Interhemispheric Linkage of Paleoclimate during the Last Glaciation Author(s): George H. Denton, Thomas V. Lowell, Calvin J. Heusser, Patricio I. Moreno, Bjørn G. Andersen, Linda E. Heusser, Christian Schlüchter, David R. Marchant Source: Geografiska Annaler. Series A, Physical Geography, Vol. 81, No. 2, Glacial and Vegetational History of the Southern Lake District of Chile (1999), pp. 107-153 Published by: Blackwell Publishing on behalf of the Swedish Society for Anthropology and Geography Stable URL: http://www.jstor.org/stable/521340 Accessed: 15/07/2009 14:35 Your use of the JSTOR archive indicates your acceptance of JSTOR's Terms and Conditions of Use, available at http://www.jstor.org/page/info/about/policies/terms.jsp. 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Swedish Society for Anthropology and Geography and Blackwell Publishing are collaborating with JSTOR to digitize, preserve and extend access to Geografiska Annaler. Series A, Physical Geography. http://www.jstor.org INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION BY G.H. DENTON', C. J. HEUSSER2, T.V. LOWELL3, P.I. MORENO4, B.G. ANDERSEN5, LINDA E. HEUSSER6, C. SCHLUCHTER7 and D.R. MARCHANT8 'Department of Geological Sciences and Institute for QuaternaryStudies, University of Maine, Orono, Maine, USA 2100 Clinton Road, Tuxedo, New York, USA 3Department of Geology, University of Cincinnati, Cincinnati, Ohio, USA 4Institute for Quaternary Studies, University of Maine, Orono, Maine, USA 5Institute for Geology, University of Oslo, Oslo, Norway 6Lamont-Doherty Earth Observatory, Palisades, New York, USA 7Institute of Geology, University of Bern, Bern, Switzerland 8Department of Geology, Boston University, Boston, Massachusetts, USA Denton, G.H., Heusser, C1L,Lowell, T.V., Moreno,P.I., Andersen,B.G., Heusser,L.E.,Schliichter,C. andMarchant,D.R., 1999: Interhemispheric linkage of paleoclimate during the last glaciation. Geogr. Ann., 81 A (2): 107-153. ABSTRACT. Combined glacial geologic and palynologic data from the southern Lake District, Seno Reloncavf, and Isla Grande de Chilo6 in middle latitudes (40°35'-42°25'S) of the Southern Hemisphere Andes suggest (1) that full-glacial or near-full-glacial climate conditions persisted from about 29,400 to 14,550 14C yr BP in late Llanquihue time, (2) that within this late Llanquihue interval mean summer temperaturewas depressed 6°- 8°C compared to modem values during major glacier advances into the outer moraine belt at 29,400,26,760,22,295-22,570, and 14,55014,805 14C yr BP, (3) that summer temperature depression was as great during early Llanquihue as during late Llanquihue time, (4) that climate deteriorated from warmer conditions during the early part to colder conditions during the later part of middle Llanquihue time, (5) that superimposed on long-term climate deterioration are Gramineae peaks on Isla Grande de Chilo6 that represent cooling at 44,520-47,110 14C yr BP (T-11), 32,105-35,764 14C yr BP (T-9), 24,895-26,019 14C yr BP(T-7), 21,430-22,774 14C yr BP (T-5), and 13,040-15,200 14C yr BP (T-3), (6) that the initial phase of the glacial/interglacial transition of the last termination involved at least two major steps, one beginning at 14,600 14C yr BP and another at 12,700-13,000 '4C yr BP, and (7) that a late-glacial climate reversal of < 2-3° C set in close to 12,200 14C yr BP, after an interval of near-interglacial warmth, and continued into Younger Dryas time. The late-glacial climate signal from the southern Chilean Lake District ties into that from proglacial Lago Mascardi in the nearby Argentine Andes, which shows rapid ice recession peaking at 12,400 14C yr BP, followed by a reversal of trend that culminated in Younger-Dryas-age glacier readvance at 11,400-10,200 14C yr BP. Many full- and late-glacial climate shifts in the southern Lake District match those from New Zealand at nearly the same Southern Hemisphere middle latitudes. At the last glacial maximum (LGM), snowline lowering relative to present-day values was nearly the same in the Southern Alps (875 m) and the Chilean Andes (1000 m). Particularly noteworthy are the new YoungerDryas-age exposure dates of the Lake Misery moraines in Geografiska Annaler * 81 A (1999) · 2 Arthur's Pass in the Southern Alps. Moreover, pollen records from the Waikato lowlands on North Island show that a major vegetation shift at close to 14,700 14C yr BP marked the beginning of the last glacial/interglacial transition (Newnham et al. 1989). The synchronous and nearly uniform lowering of snowlines in Southern Hemisphere middle-latitude mountains compared with Northern Hemisphere values suggests global cooling of about the same magnitude in both hemispheres at the LGM. When compared with paleoclimate records from the North Atlantic region, the middle-latitude Southern Hemisphere terrestrial data imply interhemispheric symmetry of the structure and timing of the last glacial/interglacial transition. In both regions atmospheric warming pulses are implicated near the beginning of Oldest Dryas time (-14,600 14C yr BP) and near the Oldest Dryas/Billing transition (-12,700-13,000 14C yr BP). The second of these warming pulses was coincident with resumption of North Atlantic thermohaline circulation similar to that of the modern mode, with strong formation of Lower North Atlantic Deep Water in the Nordic Seas. In both regions, the maximum Bolling-age warmth was achieved at 12,200-12,500 14C yr BP, and was followed by a reversal in climate trend. In the North Atlantic region, and possibly in middle latitudes of the Southern Hemisphere, this reversal culminated in a Younger-Dryas-age