOTSINTOAFORESTMANAGEMENTTYPE
IMPLIES A SIMILARITY IN TIMBER PRODUCTION WITH THE GEO
GRAPHICRANGEOFTHATFORESTMANAGEMENTTYPE&ORBOTH
OFTHESECLASSIlCATIONSYSTEMSTHEECOSYSTEMDYNAMICSOR
TIMBER PRODUCTION WITHIN A TYPE MAY BE QUITE VARIABLE
#URRENT LAND USES HAVE ALTERED POTENTIAL DISTRIBUTIONS
)NVENTORYDATAISMORELIKELYTOREPRESENTTHECURRENTDIS
TRIBUTIONOFFORESTS
5SE OF VEGETATION TYPES PRESENTS PROBLEMS BY POTEN
TIALLYIGNORINGDIFFERENCESBETWEENSPECIESANDNEWASSOCI
ATIONSANDHOWBOTHMAYAFFECTECOSYSTEMSUNDERCLIMATE
CHANGE 4HE ASSUMPTION THAT SPECIES ASSOCIATIONS WILL
REMAINCONSTANTISACONSEQUENCEOFLUMPINGSPECIESSPE
CIlCINFORMATIONINTOAhTYPEv+IRSCHBAUMAND&ISCHLIN
3PECIESASSOCIATIONSHAVEBEENVERYDIFFERENTINTHE
PASTUNDERDIFFERENTCLIMATES$AVISREPORTSPALEO
ECOLOGICAL EVIDENCE OF COMMUNITY TYPES NO LONGER PRES
ENT SUCH AS THE SPRUCEOAK WOODLAND ASSOCIATION 4HE
APPROACH OF USING FUNCTIONAL TYPES (ENDERSON3ELLERS
AND-C'UFlE7OODWARDETALINLIEUOFSPE
CIESINCLIMATECHANGEMODELINGISATTRACTIVEBECAUSETHIS
APPROACH REDUCES THE COMPUTATIONAL COMPLEXITY ASSOCI
ATEDWITHPROJECTINGEACHPLANTSPECIES(OWEVERTHEUSE
OF FUNCTIONAL TYPES OFTEN ASSUMES THAT THESE GROUPINGS
OF SPECIES WILL REMAIN TOGETHER AND RESPOND TO CLIMATE
CHANGE AS A UNIT 4HE USE OF FUNCTIONAL TYPES RAISES THE
ISSUEOFWHETHERhFUNCTIONALTYPESvPRESERVESPECIESDIF
FERENCES3OLBRIG4HEVALUEOFTHESEFUNCTIONALTYPES
IS THAT THEY GROUP SIMILAR PHYSIOLOGICAL AND ECOLOGICAL
ROLES 3OLBRIG 6INTON AND "URKE BUT SPECIES
BEHAVIORALDIFFERENCESMAYBEOVERLOOKED&OREXAMPLEIS
THERATEOFREACHINGEQUILIBRIUMWITHCLIMATETHESAMEFOR
ALLSPECIESWITHINAFUNCTIONALGROUP
3IMILARLY THE AGGREGATION OF DIVERSE TREE SPECIES INTO
FORESTMANAGEMENTTYPESPRESENTSPROBLEMSBYIGNORING
POTENTIALDIFFERENCESBETWEENCOMMERCIALTIMBERSPECIES
ANDHOWCLIMATECHANGEMIGHTALTERWOODlBERPRODUC
TIONFOREXAMPLEHOWVOLUMESMIGHTSHIFTWHENWOOD
DEVELOPSANDTHEQUALITYOFWOODUNDERALTEREDCLIMATE
%XPERIMENTALRESULTSSUGGESTTHATTHERESPONSESTOCLIMATE
CHANGEMIGHTBEGENERAIFNOTSPECIESSPECIlCSEEEARLIER
DISCUSSION
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
7ITHIN THE LAST 20! ANALYSIS THE IMPACT OF CLIMATE
CHANGEON.00WASASSESSEDUSINGTHEHISTORICALRANGEOF
TEMPERATEFORESTSlGNOTTHECURRENTDISTRIBUTIONAS
AFFECTEDBYLANDUSE4HUSTHEECOLOGICALVARIABILITYANA
LYZEDUNDERCLIMATECHANGEREPRESENTEDAGREATERECOLOGI
CALAMPLITUDETHANFORESTSCURRENTLYREPRESENT4HESPATIAL
DISTRIBUTION OF EXISTING FORESTS USING THE FOREST MANAGE
MENT TYPE CLASSIlCATION HAS BEEN RECENTLY MAPPED 7E
USEDTHISSPATIALDISTRIBUTIONTODEVELOPAMAPOFVEGETA
TION CLASSIlED BY THE SAME SYSTEM USED BY THE 6%-!0
MEMBERS4HEFORESTMANAGEMENTTYPEINFORMATIONWAS
AVAILABLE AT THE KM SCALE 7E IDENTIlED A +àCHLER
0.6 TYPE FOR EACH GRID CELL ON THE FOREST MANAGEMENT
MAPBASEDON'ARRISONETALTABLE/NCETHE
+àCHLERTYPEWASASSOCIATEDWITHEACHGRIDCELLWETHEN
USEDTHECLASSIlCATIONGIVENIN6%-!0MEMBERS
TOLINKTHECELLTOA66%'TYPETABLE4HEMAPWAS
THEN RESAMPLED TO THE KM SCALE lG 4HE lNER
SCALEOFTHISMAPALLOWSSMALLERISOLATEDPATCHESOFFOREST
TOREMAINONTHEMAPPARTICULARLYINTHE3OUTHWESTAND
INTHE'REAT0LAINS(OWEVERTHISDISTRIBUTIONCONTRASTS
WITH THE DISTRIBUTION USED IN THE LAST 20!ANALYSIS lG
INTHEDRASTICALLYREDUCEDAREAOFFORESTSINTHEEAST
ERN PART OF THE 5NITED 3TATES AND IN THE PATCHINESS OF
THE FORESTS ACROSS THE 5NITED 3TATES 4HE AREA OF FORESTS
IN THE -IDWEST REGION AND THE -ISSISSIPPI 2IVER VALLEY
DECLINESWHENTHELANDUSEINAGRICULTUREISREMOVED4HE
HOMOGENEITYOFVEGETATIONTYPESISLESSENEDINlGURE
PARTICULARLY FOR THE .EW %NGLAND STATES WHERE BOREAL
TEMPERATE CONIFEROUS AND COOL TEMPERATE MIXED FOREST
TYPESINTERMINGLEINCONTRASTTOTHEUNIFORMITYINlGURE
4HIS REEXAMINATION OF FOREST TYPES lG COULD
BEUSEDASTHEBASISFORANANALYSISOFTHEIMPACTOFCLI
MATECHANGEONFORESTPRODUCTIVITY)TWOULDLIKELYREmECT
THE POTENTIAL SHIFTS IN FOREST PRODUCTIVITY MORE CLOSELY
BECAUSECLIMATESHIFTSINREGIONSOFEXISTINGFORESTSWOULD
BEUSEDASCLIMATEINPUTTOAMODELSUCHAS4%-
0ROJECTING%COSYSTEM