cold pulse. Although changes in ocean circulation can redistributeheat between the hemispheres, they cannot alone account either for the synchronous planetarycooling of the LGM or for the synchronous interhemispheric warming steps of the abruptglacial-to-interglacial transition. Instead, the dominant interhemispheric climate linkage must feature a global atmospheric signal. The most likely source of this signal is a change in the greenhouse content of the atmosphere. We speculate that the Oldest Dryas warming pulse originated from an increase in atmospheric water-vapor production by half-precession forcing in the tropics. The major thermohaline switch near the Oldest Dryas/Bolling transition then could have triggered another increase in tropical water-vapor production to near-interglacial values. Introduction A fundamental problem of global climate dynam107 G.H. DENTON ET AL. ics is to identify mechanisms that drove late Quaternary glacial cycles. Strong statistical linkages imply that these cycles are ultimately related to the slow changes in the eccentricity of Earth's orbit and in the tilt and orientation of its spin axis. But what physical mechanisms translate the seasonality changes caused by these astronomical factors into global climate changes? And why is there a strong non-linear behavior of the late Quaternary climate system that produces a dominant 100-kyr signal? To complicate matters, the geologic record shows numerous large and abruptclimate changes. One of the most prominent of these occurred during the last termination, when global climate switched from a glacial to an interglacial mode. Beyond that, however, isotope and methane records in Greenland ice cores reveal widespread abrupt changes throughout the last glacial cycle (Johnsen et al. 1992). In addition, a pervasive millennial-scale oscillation of the climate system is commonly recognized (Denton and Karlen 1973; Grimm et al. 1993; Bond et al. 1997). To explain these observations, abrupt reorganizations of the global ocean-atmosphere system have been suggested as fundamental to Earth's climate system (Broecker and Denton 1990; Imbrie et al. 1992, 1993). Such reorganizations are thought to constitute jumps among stable modes of operation-shifts that cause changes in the greenhouse gas content or the albedo of the atmosphere. The challenge, then, is to understand how orbital forcing interacts with an ocean-atmosphere system that has a tendency to undergo abrupt mode changes. The atmosphere and ocean somehow work together to change the hydrologic cycle of the planet during abrupt reorganizations of the climate system. Although the exact linkages remain elusive, a strong possibility is that the roots of at least some abrupt changes lie within the thermohaline circulation of the ocean (Broecker and Denton 1990). Numerical modelling studies imply that several quasi-stable patterns of thermohaline circulation can exist because the dense water that sinks to the deep ocean is produced in several locations. Particularly prominent are sources in the Southern Ocean near Antarctica and in the northern North Atlantic Ocean. By altering the transport of heat from low to high latitudes, as well as across the equator, shifts among these patterns can affect regional or even hemisphere-wide climate. Although changes in ocean circulation can redistribute heat on the planet, they cannot by themselves produce the overall planetary cooling of the last glacial 108 maximum (LGM), nor the global warming of the last glacial-interglacial transition. Does the explanation of these planetary changes lie instead in the atmospheric inventory of water vapor (Broecker 1994, 1997a, b)? If so, then how are thermohaline switches coupled to the production of atmospheric water vapor, which occurs largely in the tropics? A coupling between thermohaline switches and tropical ocean-atmosphere dynamics during some late-glacial abruptclimate changes is suggested by the tight linkage of North Atlantic atmosphere and sea-surface temperatures (thermohaline variations) (Bond et al. 1993) with tropical Atlantic upwelling (tradewind strength) (Hughen et al. 1996, 1998) and monsoonal activity in the Arabian Sea (Schulz et al. 1998). Do the origins of abrupt changes always lie within the thermohaline circulation of the ocean, or is it possible that some lie solely in the non-linear production of tropical atmospheric water vapor? Carefully dated paleoclimate records can reveal critical characteristics of the climate system during abrupt changes. The chronology of such records may reveal the anatomy of a system rich in climate feedbacks, thus clarifying mechanisms of interhemispheric coupling. Of particular importance are whether the dramatic rapid changes in the North Atlantic region are manifested globally, whether their amplitude varies between the two polar hemispheres, whether there are leads and lags between the hemispheres, and finally, the phase relations among components of the climate system. To assess these interhemispheric linkages, we carried out field investigations in middle latitudes of the Southern Hemisphere in both South America and New Zealand. Our prime field area was the southern Lake District, Seno Reloncavi, and Isla Grande de Chiloe in southern Chile between 40035' and 42°25'S (Fig. 1). Here the wide, north-south trending Valle Central occurs between the Cordillera de los Andes to the east and the Cordillera de la Costa to the west. Within the northernpart of the field area, the valley floor is commonly less than 200 m in elevation and features Lago Puyehue, Lago Rupanco, and Lago Llanquihue. At the LGM these