0RODUCTIVITYAT$IFFERENT
3PATIAL3CALES
#LIMATETOPOGRAPHYVEGETATIONANDSOILSINPUTDATA
FORECOLOGICALMODELSUSEDINLARGESCALEINTEGRATEDASSESS
MENTSARETYPICALLYGRIDDEDATTHEDEGREELONGITUDEBY
DEGREELATITUDESCALE#RAMERETAL(EIMANNET
AL+ICKLIGHTERETAL-ELILLOETAL6%-!0
MEMBERS 5SE OF GRIDDED INPUT DATA IMPLICITLY
ASSUMESTHATTHEMEANORDOMINANTSURFACEFEATURESREP
RESENTTHEENTIREGRIDCELL)NHERENTINTHISASSUMPTIONIS
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
*OYCEAND.UNGESSER
THEUNCERTAINTYWITHWHICHTHISGRIDDEDVALUEREPRESENTS
THE HETEROGENEITY OF THE ACTUAL COARSE GRID CELL FEATURES
AND THE REPRESENTATIVENESS OF THIS GRIDDED VALUE WHEN
USED IN THE ECOLOGICAL MODELS TO DESCRIBE THE BIOLOGICAL
PROCESSESOPERATINGWITHINTHEGRIDCELL7HILEOPPORTU
NITIESEXISTTOMOVETHEANALYSISOFTHEIMPACTOFCLIMATE
CHANGE ON FORESTS TO A lNER GRID SCALE THIS REDUCTION IN
GRIDSIZEWOULDINCREASETHEPROCESSINGTIMEBYANORDER
OFMAGNITUDE(ENCEITISIMPORTANTTOASSESSTHEUNCER
TAINTYTHATAGGREGATIONOFCLIMATEDATACONTRIBUTESTOTHE
ESTIMATION OF FOREST PRODUCTIVITY UNDER CLIMATE CHANGE
.UNGESSER ET AL USED THE 4ERRESTRIAL %COSYSTEM
-ODEL4%-VERSION-C'UIREETALATOEVALUATE
THEUTILITYOFMOVINGTHECLIMATECHANGEIMPACTANALYSIS
FROMTHEDEGREESCALEUSEDINTHELAST20!ANALYSISTO
THEKMSCALE
4HEEFFECTOFAGGREGATIONONTHEESTIMATIONOFPRODUCTIV
ITYHASBEENSTUDIED.ETPRIMARYPRODUCTION.00ESTI
MATESDIFFEREDBYPERCENTWHENCOARSEGRAINVERSUSlNE
GRAINSOILSDATAWEREUSEDASINPUTDATAFORTHE0N%TMODEL
,ATHROPETAL0IERCEAND2UNNINGOBTAINED
OVERESTIMATESOFUPTOPERCENTIN.00FROMTHE&/2%34
"'#MODELWHENSUBGRIDVARIATIONSINCLIMATETOPOGRA
PHYSOILSANDVEGETATIONWEREAVERAGEDACROSSASERIESOF
GRAINSIZESFROMKMTODEGREE-OSTOFTHISERRORWAS
PRODUCEDBYAVERAGETEMPERATUREWHILEAVERAGETOPOGRA
PHYSOILSANDVEGETATIONTYPESALSOCONTRIBUTED
.UNGESSER ET AL EXAMINED THE IMPACT OF TWO
DIFFERENTSPATIALRESOLUTIONSONTHESIMULATEDFORESTECO
SYSTEMRESPONSESFORABASELINECLIMATEANDTWOCLIMATE
CHANGESCENARIOS4HE4%-MODELUSESSPATIALLYRESOLVED
INFORMATION ON CLIMATE MONTHLY PRECIPITATION MONTHLY
MEANAIRTEMPERATUREANDCLOUDINESSSOILTEXTUREPER
CENTSANDSILTANDCLAYVEGETATIONTYPEANDELEVATION
4HElNERESOLUTIONGRIDCELLSWEREKMINSIZEAND
OFTHESECELLSWERENESTEDWITHINACOARSERESOLUTIONGRID
CELLOFAPPROXIMATELYAHALFDEGREEINSIZEKM4HE
KMsKMRASTERDATAFORCLIMATEMONTHLYPRECIPITA
TIONMONTHLYMEANAIRTEMPERATUREWEREOBTAINEDFROM
.EILSON PERSONAL COMMUNICATION AS DESCRIBED IN $ALY
ETAL-ARKSAND.EILSON&ORTHE
KMsKMINPUTDATATHElNESCALEINPUTDATASETSFOR
CONTINUOUS VARIABLES WERE AVERAGED TO THE KM s KMRESOLUTION!VERAGINGTOTHEKMSCALERESULTSINTHE
SMOOTHING OUT OF PRECIPITATION AND TEMPERATURE VALUES
ALONGTHEGRADIENTSOFCHANGEANDTHELOSSOFlNERDETAIL
INSOMEAREASlGAND!REASOFlNESCALEPATCHI
NESS OF PRECIPITATION IN THE WESTERN MOUNTAINS AND THE
SOUTHERN COASTAL PLAINS ARE SMOOTHED OUT AT THE KM
GRID SCALE lG !NNUAL AVERAGE TEMPERATURE VALUES
AREINmUENCEDINAREASWHERETHEREISSUBSTANTIALTEMPER
ATUREVARIABILITYSUCHASAROUNDTHEMOUNTAINOUSAREASIN
THE7ESTlG
4HEHISTORICALRANGEOFTEMPERATEFORESTSWASTHESPA
TIAL EXTENT OF THE .UNGESSER STUDY +àCHLER VEGETATION
Ecosystem Productivity and the Impact of Climate Change
Figure 3.1—Forest type distribution used in the last RPA analysis (Joyce et al. 1995).
Joyce and Nungesser
56
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
Figure 3.2—VEMAP vegetation classification associated with the current distribution of forests based on USDA Forest Service inventory plots.
Ecosystem Productivity and the Impact of Climate Change
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
Joyce and Nungesser
57
Joyce and Nungesser
Ecosystem Productivity and the Impact of Climate Change
Figure 3.3—Baseline annual precipitation at 50 km scale and 10 km scales.
58
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
Ecosystem Productivity and the Impact of Climate Change
Joyce and Nungesser
Figure 3.4—Baseline annual average temperature at 50 km and 10 km scales.
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
59
*OYCEAND.UNGESSER
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
WASDIGITIZEDFROMTHEMAP+àCHLERATKM
RESOLUTION 3TEVE (ODGE PERSONAL COMMUNICATION AND
WASRECLASSIlEDTO4%-VEGETATIONTYPES&ORTHEKM
sKMINPUTDATATHElNESCALEINPUTDATASETFORVEG
ETATIONWEREAGGREGATEDTOTHEKMsKMRESOLUTION
THROUGHTHEUSEOFTHEMAJORITYRULEPROPORTIONALAGGRE
GATION METHOD OF #OSTANZA AND -AXWELL 7HILE
THE COARSE RESOLUTION GRIDS INCLUDE SIX VEGETATION TYPES
lGASEVENTHTYPETEMPERATEEVERGREENBROADLEAF
APPEAREDINONLYTHREEGRIDCELLSATTHElNERESOLUTION!LL
FORESTTYPESTHATCOMPRISEDGREATERTHANPERCENTOFTHE
TOTALAREAWITHINTHEKMGRIDSRETAINEDATLEASTPER
CENTOFTHEIRTOTALAREAWHENAGGREGATEDTOTHEKMGRID
SIZE4HERARERFORESTTYPESBOREALWETBOREALTEMPERATE
FOREST WETLAND AND TEMPERATE BROADLEAF EVERGREEN LOST
FROM TO PERCENT OF THEIR AREA UNDER THE PROPOR
TIONAL AGGREGATION RULE 2ARE TYPES DISPERSED ACROSS THE
LANDSCAPEWEREMOSTLIKELYTOBELOSTINTHEAGGREGATION
&OREXAMPLETHEBOREALGRIDCELLSIN.EW%NGLANDDISAP
PEAR AT THE KM SCALE lG 3IMILARLY SOME OF THE
FORESTEDBOREALWETLANDFORESTSINTHENORTHERN-IDWEST
ARELOSTATTHEKMSCALE
!NAGGREGATIONERRORWASCOMPUTEDASTHEDIFFERENCE
OFTHEKMESTIMATEOF.00ANDTHEKMESTIMATEOF
.00DIVIDEDBYTHEKMESTIMATEOF.004HISAGGREGA
TION ERROR WAS COMPUTED FOR THE BASELINE RUNS BASELINE
CLIMATEAND#/LEVELSOFMEQL
4ABLEˆ.ETPRIMARYPRODUCTION.00FORBASELINECLIMATE
6ALUESARENETPRIMARYPRODUCTIONING#MREPORTEDVALUESARE
MEANSANDSTANDARDDEVIATIONSAREINPARENTHESESFOLLOWEDBY
RANGES4HEhNvISTHENUMBEROFCOARSERESOLUTIONGRIDCELLSIN
EACHFORESTTYPE
&OREST4YPEN
2ESOLUTION
.00
"OREAL
&INE
#OARSE
n
n
&ORESTEDBOREAL
WETLAND
&INE
#OARSE
n
n
-IXEDTEMPERATE
&INE
#OARSE
n
n
#ONIFERS
&INE
#OARSE
n
n
$ECIDUOUS
&INE
#OARSE
n
n
4EMPERATEFORESTED
WETLAND
&INE
#OARSE
n
n
!,,&/2%343
&INE
#OARSE
n
n
INBOREALANDFORESTEDBOREALWETLANDFORESTSWITHPOSI
TIVEVALUESOFANDPERCENTRESPECTIVELY%STIMATES
EB = 100 ∗ NPP10 − NPP50 / NPP10 OF .00 FOR BOREAL FOREST AT THE KM GRID SCALE WERE
LESSTHANTHEKMESTIMATEBYAPPROXIMATELYG#M
d
=
100
∗
NPP
(
WHERE %" IS THE RELATIVE AGGREGATION ERROR AND GQ50 IS
TABLE
GQ50 − NPP50 ) / NPP50
THE AVERAGE OF THE MEAN ANNUAL .00IN G#M YR ESTI
4HE CLIMATE CHANGE SCENARIOS TEMPERATURE PRECIPI
MATESFORTHElNEGRIDCELLSAND.00ISTHEESTIMATE
TATION AT THE KM SCALE WERE BASED ON TWO '#-S
OFTHE.00ATTHECOARSEGRIDSCALE
USED IN THE LAST 20! ANALYSIS THE 'EOPHYSICAL &LUID
4HE .00RESULTS FOR THE KM REmECT lNER SCALE PAT
$YNAMICS ,ABORATORY 1mUX '&$,1 -ANABE AND
TERNSTHANTHEKMRESULTSlGBUTTHESEPATTERNS
7ETHERALD AND /REGON 3TATE 5NIVERSITY /35
ARENOTSUFlCIENTTOGENERATELARGEAGGREGATIONERRORSAT
MODELS3CHLESINGERAND:HAO5SINGTHESAMEPRO
THENATIONALFORESTTYPEORGRIDCELLSCALE4HE.00ESTI
TOCOLASFORBASELINECLIMATElNERESOLUTIONCLIMATEINPUT
MATEFORALLFORESTSATTHENATIONALEXTENTDIFFEREDBYLESS
DATAWEREAVERAGEDWITHINACOARSEGRIDTOSERVEASCOARSE
THAN G#M BETWEEN THE lNE AND COARSE RESOLUTION
RESOLUTION INPUTS TO 4%- 6ALUES FOR FOREST TYPE ELEVA
TIONANDSOILSREMAINUNCHANGEDFROMTHEBASELINESIM
SCALESVERSUSG#MTABLE!GGREGATION
ULATION 4HE CLIMATE CHANGE SCENARIOS INCLUDED A #/
ERRORBASEDONTHESEGRIDCELLSISVERYSMALLANDNEGA
TIVEn%STIMATESOF.00ATTHEKMGRIDSCALEDIF
CONCENTRATIONOFPPMV!GGREGATIONERRORIN.00FOR
FEREDFROMTHECORRESPONDINGAVERAGEFORTHEKMGRID
THE TWO CLIMATE CHANGE SCENARIOS WAS COMPUTED IN THE
CELLS BY LESS THAN PERCENT ACROSS MOST OF THE HISTORIC
SAMEMANNERASTHEBASELINEAGGREGATIONERROR
RANGEOFTEMPERATEFORESTS4HESMALLESTAGGREGATIONERROR
'RIDLEVELRESPONSEOFNETPRIMARYPRODUCTIVITYTOTHE
WASFOUNDGENERALLYTHROUGHOUTTHE%ASTAND3OUTHEAST
CLIMATECHANGESCENARIOSWERECALCULATEDAS
ASWELLASINTHEWESTERNMOUNTAINS2ARELYWEREAGGRE
d OS10 = 100 ∗ NPPOS10 − NPP10 / NPP10 , GATION ERROR DIFFERENCES GREATER THAN PERCENT AT THE
INDIVIDUALGRIDCELLLEVEL4HESELARGERAGGREGATIONERRORS
d OS50 = 100 ∗ ( NPPOS50 − NPP50 ) / NPP50 , OCCURREDAROUNDTHE'REAT,AKESINNORTHERN.EW%NG
LAND AND IN THE 2OCKY -OUNTAINS "Y FOREST TYPE THE
d GQ10 = 100 ∗ NPPGQ10 − NPP10 / NPP10 , and AGGREGATION ERRORS WERE STILL SMALL LESS THAN PERCENT
%STIMATESOF.00DIFFEREDBYLESSTHANG#MINMOST
d GQ50 = 100 ∗ ( NPPGQ50 − NPP50 ) / NPP50 CASES TABLE 4HE LARGEST AGGREGATION ERROR OCCURRED
(
)
(
)
(
)
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
Ecosystem Productivity and the Impact of Climate Change
Joyce and Nungesser
Figure 3.5—Vegetation types mapped at 50 km and 10 km scales.
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
61
Joyce and Nungesser
Ecosystem Productivity and the Impact of Climate Change
Figure 3.6—Baseline net primary productivity (NPP) at 50 km and 10 km scales.