lakes were filled with piedmont ice lobes fed by an extensive icefield and mountain-glacier system in the adjacent Andes. Today the southern part of the valley within the field area is submerged below sea level in the marine basins of Seno Reloncavi, Golfo de Ancud, and northern Golfo Corcovado. These marine basins also were occupied at the LGM by lobes of piedmont glaciers that adGeografiska Annaler * 81 A (1999) - 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION O Ca, Fig. 1. Index map of the field area in the southern Lake District of the Chilean Andes. Valle Central is a large north-south trending depression filled with lakes and gulfs. The schematic map of the Llanquihue moraine system is derived from the four glacial geomorphological map plates in Andersen et al. (1999). The ice extent for the LGM shown in the inset is from Hollin and Schilling (1981). The numbers refer to sites discussed in the text as follows: 1, Canal de la Puntilla site; 2, Puerto Octay site; 3, FrutillarBajo site; 4, Puerto Phillippi site; 5, Fundo Llanquihue site; 6, crossroad site; 7, railroad bridge site; 8, Calle Santa Rosa site; 9, Northwest Bluff site; 10, Bella Vista Park site; 11, Bella Vista Bluff site; 12, Canal Tenglo site; 13, Punta Penas site; 14, western Puerto Montt site, 15, Huelmo site; 16, Calbuco site; 17, Taiquem6 site; 18, Dalcahue site. These sites are described in Denton et al. (1999b), Heusser et al. (1999), and Moreno et al. (1999). In addition, 19 is the Cuesta Moraga site in the Chilean Andes (Heusser 1990) and 20 is the Lago Mascardi site near Mt. Tronador in the Argentine Andes (Ariztegui et al. 1997). Geografiska Annaler · 81 A (1999) · 2 109 G.H.DENTONETAL. vanced westward from the Andes onto both the mainland and Isla Grande de Chiloe. As a result, Valle Central and Isla Grande de Chiloe both feature well-preserved moraine-and-outwash systems of the Llanquihue (last) and older glaciations. Overall, the spatial distribution of Llanquihue-age moraines shows that the lowland piedmont lobes at the LGM reached fartherwestward with increasing distance to the south, until they overrode southern Isla Grande de Chilo6 and passed into the open Pacific Ocean. The map pattern and radiocarbon chronology of these Llanquihue moraines are given in Andersen et al. (1999) and Denton et al. (1999). Andean glaciers east of the southern Lake District are now confined largely to the high volcanoes that surmount the lower crystalline peaks. However, east of Isla Grande de Chiloe small glaciers are widespread on the lower crystalline Andean peaks (frontispiece), and icefields mantle the higher volcanoes. Much of Valle Central and eastern Isla Grande de Chiloe has been cleared of forest since European settlement. But the natural vegetation is Lowland Deciduous Evergreen Forest in the lowlands of the southern Lake District, and was Valdivian Evergreen Forest in eastern Isla Grande de Chiloe. In the Lake District, this transition occurs at about 41°S at intermediate and low elevations in the mountains. In the Andes east of the southern Lake District, successively higher vegetation zones are composed of North Patagonian Evergreen Forest, and Subantarctic Deciduous Beech Forest to treeline at 1250 m. Above treeline are high-elevation Andean shrubs and herbs. To the west of the southern Lake District, and in western Isla Grande de Chiloe, the coastal range exhibits the following vegetation belts: North Patagonian Evergreen Forest, SubantarcticEvergreen Forest, and Magellanic Moorland. As well as being defined by increasing elevation in mountain ranges on both flanks of the field area, these vegetation belts occur in distinctive patterns southward through the Chilean channels to Cape Horn. Descriptions of the vegetation belts, their climate regimes in and near the field area, and their occurrence farther south in Chile at lower elevations appear in Heusser et al. (1999) and Moreno et al. (1999). We chose the area of the southern Chilean Lake District, Seno Reloncavi, and Isla Grande de Chiloe for detailed study for five reasons. First, the area lies in the middle latitudes of the Southern Hemisphere, where the orbital seasonality signal is nearly out of phase with that of similar latitudes of the 110 Northern Hemisphere that have produced most paleoclimate records of the last glacial/interglacial transition. Second, the field area is within the zone of dominant Southern Hemisphere westerlies and thus ideally situated to monitor any northwardshift of these westerlies during the LGM. Moreover, the field area is on the western flank of the Andes, and hence here the westerlies have not been affected by topographic obstacles (such as the large ice sheets that are so important in the westerlies belt of the Northern Hemisphere). Third, the field area is far removed from the North Atlantic basin, so commonly deemed the critical location on the planet for triggering climate change. Also, unlike the North Atlantic region, the area is distant from major sources of deepwater formation that can greatly affect local ocean heat transport. Fourth, the field area is at low elevations between central Isla Grande de Chiloe (42°40'S) and the northern border of the Lake District (37°S), where terminal moraine belts dating to the LGM are accessible for study and radiocarbon dating. Fifth, the field area is in the runout zone for numerous pyroclastic flows from Andean volcanoes. These pyroclastic flows mantle the moraine belts and outwash plains. They also afford sediment for interdriftdeposits, as well as for the infilling of lakes and mires. To piece together an overall paleoclimate record, we combined glacial-geologic data with pollen analysis of sediment cores from mires on Llanquihue moraines and from interdrift organic deposits. Four glacial morphologic maps were constructed for the Llanquihue moraine belts between 40°35' and 42°25'S, covering the regions of Lago Puyehue, Lago Rupanco, Lago Llanquihue, Seno Reloncavi, Golfo de Ancud, and northern Golfo Corcovado (Andersen et al. 1999). The chronology of the glacial deposits is based on more than 450 new radiocarbon dates of samples from stratigraphic sections and from the base of mires on moraine belts (Denton et al. 1999). The reconstruction of the paleovegetation is based on pollen analysis of numerous cores from surface mires and of several organic interdrift horizons (Heusser et al. 1999; Moreno et al. 1999), with extensive additional radiocarbon control. The cores were obtained with a Wright square-rod piston corer. Multiple overlapping cores were collected within 1 m of each other at many sites to avoid problems associated with core breaks; these closely spaced cores were easily correlated by magnetic susceptibility. Master cores at Fundo Llanquihue (Heusser et al. 1999) and Canal de la Puntilla (Moreno et al. 1999) Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION a. Southern Lake District, Chile (40o30'- 42°25'S; 72o25'- 73045'W) Mean Summer Temperature b. Southern Lake District, Chile (40030'- 42025'S; 72°25'- 73°45'W) Mean Summer Temperature 14°C 8°C I 10°C 12°C 160C I I C. Lago Mascardi, Argentina (41017'S; 71°35'W) Median Grain Size (pm) 3 4 5 6 7 8 - . < i - < 0 - - . II 0 . -'s o ~ - _Ct (r L. N <-Glacier 1- I I advance- I Drald bev R. D. Klv Jr. t1997 Fig. 2. Paleoclimate records for the southern Lake District of Chile and for Lago Mascardi in the adjacent Argentine Andes. Panels (a) and (b) show a schematic representationof paleoclimate relative to the approximate climate limits of present-day vegetation belts whose majorconstituents can be recognized in radiocarbon-datedpollen records. The reconstruction for middle Llanquihue time is largely from Taiquem6 (Heusser et al. 1999), and hence the labels represent Taiquem6 pollen zones. The detailed fluctuations in middle Llanquihue time come from the percentage of grass pollen (Heusser et al. 1999). We do not know the terminal position of piedmont ice margins through this portion of Llanquihue time. The reconstruction for late Llanquihue time between 30,000 and 14,550 4C yr BP shows peaks of glacier expansion into the outer Llanquihue moraine belts (Denton et al. 1999b). These peaks are placed on the diagram by assuming that the glacier maxima correspond with the most extreme vegetational environment. Such an assumption is generally consistent with the similar estimates for the lowering of snowline and treeline at the times of the most severe climate deterioration during the LGM (Porter 1981; Heusser et al. 1999; Moreno et al. 1999). A possible exception, and hence a potential weakness in panels (a) and (b), is that the glacial record shows a maximum close to 29,400 14C yr BP, whereas the pollen records from Taiquem6 and Dalcahue do not show a fully developed SubantarcticParklanduntil several thousand years later (Heusser et al. 1999). The detailed pollen diagrams from the Fundo Llanquihue and Canal de la Puntilla sites both show continued severe conditions during much of the LGM, as depicted by the dashed line between glacier maxima. The portion of the late Llanquihue record between 14,600 and 10,000 14C yr BP is from pollen records at Fundo Llanquihue (Heusser et al. 1999), Canal de la Puntilla (Moreno et al. 1999), Huelmo (Moreno 1998), and Taiquemo6 (Heusser et al. 1999). Two major warming steps terminated the LGM. We further subdivide the first step into an early warming pulse at 14,600 14C yr BP (rise of Nothofagus at Canal de la Puntilla and Huelmo) and a later warming pulse at 14,000 14Cyr BP (invasion of Subantarctic Parkland by thermophilic trees at many sites). The marked warming to near-Holocene values centered at 12,200-12,500 14C yr BP, followed by a late-glacial climate reversal, is shown at all pollen sites. Because of fire disturbance at many sites the record for the end of this reversal is shown by a dashed line, representing the situation at Taiquemo6.However, a Younger Dryas signal is shown in the sediment record from Lago Mascardi, a proglacial lake near Mt. Tronador in the Argentine Andes (Ariztegui et al. 1997) only 15 km east of our key sites of Fundo Llanquihue and Canal de la Puntilla (Fig. 1). The Lago Mascardi sediment record can be locked into the pollen records from Fundo Llanquihue and Canal de la Puntilla, because all show a distinctive warm peak at 12,200-12,500 4C yr BP, followed by a reversal in climate trend. The subsequent record from Lago Mascardi suggests a Younger-Dryas-age readvance of the Tronador ice cap (Ariztegui et al. 1997). This is similar to the situation on Isla Grande de Chilo6, where the Taiquem6 pollen record escaped the influence of fire and shows episodic cooling between 11,360 and 10,355 14C yr BP (Heusser et al. 1999). were analysed for pollen at intervals of 1 cm or less, and the chronology of each was controlled with more than 25 radiocarbon dates. Subsidiary cores were analysed at more widely spaced intervals and the chronology controlled with fewer radiocarbon dates (Heusser et al. 1999). The organic layers revealed in exposures at the Canal Tenglo and DalcGeografiska Annaler · 81 A (1999) - 2 ahue sites were also analysed at intervals of 1 cm or less; organic horizons in the other sections were less closely controlled (Heusser et al. 1999). Chilean