62
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
WHERE D/3 D/3 D'1 AND D'1 ARE .00 RESPONSES
FOREACHlNED?ANDCOARSED?RESOLUTIONGRIDTOTHE
/35ANDTHE'&$,1CLIMATESCENARIOS4HISMETRICISTHE
DELTATHATISPASSEDTOTHEFORESTSECTORMODELANDUSEDTO
MODIFYTIMBERVOLUMEINCREASESINTHEINVENTORYMODEL
.ETPRIMARYPRODUCTIVITYFORALLFORESTSINCREASEDUNDER
BOTHCLIMATECHANGESCENARIOSATBOTHRESOLUTIONS5NDER
THE/35CLIMATE.00OFALLFORESTSINCREASEDAPPROXIMATELY
PERCENT ABOVE THE BASELINE .00 RESPONSE WHEREAS
UNDER THE '&$,1 CLIMATE .00 OF ALL FORESTS INCREASED
LESSTHANPERCENTlG7ITHINEACHCLIMATECHANGE
SCENARIOTHERESPONSETOCLIMATECHANGEATTHECOARSERES
OLUTIONDIFFEREDBYLESSTHANPERCENTFROMTHERESPONSE
ATTHElNERRESOLUTION!GGREGATIONERRORFORALLFORESTSFOR
THE/35CLIMATEnPERCENTWASSIMILARTOTHEERRORFOR
THE'&$,1CLIMATEnPERCENTANDBOTHRESULTSWERE
SIMILARTOTHEAGGREGATIONERROROFTHEBASELINECLIMATEn
PERCENT 4HE SPATIAL PATTERNS OF THESE AGGREGATION ERRORS
WERE SIMILAR BETWEEN THE TWO CLIMATE SCENARIOS AND THE
BASELINEACROSSTHEHISTORICRANGEOFTEMPERATEFORESTS
7ITHINFORESTTYPES.00INCREASEDFROMTOPER
CENT VARYING BY CLIMATE SCENARIO 4HE .00RESPONSE FOR
BOREAL FORESTS AND FORESTED BOREAL WETLANDS WAS TO PERCENT GREATER UNDER THE '&$,1 CLIMATE THAN UNDER
THE/35CLIMATE(OWEVERFORMIXEDTEMPERATEDECID
UOUS AND TEMPERATE FORESTED WETLAND .00 UNDER THE
/35 CLIMATE WAS TO PERCENT GREATER THAN UNDER
THE'&$,1CLIMATE7ITHINEACHCLIMATESCENARIOFOREST
.00RESPONSESATTHECOARSESCALEDIFFEREDBYLESSTHAN
PERCENTFROMTHE.00RESPONSETOCLIMATECHANGEATTHE
lNERRESOLUTIONFORALLFORESTTYPESEXCEPTCONIFERWHICH
DIFFERED BY LESS THAN PERCENT 4HE ERROR IN ALL THREE
CLIMATE SCENARIOS WAS HIGHEST IN THE BOREAL FORESTS AND
FORESTED BOREAL WETLANDS AND PERCENT RESPEC
TIVELY 4HE ABSOLUTE VALUE OF THE AGGREGATION ERROR FOR
THE OTHER FORESTS WAS LESS THAN PERCENT 'EOGRAPHI
CALLYAGGREGATIONERRORFORBOTH/35AND'&$,1ISCON
CENTRATED IN THE SAME AREAS AS THAT OF BASELINE CLIMATE
AROUND THE 'REAT ,AKES IN .EW %NGLAND AND IN THE
2OCKY -OUNTAINS 4HE SMALLEST AGGREGATION ERROR LESS
THAN PERCENT WAS FOUND IN THE 3OUTH AND THE -ID
!TLANTICANAREATHATHADTHEGREATESTDIFFERENCESINTHE
.00RESPONSETOCLIMATECHANGEnTOPERCENT
2ELATIVETOTHEBASELINETHEPERCENTINCREASESORDECREASES
IN .00ARE SIMILAR ACROSS THE lNE AND COARSE RESOLUTIONS
WITHIN A CLIMATE CHANGE SCENARIO BUT DIFFER SIGNIlCANTLY
ACROSS SCENARIOS lG 0ERCENT INCREASES IN .00 WERE
SIMILARUNDERBOTHCLIMATESCENARIOSINTHE7ESTBUTTHE
RESPONSESINTHE3OUTHAND-ID!TLANTICFORESTSWEREDRA
MATICALLYDIFFERENT&ORTHE'&$,1CLIMATETHESOUTHERN
ANDMID!TLANTICFOREST.00DECLINEDUPTOPERCENTREL
ATIVETOTHEBASELINE.00WHEREASUNDERTHE/35CLIMATE
THESEFORESTSINCREASEDIN.00FROMTOPERCENT
4HE KM GRIDCELL RESOLUTION IS MOST RELEVANT TO
STANDLEVEL FOREST MANAGERS !GGREGATION ERROR AT THIS
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
*OYCEAND.UNGESSER
SCALE IS LESS THAN PERCENT TWO STANDARD DEVIATIONS
WHICHISWITHINTHEPERCENTPRECISIONTHATSTANDLEVEL
.00ISMEASURED/URANALYSESINDICATETHATAGGREGATION
ERROR IS THE LARGEST IN TRANSITION REGIONS AND IN REGIONS
WITHSUBSTANTIALVARIABILITYINAIRTEMPERATURE!GGREGA
TIONERRORISPRIMARILYASSOCIATEDWITHTHEREPRESENTATION
OF A MOSAIC OF FOREST TYPES WITH A SINGLE FOREST TYPE AT
KM RESOLUTION 4HIS SOURCE OF AGGREGATION ERROR CAN
EASILY BE MINIMIZED BY MAKING .00ESTIMATES FOR EACH
FORESTTYPEWITHINAKMGRIDCELLANDAGGREGATINGESTI
MATESBASEDONTHEPROPORTIONOFEACHFORESTTYPEWITHIN
THEGRIDCELL4HISAPPROACHHASBEENUSEDBY"ONAN
AS A MEANS OF REPRESENTING VEGETATIVE HETEROGENEITY FOR
ESTIMATINGCARBONWATERANDENERGYEXCHANGEINTHESUR
FACEBOUNDARYLAYEROFGENERALCIRCULATIONMODELS0IERCE
AND2UNNINGALSOFOUNDTHATAVERAGINGTEMPERA
TURESUBSTANTIALLYINmUENCEDAGGREGATIONERRORINREGIONS
OFCOMPLEXTERRAIN)TMAYBEPOSSIBLETOACHIEVECOMPUTA
TIONALEFlCIENCYATKMRESOLUTIONBYAGGREGATINGTEM
PERATUREFORALIMITEDNUMBEROFELEVATIONBANDSMAKING
.00ESTIMATESFOREACHELEVATIONBANDANDAGGREGATING
ESTIMATESBASEDONTHEPROPORTIONOFEACHELEVATIONBAND
WITHINTHEGRIDCELL.UNGESSERETAL
4HERESOLUTIONOFFORESTTYPESISMOSTRELEVANTTOCOUN
TRYSPECIlC ECONOMIC ASSESSMENTS OF THE IMPACTS OF CLI
MATE CHANGE ON TIMBER RESOURCES &OR EXAMPLE RELATIVE
CLIMATICRESPONSESOF.00FORDIFFERENTFORESTTYPESINDIF
FERENTREGIONSWEREUSEDASINPUTSFORTHELAST20!*OYCE
ET AL %XCEPT FOR BOREAL FORESTS AND FORESTED BOREAL
WETLANDSTHEMEANAGGREGATIONERROROFBASELINE.00ESTI
MATESISLESSTHANPERCENTFOREACHFORESTTYPE)NCONTRAST
AGGREGATION ERROR FOR BOREAL FORESTS AND FORESTED BOREAL
WETLANDSISAPPROXIMATELYPERCENT4HISLEVELOFERRORIS
CAUSEDPRIMARILYBYTHEOVERREPRESENTATIONOFBOREALAND
BOREALWETLANDFORESTSANDUNDERREPRESENTATIONOFMORE
HIGHLYPRODUCTIVEFORESTSINTHEKMSIMULATION
)NCOMPARISONTOTHEBASELINESIMULATIONSMEANAGGRE
GATIONERRORFORTHEABSOLUTEESTIMATESOFEACHFORESTTYPE
INTHECLIMATECHANGESIMULATIONSISSIMILAREXCEPTFORTEM
PERATE CONIFER FOREST n PERCENT FOR /35 AND n PER
CENTFOR'&$,VSnPERCENTFORBASELINECLIMATE3IMILAR
TOAGGREGATIONERRORTHERELATIVERESPONSESOF.00ATEACH
RESOLUTIONARESIMILAREXCEPTFORCONIFERFORESTSAND
PERCENT LOWER RESPONSE FOR THE lNE RESOLUTION /35 AND
'&$,SIMULATIONS 4HE NEGATIVE AGGREGATION ERRORS AND
LOWER RESPONSES FOR TEMPERATE CONIFER FORESTS ARE ASSOCI
ATED WITH THE AVERAGING OF TEMPERATURE IN THE COMPLEX
TERRAIN IN THE NORTHERN 2OCKY -OUNTAINS AND IN WESTERN
7ASHINGTON /REGON AND #ALIFORNIA "ECAUSE DIFFERENCES
BETWEENTHERESPONSESOF.00ATDIFFERENTRESOLUTIONSARE
SMALLCOMPAREDWITHTHERESPONSESTODIFFERENTCLIMATESCE
NARIOSTHEYCOULDBEIGNOREDINIMPACTASSESSMENTSTHAT
EVALUATESENSITIVITIESTODIFFERENTCLIMATECHANGESCENARIOS
4HENATIONALRESOLUTIONISMOSTRELEVANTTOGLOBALECO
NOMIC ASSESSMENTS OF THE IMPACTS OF CLIMATE CHANGE ON
Joyce and Nungesser
Ecosystem Productivity and the Impact of Climate Change
Figure 3.7—Net primary productivity at 50 km and 10 km scales from the GFDL-Q climate scenario.