Andes paleoclimate record Figure2 shows a schematicrepresentationof the 111 G.H.DENTONETAL. paleoclimatechangesinferredfrommorainechronologies and pollen recordsfrom the area of the southernChileanLake District, Seno Reloncavi, andIslaGrandede Chiloe.Theresultsaredisplayed in relationto paleotemperature estimatesfrompolin termsof fluctuationsof len diagrams,interpreted broadvegetationbeltsthroughtime (Heusseret al. 1999;Morenoet al. 1999).Climateparametersfor each broad modern vegetationbelt in southern Chilearereviewedin Heusseret al. (1999).Theraoutermorainebeltsof Llanquihue diocarbon-dated age arerepresentedin this diagramso thatthe glacier maximacorrespondto the mostextremevegetationalenvironment,an assumptionconsistentnot only withthe chronologiesof thepollensequences andthe morainebelts, butalso with the similarestimatesfor the loweringof snowlineandtreelineat the times of the most severeclimatedeterioration during the LGM (Porter 1981; Hubbard1997; Heusseretal. 1999;Morenoetal. 1999).Finally,the diagramindicatesthe roughboundariesfor early, middle,andlateLlanquihuetime,basedlargelyon usagedevelopedin NorthAmerica(Dreimanisand Goldthwait1973). Marineisotope stage (MIS) 4 correspondsroughlyto earlyLlanquihuetime,MIS 3 to middleLlanquihuetime,MIS2 to lateLlanquihue time, andMIS 1 to the Holocene. A potentialproblemwith our reconstructionis thatthe SubantarcticParklandthatoccurredat all the pollen sites at the LGM does not have an unambiguous modern analog. This parklandwas closest in characterto theMagellanicMoorland,as the two communitiesshare many taxa. We think that the differencebetween the two communities results from the thick and extensive alluvial fill, cappedby outwashplains,of ValleCentralandof easternIsla Grandede Chiloe.A similargeologic settingdoes notexist in thepresent-dayMagellanic Moorlandof theouter,cold andwet, rockycoastof southernmostChile. The result is that in glacial times grassesand compositesbecamewidespread within the SubantarcticParkland,where conditions were suitablefor theirgrowthandreproduction on well-drainedoutwashplains;at the same time,elementsof MagellanicMoorlandfloramade upthelowlandvegetationin boggyareaswithinthe poorly drainedmorainebelt. In other words, the present-dayMagellanicMoorlanddoes not serve as a strictmodem analogfor the ice-age Subantarctic Parklandbecause of geologic differences, and yet the two are thoughtto representthe same climateregimebecausetheyhaveso manycharacteristicplanttaxain common. 112 An importantassumptionof our conceptual frameworkis thatthelow-slopingformerpiedmont glaciers of the southernLake District, Seno Reloncavi, Golfo de Ancud, and Golfo Corcovado werereliableindicatorsof climatechanges.Inotherwords,we assumethatthebehaviorof suchlobes was dominatedby changes in surfaceconditions imposed by climate (large variationsof ablation and accumulationzones on such low-slopingsurfaces), as opposedto changes in basal conditions (frombasalhydrologyanddeformingbeds). A contraryview holdsthatsuchlow-slopingglacier lobes are inherentlyunstablewherethey rest on deformingsediment.This concept arose from the behaviorof ice streamsthatdrainthe interior ice reservoirof the marine-basedWest Antarctic Ice Sheetinto the Ross Ice Shelf (Alley andWhillans 1991a;MacAyeal1993).There,of course,the ice surfacedoes not have an ablationzone, the ice streamshave low profiles,and at least Ice Stream B rests on till. It has been postulatedthatmost of thefastflowof Ice StreamB resultsfromdeformationof this basaltill layer(Alley et al. 1987;Alley and Whillans 1991b).These ice streamsundergo episodic flow. For example,Ice StreamC is stagnanttoday but apparentlywas fast-flowingabout 200 yearsago (Alley andWhillans1991a). Generations of ice streamsseem to replaceeach other, and it may be thatthe WestAntarcticIce Sheet is still collapsingslowly as theseice streamsmigrate headwardinto the interiorice reservoir.An extension of thisconceptis thatsectorsof theLaurentide Ice Sheetrestingon sedimentarybedrock,particularlyalongthelow-slopinglobatesouthernmargin, exhibitedsimilarunstablebehaviorthoughtto be associatedwithdeformingsubglacialbeds.In fact, Clark(1994) suggestedthat inherentinstabilities wherethe LaurentideIce Sheet overrodedeforming sedimentled to irregularmarginalfluctuations which, throughtheirimpacton thermohalinecirculation, caused the abruptclimate oscillations seen in paleoclimaterecordsfromtheNorthAtlantic region.MacAyeal(1993) postulatedthatcyclic behaviorof the interiorLaurentideIce Sheetfrom alternatingthawed and frozen basal conditions caused surges of carbonate-bearingice through HudsonStrait,resultingin depositionof detrital, carbonatelayersin the NorthAtlanticOceanduring Heinrichevents.The key point is that,by this concept,therapidmarginaloscillationsof theLaurentideIce Sheetarenotthoughtto haveoriginated in climate.It shouldalso be pointedout thatother mechanismshave previously been proposedfor Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGEOF PALEOCLIMATE DURINGTHELASTGLACIATION some advancesof individuallobes of the southern LaurentideIce Sheet (Prest 1970; Wright 1973; Claytonet al. 1985). Claytonet al. (1989) counteredthatnearlyintact stratigraphicsequencesof coherentglacial units, including some with delicate sedimentarystructures, is strongevidence againstpervasivedeformationof the bed beneathformerLaurentideice lobes in the GreatLakesregion.Instead,they suggested that these lobes are low sloping because they are supportedby high basalpore-waterpressure within the underlyingsediments,with rapid flow from sliding at the ice-sediment contact.In thisway thebasalslidingis akinto a thrustfaultfacilitatedby high waterpressure.Becausebasaltill canbe draggedbeneaththeice, deformationwithin the till is a consequenceof, not the cause of, rapid ice flow.In fact,it now turnsoutthatthis argument mayalso applyto the low-slopingflankof theWest AntarcticIce Sheetin theRoss Embaymentsector, whereKamband Engelhardt(1998) reportedthat the flow mechanism beneath fast-moving Ice StreamB is largelybasalslidingat the ice-till contact,ratherthandeformationwithinthe underlying till layer. In view of this situation,we mappedandradiocarbondatedthe morainebelts of six formerAndean piedmontglaciersthroughnearly2° of latitude to ensure that the glacier fluctuationswere widespreadandhence relatedto regionalclimate, ratherthanrestrictedto the local dynamicsof one lobe (Andersenet al. 1999;Dentonet al. 1999). In like manner,we analysedpollen cores over a similar range of latitudeto ensure that the recorded vegetationchanges were representativeof the region ratherthana single site (Heusseret al. 1999; Morenoet al. 1999).These two datasets reinforce our assumptionthatthe majorglacierfluctuations were a responseto climatechange.Moreover,we were not able to find a layer of pervasivelydeformed sedimentwithin the Lago Llanquihueor Seno Reloncavibasins(TurbekandLowell 1999). The firstimportantpointto drawfromFig. 2 is that the full-glacial summertemperaturereconstructedfrom paleovegetationwas depressed68°C comparedto modem valuesat this latitudeof the Chilean Andes, an estimate consistent with snowline depression of about 1000 m from its presentpositionof 1900-2100 m in theVolcainCalbuco andLago Todoslos Santossectorof the Andes (Porter 1981; Hubbard 1997). Pollen sites within 200 m elevation of present-daysea level alongnearly2° of latitudeuniformlyshowan open Geografiska Annaler · 81 A (1999) - 2 SubantarcticParklandenvironmentat the LGM. The paucityof trees suggeststhatthese low-lying sites wereall thenneartreeline.The implicationis thattreelineloweredabout 1000 m or more at the LGMfromits present-daylimit at close to 1250 m in the adjacentAndes. Thus the temperaturedepressionaffectednot only higherelevationsnear 2000 m (snowline),butalso thelow-elevationfloor of Valle Centralwithin a few hundredmetersof present-daysea level (treeline). A secondmajorpointderivedfromFig. 2 is that the depression of summer temperaturewas as greatduringearlyLlanquihueas duringlate Llanquihuetime. This conclusioncomes fromthe moraine morphologyand pollen stratigraphyat the Taiquemosite in Isla Grandede Chiloe. The glacial morphologicmapnearthe Taiquem6site (fig. 5 in Andersen et al. 1999) depicts a complex north-south trending Llanquihue-age moraine belt, which is little-weatheredin sharpcontrastto the older distal morainebelt. The age of a readvance into this morainebelt at the Dalcahuesite (18 in Fig. 1) comes froma meanradiocarbondate of 14,805 14Cyr BP for 34 samplesfrom a paleolandsurfaceburiedby a coarsening-upwardglaciofluvialsequencethatculminatesin basal lodgment till with large (up to 1 m diameter)striated granitebouldersof Andeanprovenance(Dentonet al. 1999). The paleolandsurfaceis at the top of a layerof organicvolcanicfine sandandsilt with an extensiveseries of radiocarbondates thatextends back in sequence to 30,070 14Cyr BP (A-7685) (Heusseret al. 1999;Dentonet al. 1999).Henceat Dalcahue,andby extensionat all localities in the Llanquihuemorainebelt at andwest of Dalcahue, the advanceat 14,805 '4C yr BP was the most extensive of late Llanquihuetime. An organicclast datedto 14,820+450 (QL-4532),as well as older organic clasts dating to 19,840+180 '4C yr BP (UGA-6979) and 21,080+220 '4C yr BP (UGA6972), are reworkedinto ice-contactgravelsnear the upperlip of a prominentice-contactslope 1.5 kmnorthof theDalcahuesite (Dentonet al. 1999). This morainecan be tracednorthwardand shown to lie 4 km east of the Taiquem6pollen site (Andersenet al. 1999).The Taiquem6mire,in turn,is located within a topographicdepression on the outermostLlanquihuemoraine ridge in eastern Isla Grandede Chiloe. Thus, in the region of the Taiquem6and Dalcahue sites, all but the outermost ridges in this morainebelt are late-Llanquihue in age anddatefromclose to 14,805 '4Cyr BP (Dentonet al. 1999). 113 G.H. DENTON ET AL. The pollen record of the Taiquem6 mire comes from a core 6.55 m long that penetrates to the glacial sediments of the outermost Llanquihue moraine (fig. 20 in Heusser et al. 1999). The chronology of the Taiquem6 pollen record is controlled by 30 radiocarbon dates. The lower 55 cm of the core dates to > 49,892 '4C yr BP (AA-14770). The level at 525 cm is dated to 47,110±289314C yr BP (AA14767), which is most likely a minimum age. The point here is that a substantial portion of the lower part of Taiquem6 core is close to or beyond the limit of radiocarbon dating. The pollen record for the Taiquem6 core, given in fig. 20 of Heusser et al. (1999), shows that a closed-canopy North Patagonian Evergreen Forest had come into existence by about 13,000 '4C yr BP during the last glacial/interglacial transition, with the peak of late-glacial forest development at 12,200 14C yr BP. This forest featured a suite of thermophilic trees, including Lomatia, Myrtaceae, and Maytenus, Gramineae, and even the cold-tolerant elements of the North Patagonian Evergreen Forest, are at very low levels. Such interglacial conditions are not again encountered deeper in the Taiquem6 core. There is no evidence from the pollen record, the lithology, or the radiocarbon chronology for hiatuses in the Taiquem6 core. Therefore, we infer that the outer Llanquihue-age moraine at Taiquem6 is younger than the penultimate