64
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
Joyce and Nungesser
Figure 3.8—Forest responses to two climate change scenarios (OSU, GFDL-Q) at 10 km (fine grid) and 50 km (coarse grid) scales.
Ecosystem Productivity and the Impact of Climate Change
USDA Forest Service Gen. Tech. Rep. RMRS–GTR–59. 2000.
65
*OYCEAND.UNGESSER
TIMBERRESOURCES4HERELATIVECLIMATICRESPONSESOF.00
OF HARDWOODS AND SOFTWOODS FOR DIFFERENT REGIONS WERE
USED AS INPUTS FOR A NATIONAL ASSESSMENT 0EREZ'ARCIA
ET AL &OR THE CONTERMINOUS 5NITED 3TATES AGGRE
GATION ERROR WAS PERCENT IN THE BASELINE SIMULATION
PERCENTINTHE/35SIMULATIONANDPERCENTINTHE
'&$,SIMULATION2ESPONSESOF.00ACROSSTHETWOSPA
TIALSCALESWEREANDPERCENTLOWERFORTHE/35AND
'&$,SIMULATIONS"ECAUSETHESEDIFFERENCESAREMINUS
CULE IN COMPARISON TO THE NATIONAL .00RESPONSES THEY
CANBEIGNOREDINIMPACTASSESSMENTSTHATEVALUATESENSI
TIVITIESTODIFFERENTCLIMATECHANGESCENARIOS
4HESE RESULTS INDICATE THAT .00 RESPONSES OF 4%-
TO PROJECTED CLIMATE CHANGE ARE INSENSITIVE TO THE RESO
LUTION OF INPUTS BUT THAT AGGREGATION ERROR OF ABSOLUTE
.00ESTIMATESISSENSITIVETOTHERESOLUTIONOFINPUTSFOR
SOME SITUATIONS %XCEPT FOR TRANSITION AREAS AND REGIONS
WITH SUBSTANTIAL TEMPERATURE VARIABILITY THESE SIMULA
TIONSINDICATETHATTHEUSEOFORESOLUTIONPROVIDESAN
ACCEPTABLELEVELOFAGGREGATIONERRORATTHETHREESCALESOF
ANALYSISINTHISSTUDY
)TISIMPORTANTTORECOGNIZETHATTHECONCLUSIONSINTHIS
STUDYAREBASEDONTWORESOLUTIONSANDONEBIOGEOCHEM
ISTRY MODEL 0IERCE AND 2UNNING USED A DIFFERENT
BIOGEOCHEMISTRYMODELTOSIMULATE.00FORVARIOUSRES
OLUTIONS RANGING FROM KM TO KM IN A REGION OF
COMPLEX TOPOGRAPHY !T THE COARSEST SCALE THEY FOUND
COARSERESOLUTION .00 WAS OVERESTIMATED BY UP TO PERCENTRELATIVETO.00ESTIMATESATTHElNESTRESOLUTION
4HERESULTSOF.UNGESSERETALQUALITATIVELYAGREE
WITHTHOSEOF0IERCEAND2UNNING
"ECAUSEMOSTLARGESCALEBIOGEOCHEMISTRYMODELSARE
PARAMETERIZED WITH STANDLEVEL DATA A SYSTEMATIC ANAL
YSIS OF AGGREGATION ERROR WITH SEVERAL BIOGEOCHEMISTRY
MODELS ACROSS A RANGE OF SPATIAL RESOLUTIONS FROM STAND
TO EG M TO KM TO KM TO KM
SHOULDBEUNDERTAKENINDIFFERENTFORESTREGIONSTODETER
MINE WHETHER OUR CONCLUSIONS AND THOSE OF 0IERCE AND
2UNNINGAREROBUST
&INALLY IT IS IMPORTANT TO VERIFY THE CONCLUSION ABOUT
THE INSENSITIVITY OF .00 RESPONSES TO THE RESOLUTION OF
INPUTSWITHOTHERBIOGEOCHEMISTRYMODELS"YCLARIFYING
THE SCALING ISSUES ASSOCIATED WITH .00 ESTIMATES AND
RESPONSESTHESESUGGESTEDSTUDIESWOULDIMPROVEIMPACT
ASSESSMENTSTHATRELYONTHEESTIMATESOFLARGESCALEECO
LOGICALMODELS
#ONCLUSIONS
!NALYSES OF THE IMPACT OF CLIMATE CHANGE ON FOREST
PRODUCTIVITYBASEDONEXPERIMENTALRESEARCHANDMODEL
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
INGWOULDSUGGESTTHATFORESTPRODUCTIVITYMAYINCREASE
UNDER ELEVATED CARBON DIOXIDE BUT THAT THE LOCAL CON
DITIONS OF MOISTURE STRESS AND NUTRIENT AVAILABILITY WILL
STRONGLYTEMPERANYRESPONSE0ROJECTEDINCREASESINPRO
DUCTIVITY FROM CARBON DIOXIDE FERTILIZATION APPEAR TO BE
WITHINTHESAMEMAGNITUDEASPOTENTIALINCREASESINPRO
DUCTIVITY FROM TIMBER MANAGEMENT TREATMENTS 2ElNE
MENTS IN THE ANALYSIS SUCH AS ANALYZING THE IMPACT AT A
lNERSCALEDONOTAPPEARTOALTERTHERESULTSFROMTHELAST
20!ANALYSIS)NCORPORATINGLANDUSECHANGESAPPEARSTO
BEACRITICALNEXTSTEPINTHEANALYSISOFTHEIMPACTOFCLI
MATECHANGEONFORESTPRODUCTIVITY
!CKNOWLEDGMENTS
7EWOULDLIKETORECOGNIZE$AVE-C'UIREFORHISCON
TRIBUTIONS TO THE AGGREGATION RESEARCH DESCRIBED IN THIS
CHAPTER3COTT0OWELLAND!LICIA,IZARRAGAPROVIDED')3
SUPPORTAND-ONICA%NGELPROVIDEDPROGRAMMINGSUP
PORT+AREN&ASSNACHTASSISTEDWITHTHELITERATUREREVIEW
7E GREATLY APPRECIATE THE EDITORIAL REVIEW BY 2OBERT
(AMREANDTHESTATISTICALREVIEWOF2UDY+ING7EALSO