interglaciation. Given these age constraints, we consider it highly likely that this outer moraine was deposited during early Llanquihue time (probably at the MIS 4 maximum), as indicated in Fig. 2. We do not have direct information on the age of the outer Llanquihue-age moraine belt from Taiquem6 northwardto the middle reaches of the Llanquihueage moraines west of Seno Reloncavi. But the radiocarbon chronology given in Denton et al. (1999) shows that the outer moraine belt fringing northern Seno Reloncavi, Lago Llanquihue, and Lago Rupanco is late Llanquihue in age. However, middle or early Llanquihue advances into subsidiary moraine belts west of Lago Llanquihue are suggested by virtually unweathered till or outwash beneath radiocarbon-datedinterdriftorganic silt-andsand deposits in exposures at Puerto Octay, at Frutillar Bajo, and west of Puerto Varas (Denton et al. 1999). A third major point illustrated in Fig. 2 is that the climate deteriorated from warmer conditions during the early partto colder conditions during the later partof middle Llanquihue time (MIS 3). The pollen record from Taiquemo again affords most of the 114 pertinent evidence (fig. 20 in Heusser et al. 1999). Throughout much of middle Llanquihue time, from > 49,892 14Cyr BP (AA-14770) until about 26,019 14Cyr BP (AA-14758), the Taiquem6 mire in eastern Isla Grande de Chiloe recorded elements of Subantarctic Evergreen Forest, except during the times of the Gramineae peaks described below. During zones T-12 and T-14, prior to 47,110 14Cyr BP, Nothofagus occurs in association with arboreal elements such as Podocarpus nubigena, Pilgerodendron-type, and Pseudopanax laetevirens, along with low frequencies of Gramineae, Myrtaceae, Maytenus, and Drimys winteri. This assemblage implies the presence of Subantarctic Evergreen Forest in the early phases of middle Llanquihue time. Later in middle Llanquihue time, a diminished SubantarcticEvergreen Forest persisted until between 44,520 and 35,764 '4C yr BP; but Subantarctic Parkland expanded thereafter, with the last remnants of evergreen forest disappearing during zone T-8 between 32,105 14Cyr BP and 26,019 14C yr BP. Subantarctic Parkland then dominated until about 13,000 '4C yr BP.At the nearby Dalcahue site, remnants of Subantarctic Evergreen Forest lasted until 25,176 14Cyr BP, then to be replaced by Subantarctic Parkland (Heusser et al. 1999). A striking feature of the Taiquem6 pollen record is the succession of Gramineae peaks through middle and late Llanquihue time (Table 1) (figs 20 and 29 in Heusser et al. 1999). The Gramineae peaks during pollen zones T-9, T- 1, and T- 13 were each accompanied by declines in Podocarpus nubigena and Pilgerodendron type. The vegetation during each of these zones is interpreted as being characteristic of phases of SubantarcticParklandenvironment superimposed on the general deterioration of Subantarctic Evergreen Forest. The durations of the youngest two of these middle Llanquihue grass pollen events are 44,520-47,110 14Cyr BP for zone T-11 and 32,105-35,764 '4C yr BP for zone T-9. Grass pollen events also occurred during late Llanquihue time at 24,895-26,019 '4C yr BP for zone T7, 21,430-22,774 14C yr BP for zone T-5, and 13,040-15,200 '4C yr BP for zone T-3 (Heusser et al. 1999). In the southern Lake District, the pollen records for middle Llanquihue time extend back only to 36,960-39,340 '4C yr BP at the Frutillar Bajo and Puerto Octay sites, and intermittently to 39,660 14C yr BP at the Canal Tenglo site (Heusser et al. 1999). These records come not from a continuous core as at Taiquem6, but from interdrift organic silt-andfine-sand deposits. They cover, then, only the later Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION Table 1. Comparison of the chronology of Heinrich events in the North Atlantic Ocean (40-60°N), Gramineae maxima at Taiquemo, IslaGrandede Chilo6(42°10'S),andglaciermaximain the southernLakeDistrict-Isla Grandede Chilo6on the westernborderof the ChileanAndes(40°35'-42°25'S).All ages arein 14Cyr BP. Heinrichevents NorthAtlanticOcean Gramineaeevents IslaGrandede Chiloe Glaciermaxima ChileanAndes (Elliot et al. 1998) (Heusser et al. 1999) (Denton et al. 1999b) H-1 13,200-15,000 H-2 20,200-22,200 T-3 13,040-15,200 T-5 21,430-22,774 H-3 26,000-27,700 H-4 34,200-35,200 (32,200-35,200) H-5 44,220 T-7 24,895-26,019 T-9 32,105-35,764 14,550-14,805 22,295-22,570 21,000 (?) 26,760 - T-11 44,520-47,110 1. The table shows the durations (rather than just the peaks) of Heinrich lithic events. The duration of Heinrich events 1, 2, 3, and 4 aretakenfromthe averageradiocarbon ages of the baseandtopof the lithiclayersin severalcoresover20° of latitudein the North AtlanticOcean,as plottedin fig. 5 of Elliotet al. (1998). An alternatechronologyfor the durationof H-4 (givenin parentheses)is fromthe combinedlithicand 18O0signalin fig. 9 of Elliotet al. (1998). Previousage estimatesfor the H-4 eventareabout35,100 14Cyr. BP(Cortijo et al. 1997) and about 35,500 14Cyr. BP(Bond et al. 1993). The radiocarbon age estimate for H-5 is from Elliot et al. (1998).Radiocarbon datesfromNorthAtlanticmarine-sediment coresarecorrectedby 400 yearsforthesurface-water marine reservoireffect. 2. The Gramineae events show the duration, not just the peaks, of pulses of enhanced grass pollen in the Taiquemo vegetation record, IslaGrandede Chiloe. 3. Thetableshowsonly the peaksof Chileanglacieradvances,not the durationsof the expansionevents. 