THANK$OMINIQUE"ACHELETFORHERHELPFULREVIEW
4HISWORKWASSUPPORTEDBYTHE53$!&OREST3ERVICE
2ESOURCES0ROGRAMAND!SSESSMENTSTAFFANDTHE.ORTH
EASTERN'LOBAL#HANGE0ROGRAM
,ITERATURE#ITED
!TKINSON#*4AYLOR*-7ILKINS$;ETAL=%FFECTSOFELEVATED
#/ ON CHLOROPLAST COMPONENTS GAS EXCHANGE AND GROWTH OF OAK
ANDCHERRY4REE0HYSIOLOGYn
"ERNTSON'-"AZZAZ&!4HEALLOMETRYOFROOTPRODUCTIONAND
LOSS IN SEEDLINGS OF !CER RUBRUM !CERACEAE AND "ETULA PAPYRIFERA
"ETULACEAEIMPLICATIONSFORROOTDYNAMICSINELEVATED#/!MERI
CAN*OURNALOF"OTANYn
"IRDSEY2ICHARD!0LANTINGA!NDREW*(EATH,INDA30ASTAND
PROSPECTIVECARBONSTORAGEIN5NITED3TATESFORESTS&OREST%COLOGY
AND-ANAGEMENTn
"IRDSEY2!(EATH,3#ARBONCHANGESIN53&ORESTS)N*OYCE
,!ED#LIMATECHANGEANDTHEPRODUCTIVITYOF!MERICASFORESTS
'EN4ECH2EP2-&ORT#OLLINS#/53$EPARTMENTOF!GRI
CULTURE &OREST 3ERVICE 2OCKY -OUNTAIN &OREST AND 2ANGE %XPERI
MENT3TATIONn
"ONAN'",ANDATMOSPHERE#/EXCHANGESIMULATEDBYALAND
SURFACEPROCESSMODELCOUPLEDTOANATMOSPHERICGENERALCIRCULATION
MODEL*OURNALOF'EOPHYSICAL2ESEARCHn
#EULEMANS23HAO"9*IANG8.;ETAL=&IRSTANDSECOND
YEARABOVEGROUNDGROWTHANDPRODUCTIVITYOFTWO0OPULUSHYBRIDS
GROWNATAMBIENTANDELEVATED#/4REE0HYSIOLOGYn
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
#EULEMANS 2 -OUSSEAU - 4ANSLEY 2EVIEW .O %FFECTS OF
ELEVATEDATMOSPHERIC#/ONWOODYPLANTS.EW0HYTOLOGIST
n
#OSTANZA 2 -AXWELL 4 2ESOLUTION AND PREDICTABILITY AN
APPROACHTOTHESCALINGPROBLEM,ANDSCAPE%COLOGYn
#RAMER 7 +ICKLIGHTER $7 "ONDEAU ! ;ET AL= #OMPARING
GLOBAL MODELS OF TERRESTRIAL NET PRIMARY PRODUCTIVITY .00 OVER
VIEW AND KEY RESULTS 'LOBAL #HANGE "IOLOGY 3UPPLEMENT n
#URTIS036OGEL#30REGITZER+3;ETAL=)NTERACTINGEFFECTS
OFSOILFERTILITYANDATMOSPHERIC#/ONLEAFAREAGROWTHANDCARBON
GAINPHYSIOLOGYOF0OPULUSXEURAMERICANA$ODE'UINIER.EW0HY
TOLOGISTn
#URTIS037ANG8!METAANALYSISOFELEVATED#/ EFFECTSON
WOODYPLANTMASSFORMANDPHYSIOLOGY/ECOLOGIAn
$ALY # .EILSON 20 0HILLIPS $, ! STATISTICALTOPOGRAPHIC
MODEL FOR MAPPING CLIMATOLOGICAL PRECIPITATION OVER MOUNTAINOUS
TERRAIN*OURNALOF!PPLIED-ETEOROLOGYn
$AVIS - ,AGS IN VEGETATION RESPONSE TO GREENHOUSE WARMING
#LIMATE#HANGEn
$E,UCIA %( #ALLAWAY 2- 3CHLESINGER 7( /FFSETTING
CHANGESINBIOMASSALLOCATIONANDPHOTOSYNTHESISINPONDEROSAPINE
0INUSPONDEROSAINRESPONSETOCLIMATECHANGE4REE0HYSIOLOGY
n
%YRE&(ED&ORESTCOVERTYPESOFTHE5NITED3TATESAND#ANADA
7ASHINGTON$#3OCIETYOF!MERICAN&ORESTERS
'ARRISON'!"JUGSTAD!*$UNCAN$!,EWIS-%3MITH$
26EGETATIONANDENVIRONMENTALFEATURESOFFORESTANDRANGE
ECOSYSTEMS 53$! &OREST 3ERVICE !GRICULTURE (ANDBOOK .O 7ASHINGTON$#53'OVERNMENT0RINTING/FlCE
'ORISSEN ! +UIKMAN 0* 6AN $E "EEK ( #ARBON ALLOCATION
ANDWATERUSEINJUVENILE$OUGLASlRUNDERELEVATED#/.EW0HY
TOLOGISTn
'RIFlN +, 7INNER 7% 3TRAIN "2 'ROWTH AND DRY MATTER
PARTITIONINGINLOBLOLLYANDPONDEROSAPINESEEDLINGSINRESPONSETO
CARBONANDNITROGENAVAILABILITY.EW0HYTOLOGISTn
(EIMANN-%SSER'(AZELTINE!;ETAL=%VALUATIONOFTERRES
TRIALCARBONCYCLEMODELSTHROUGHSIMULATIONSOFTHESEASONALCYCLE
OFATMOSPHERIC#/&IRSTRESULTSOFAMODELINTERCOMPARISONSTUDY
'LOBAL"IOGEOCHEMICAL#YCLESn
(ENDERSON3ELLERS ! -C'UFlE + 'LOBAL CLIMATE MODELS AND
@DYNAMICVEGETATIONCHANGES'LOBAL#HANGE"IOLOGYn
(IBBS $% #HAN 33 #ASTELLANO - ;ET AL= 2ESPONSE OF RED
ALDERSEEDLINGSTO#/ENRICHMENTANDWATERSTRESS.EW0HYTOLO
GISTn
)NEICHEN+7IEMKEN67IEMKEN!3HOOTSROOTSANDECTOMY
CORRHIZAFORMATIONOFPINESEEDLINGSATELEVATEDATMOSPHERICCARBON
DIOXIDE0LANT#ELLAND%NVIRONMENTn
*OYCE ,! ED 0RODUCTIVITY OF !MERICAS &ORESTS AND #LIMATE
#HANGE'EN4ECH2EP2-&ORT#OLLINS#/53$EPARTMENT
OF !GRICULTURE &OREST 3ERVICE 2OCKY -OUNTAIN &OREST AND 2ANGE
%XPERIMENT3TATIONP
*OYCE ,! -ILLS *2 (EATH ,3 ;ET AL= &OREST SECTOR IMPACTS
FROMCHANGESINFORESTPRODUCTIVITYUNDERCLIMATECHANGE*OURNALOF
"IOGEOGRAPHYn
*OYCE ,! "IRDSEY 2 -ILLS * (EATH , 0ROGRESS TOWARD AN
INTEGRATED MODEL OF THE EFFECTS OF GLOBAL CHANGE ON 5NITED 3TATES
FORESTS )N "IRDSEY 2 -ICKLER 2 3ANDBERG $ ;ET AL= EDS 53$!&OREST3ERVICE'LOBAL#HANGE2ESEARCH0ROGRAM(IGHLIGHTS
n'EN4ECH2EP.%2ADNOR0!53$EPARTMENTOF
!GRICULTURE&OREST3ERVICE.ORTHEASTERN&OREST%XPERIMENT3TATION
n
+ICKLIGHTER$7"ONDEAU!3CHLOSS!,;ETAL=#OMPARING
GLOBALMODELSOFTERRESTRIALNETPRIMARYPRODUCTIVITY.00'LOBAL
PATTERNANDDIFFERENTIATIONBYMAJORBIOMES'LOBAL#HANGE"IOLOGY
3UPPLEMENTn
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
*OYCEAND.UNGESSER
+ING*34HOMAS2"3TRAIN"2'ROWTHANDCARBONACCUMULA
TIONINROOTSYSTEMSOF0INUSTAEDAAND0INUSPONDEROSASEEDLINGSAS
AFFECTEDBYVARYING#/TEMPERATUREANDNITROGEN4REE0HYSIOLOGY
n
+IRSCHBAUM-5&&ISCHLIN!EDS#LIMATECHANGEIMPACTSON
FORESTS)N7ATSON24:INYOWERA-#-OSS2($OKKEN$*
EDS#LIMATE#HANGE)MPACTS!DAPTATIONSAND-ITIGATIONOF
#LIMATE #HANGE 3CIENTIlC4ECHNICAL !NALYSES )NTERGOVERNMENTAL
0ANEL ON #LIMATE #HANGE #AMBRIDGE 5+ #AMBRIDGE 5NIVERSITY
0RESSn#HAPTER
+àCHLER!70OTENTIALNATURALVEGETATIONOFCONTERMINOUS5NITED
3TATES!MERICAN'EOGRAPHIC3OCIETY3PEC0UBLICATION.O.EW
9ORKP
+àCHLER!7 0OTENTIAL NATURAL VEGETATION OF THE 5NITED 3TATES
NDED.EW9ORK!MERICAN'EOGRAPHIC3OCIETY
+àCHLER!70OTENTIALNATURALVEGETATIONOFTHE5NITED3TATES
7ASHINGTON$#53$!&OREST3ERVICE2!2%))-AP"
,ATHROP2'*R!BER*$"OGNAR*!3PATIALVARIABILITYOFDIGI
TALSOILMAPSANDITSIMPACTONREGIONALECOSYSTEMMODELING%CO
LOGICAL-ODELLINGn
,INDROTH2,+INNEY++0LATZ#,2ESPONSEOFDECIDUOUS
TREES TO ELEVATED ATMOSPHERIC #/ PRODUCTIVITY PHYTOCHEMISTRY
ANDINSECTPERFORMANCE%COLOGYn
,ORD$-ORISSETTE3!LLAIRE*)NmUENCEDELINTENSITELUMIN
EUSEDELATEMPERATURENOCTURNEDELAIRETDELACONCENTRATIONEN
#/SURLACROISSANCEDESEMISDEPINETTENOIRE0ICEAMARIANAPRODU
ITSENRECIPIENTSENSERRES#ANADIAN*OURNALOF&OREST2ESEARCH
n
-ANABE 3 7ETHERALD 24 ,ARGE SCALE CHANGES IN SOIL WETNESS
INDUCED BY AN INCREASE IN CARBON DIOXIDE *OURNAL OF!TMOSPHERIC
3CIENCESn
-ARKS$4HESENSITIVITYOFPOTENTIALEVAPOTRANSPIRATIONTOCLIMATE
CHANGEOVERTHECONTINENTAL5NITED3TATES)N'OCINSKI(-ARKS
$ 4URNER $0 EDS "IOSPHERIC FEEDBACKS TO CLIMATE CHANGE THE
SENSITIVITYOFREGIONALTRACEGASEMISSIONSEVAPOTRANSPIRATIONAND
ENERGYBALANCETOVEGETATIONREDISTRIBUTION%0!#OR
VALLIS/253%NVIRONMENTAL0ROTECTION!GENCY)6nTO)6n
-C'UIRE !$ -ELILLO * - *OYCE , ! ;ET AL= )NTERACTIONS
BETWEEN CARBON AND NITROGEN DYNAMICS IN ESTIMATING NET PRIMARY
PRODUCTIVITYFORPOTENTIALVEGETATIONIN.ORTH!MERICA'LOBAL"IO
GEOCHEMICAL#YCLESn
-C'UIRE!$-ELILLO*-+ICKLIGHTER$7;ETAL=A%QUILIB
RIUMRESPONSESOFSOILCARBONTOCLIMATECHANGEEMPIRICALANDPRO
CESSBASEDESTIMATES*OURNALOF"IOGEOGRAPHYn
-C'UIRE !$ -ELILLO *- *OYCE ,! B 4HE ROLE OF NITROGEN
INTHERESPONSEOFFORESTNETPRIMARYPRODUCTIONTOELEVATEDATMO
SPHERICCARBONDIOXIDE!NNUAL2EVIEWOF%COLOGYAND3YSTEMATICS
n
-ELILLO*--C'UIRE!$+ICKLIGHTER$7;ETAL='LOBALCLI
MATE CHANGE AND TERRESTRIAL NET PRIMARY PRODUCTION .ATURE n
-IAO 3 !CORN MASS AND SEEDLING GROWTH IN 1UERCUS RUBRA IN
RESPONSETOELEVATED#/*OURNALOF6EGETATION3CIENCEn
.EILSON20!MODELFORPREDICTINGCONTINENTALSCALEVEGETATIONDIS
TRIBUTIONANDWATERBALANCE%COLOGICAL!PPLICATIONSn
.OBLE2*ENSEN+&2UFF"3,OATS+2ESPONSEOF!CERSAC
CHARUMSEEDLINGSTOELEVATEDCARBONDIOXIDEANDOZONE/HIO*OUR
NALOF3CIENCEn
.ORBY2*'UNDERSON#!7ULLSCHLEGER3$;ETAL=0RODUC
TIVITY AND COMPENSATORY RESPONSES OF YELLOWPOPLAR TREES IN ELE
VATED#/.ATUREn
.UNGESSER -+ *OYCE ,! -C'UIRE !$ %FFECTS OF SPATIAL
AGGREGATION ON PREDICTIONS OF FOREST CLIMATE CHANGE RESPONSE #LI
MATE2ESEARCHn
0AN9-ELILLO*--C'UIRE!$;ETAL=-ODELEDRESPONSES
OFTERRESTRIALECOSYSTEMSTOELEVATEDATMOSPHERIC#/ACOMPARISON
*OYCEAND.UNGESSER
OF SIMULATIONS BY THE BIOGEOCHEMISTRY MODELS OF THE 6EGETATION
%COSYSTEM-ODELINGAND!NALYSIS0ROJECT6%-!0/ECOLOGIA
n
0ARTON7*3CHIMEL$3#OLE#6;ETAL=!NALYSISOFFACTORS
CONTROLLINGSOILORGANICMATTERLEVELSIN'REAT0LAINSGRASSLANDS3OIL
3CIENCE3OCIETYOF!MERICA*OURNALn
0ARTON7*3CURLOCK*-//JIMA$3;ETAL=/BSERVATIONSAND
MODELINGOFBIOMASSANDSOILORGANICMATTERDYNAMICSFORTHEGRASS
LANDBIOMEWORLDWIDE'LOBAL"IOGEOCHEMISTRY#YCLESn
0EREZ'ARCIA * *OYCE ,! "INKLEY #3 ;ET AL= %CONOMIC
IMPACTSOFCLIMATICCHANGEONTHEGLOBALFORESTSECTORANINTEGRATED
ECOLOGICALECONOMIC ASSESSMENT #RITICAL 2EVIEWS IN %NVIRONMENT
AND4ECHNOLOGY3PECIALn
0IERCE ,, 2UNNING 37 4HE EFFECTS OF AGGREGATING SUBGRID
LAND SURFACE VARIATION ON LARGESCALE ESTIMATES OF NET PRIMARY PRO
DUCTION,ANDSCAPE%COLOGYn
0OLLE ! 0lRRMANN 4 #HAKRABARTI 3 ;ET AL= 4HE EFFECTS OF
ENHANCED OZONE AND ENHANCED CARBON DIOXIDE CONCENTRATIONS ON
BIOMASS PIGMENTS AND ANTIOXIDATIVE ENZYMES IN SPRUCE NEEDLES
0ICEAALBIES,0LANT#ELLAND%NVIRONMENTn
2OCHEFORT , "AZZAZ &! 'ROWTH RESPONSE TO ELEVATED #/ IN
SEEDLINGS OF FOUR COOCCURRING BIRCH SPECIES #ANADIAN *OURNAL OF
&ORESTRY2ESEARCHn
2UNNING 37 4ESTING &/2%34"'# ECOSYSTEM PROCESS SIMULA
TIONSACROSSACLIMATICGRADIENTIN/REGON%COLOGICAL!PPLICATIONS
n
2UNNING37#OUGHLAN*#!GENERALMODELOFFORESTECOSYSTEM
PROCESSES FOR REGIONAL APPLICATIONS ) (YDROLOGIC BALANCE CANOPY
GASEXCHANGEANDPRIMARYPRODUCTIONPROCESSES%COLOGICAL-ODEL
INGn
3AXE ( %LLSWORTH $3 (EATH * 4ANSLEY 2EVIEW .O 4REE
ANDFORESTFUNCTIONINGINANENRICHED#/ATMOSPHERE.EW0HYTOL
OGISTn
3CHLESINGER-%:HAO:3EASONALCLIMATICCHANGESINDUCEDBY
DOUBLED #/ AS SIMULATED BY THE /35 ATMOSPHERIC '#-MIXED
LAYEROCEANMODEL*OURNALOF#LIMATEn
3OLBRIG / 4 0LANT TRAITS AND ADAPTIVE STRATEGIES THEIR ROLE IN
ECOSYSTEMFUNCTION)N"IODIVERSITYAND%COSYSTEM&UNCTION%$
3CHULZE AND (! -OONEY EDS .EW 9ORK 3PRINGER6ERLAG 0AGE
n#HAPTER
%COSYSTEM0RODUCTIVITYANDTHE)MPACTOF#LIMATE#HANGE
4ISSUE$44HOMAS2"3TRAIN"2'ROWTHANDPHOTOSYNTHESIS
OF LOBLOLLY PINE 0INUS TAEDA AFTER EXPOSURE TO ELEVATED #/ FOR MONTHSINTHElELD4REE0HYSIOLOGYn
4OWNEND*%FFECTSOFELEVATED#/WATERANDNUTRIENTSON0ICEA
SITCHENSIS"ONG#ARRSEEDLINGS.EW0HYTOLOGISTn
53$!&OREST3ERVICE20!!SSESSMENTOFTHE&ORESTAND2ANGE
LAND 3ITUATION IN THE 5NITED 3TATESˆ 5PDATE &OREST 2ESOURCE
2EPORT .O 7ASHINGTON $# 53 $EPARTMENT OF !GRICULTURE
&OREST3ERVICEP
6ASIEVICH * - !LIG 2 * /PPORTUNITIES TO INCREASE TIMBER
GROWTHANDCARBONSTORAGEONTIMBERLANDSINTHECONTIGUOUS5NITED
3TATES)N&ORESTSAND'LOBAL#HANGE6OLUME&OREST-ANAGEMENT
/PPORTUNITIESFOR-ITIGATING#ARBON%MISSIONS3AMPSON2.AND
(AIR$EDS7ASHINGTON$#!MERICAN&ORESTSPAGESn
6%-!0-EMBERS6EGETATIONECOSYSTEMMODELINGANDANALYSIS
PROJECT #OMPARING BIOGEOGRAPHY AND BIOGEOCHEMISTRY MODELS IN
A CONTINENTALSCALE STUDY OF TERRESTRIAL ECOSYSTEM RESPONSES TO CLI
MATECHANGEAND#/DOUBLING'LOBAL"IOGEOCHEMICAL#YCLES
n
6INTON -! "URKE ) # )NTERACTIONS BETWEEN INDIVIDUAL PLANT
SPECIES AND SOIL NUTRIENT STATUS IN SHORTGRASS STEPPE %COLOGY n
7AYNE0-"AZZAZ&!,IGHTACQUISITIONANDGROWTHBYCOM
PETINGINDIVIDUALSIN#/ENRICHEDATMOSPHERESCONSEQUENCESFOR
SIZE STRUCTURE IN REGENERATING BIRCH STANDS *OURNAL OF %COLOGY n
7ILKINS$6AN/OSTEN**"ESFORD24%FFECTSOFELEVATED#/
ON GROWTH AND CHLOROPLAST PROTEINS IN 0RUNUS AVIUM 4REE 0HYSIOL
OGYn
7OODWARD&)3MITH4-%MANUEL72!GLOBALLANDPRI
MARYPRODUCTIVITYANDPHYTOGEOGRAPHYMODEL'LOBAL"IOGEOCHEM
ICAL#YCLESn
8IAO 8 +ICKLIGHTER $7 -ELILLO *- ;ET AL= ,INKING A
GLOBAL TERRESTRIAL BIOGEOCHEMICAL MODEL WITH A DIMENSIONAL CLI
MATEMODELIMPLICATIONSFORTHEGLOBALCARBONBUDGET4ELLUSB
n
9AKIMCHUK2(ODDINOTT*4HEINmUENCEOFULTRAVIOLET"LIGHT
AND CARBON DIOXIDE ENRICHMENT ON THE GROWTH AND PHYSIOLOGY
OF SEEDLINGS OF THREE CONIFER SPECIES #ANADIAN *OURNAL OF &OREST
2ESEARCHn
53$!&OREST3ERVICE'EN4ECH2EP2-23n'42n
Download