4. Thedateof 21,000 14Cyr. BP(?) listedunderGlacierMaximain the ChileanAndesrefersto the alternatechronologyof Dentonet al. (1999). phase of the much longer middle Llanquihue record farthersouth at Taiquem6, when Subantarctic Evergreen Forest was giving way to Subantarctic Parkland. The three records from stratigraphic sections in the southern Lake District show the continued presence of Subantarctic Parkland during the later phase of middle Llanquihue time, beginning at least by 36,960-39,340 '4C yr BP. These data indicate that during the later part of middle Llanquihue time, there was a transition zone in eastern Isla Grande de Chiloe between Subantarctic Evergreen Forest and Subantarctic Parkland, while Subantarctic Parkland dominated the southern Lake District. The trend evident from the vegetation record at Taiquem6 is of increasingly cold climate beginning after pollen zone T-12 (fig. 20 of Heusser et al. 1999) and continuing into the beginning of late Llanquihue time. That overall cold and wet climate persisted throughout middle Llanquihue time is indicated by the low percentages of the thermophilic arboreal species of the North Patagonian Evergreen Forest, even during early phases of middle Llanquihue time, and by the continued presence of Lepidothamnusfonkii, Astelia pumila, and Donatiafascicularis. Superimposed on this overall cooling trend are stadial-interstadial fluctuations. Subantarctic Parkland conditions marked the stadials Geografiska Annaler * 81 A (1999) · 2 (T-13, T-11, and T-9 in fig. 20 of Heusser et al. 1999), and reversion toward Subantarctic Evergreen Forest characterized the interstadials (T-14, T-12, T-10, and T-8 in fig. 20 of Heusser et al. 1999). Subantarctic Evergreen Forest diminished in dominance during successive interstadials. This fluctuating deterioration of climate does not leave a sharp delineation between middle and late Llanquihue time. Rather, the impression from the increase in frequency of Gramineae in the Taiquem6 pollen record is that the climate deterioration continued toward a culmination of cold conditions about 21,900 14Cyr BP in late Llanquihue time. The extent of Andean piedmont ice lobes during middle Llanquihue time is not well constrained. The Lago Llanquihue piedmont lobe did not advance into the outer moraine belt at the Puerto Octay site between 29,363 and prior to 39,340 14Cyr BP or at the Frutillar Bajo site between 26,760 and prior to 36,960 14Cyr BP. Nor is there evidence for nearby Andean ice at the Canal Tenglo section alongside Seno Reloncavi between 29,385 and prior to 39,660 14Cyr BP (Denton et al. 1999). These three sites are all at or near the ice-contact slopes that rise above Lago Llanquihue or Seno Reloncavi at the proximal margin of the outer Llanquihue moraine belt. The first definitive evidence that middle Llanquihue climate had deteriorated enough to 115 G.H. DENTON ET AL. send Andean piedmont glaciers into the outer Llanquihue-age moraine belt is outwash deposited about 29,400 '4C yr BP at the Puerto Octay and Canal Tenglo sites. A fourth major point shown in Fig. 2 is that climate fluctuated within a narrow range close to fullglacial conditions from about 29,400 to about 14,550 '4C yr BP. Throughout this long interval Subantarctic Parkland environments persisted in the southern Lake District (Heusser et al. 1999; Moreno et al. 1999). Andean piedmont glacier lobes repeatedly advanced into the outer Llanquihue moraine belt during this long interval. As mentioned above, during these peaks of the LGM, we estimate a mean summer temperatureabout 6-8°C lower than at present. The only discrepancy between the pollen and glacial records is near the beginning of the LGM. The glacial record implies that the earliest major advance occurred close to 29,400 14Cyr BP. However, the pollen records from Isla Grande de Chiloe suggest that full-glacial conditions may not have been achieved until about 26,000 '4C yr BP. The last remnants of Subantarctic Evergreen Forest disappeared at Taiquemo between 32,105 14Cyr BP and 26,019 14Cyr BP, and at Dalcahue about 25,176 14Cyr BP (Heusser et al. 1999). The anatomy of the youngest piedmont glacier advances is depicted in Fig. 2 and in Table 1 from details in Denton et al. (1999). The first of these is dated to 22,460 14C yr BP for the Lago Rupanco lobe; between 20,890 14Cyr BPand 23,020 14Cyr BP for the Lago Llanquihue lobe; 22,570 14Cyr BP for the Seno Reloncavi lobe; and 22,295 14Cyr BP for the northern part of the Golfo Corcovado lobe. Basal dates from mires on the moraine belts near Lago Llanquihue and Seno Reloncavi indicate recession after the culmination of this advance (Denton et al. 1999). Note that new radiocarbon dates reported in Denton etal. (1999) have caused the advance of most lobes to be placed at 22,295-22,570 14Cyr BP, ratherthan at the somewhat younger ages reported in Lowell et al. (1995). See Denton et al. (1999) for the current inventory of radiocarbon dates associated with this advance, as well as alternative interpretationsof available dates. There may also have been advances to the edge of the kame terraces alongside the western shore of Lago Llanquihue shortly before 17,800 '4C yr BP and again shortly before 15,730 14Cyr BP. The youngest readvance of the LGM is radiocarbon dated near Lago Llanquihue to 14,650 14C yr BP (Puerto Phillippi site), 14,869 14Cyr BP(Llan116 quihue site), 14,882 14C yr BP (Northwest Bluff site), 14,540 14C yr BP (Bella Vista Bluff site), 14,550-14,613 14Cyr BP (railroadbridge site), and 14,820 14Cyr BP (Calle Santa Rosa site) (Denton et al. 1999). Near Seno Reloncavf it is dated to 15,220 14Cyr BP at Isla Maillen, 14,879 14Cyr BP at Punta Penas, and shortly after 15,040 14Cyr BP at the top of the ice-contact slope beside Canal Tenglo. For the Golfo de Ancud lobe, this youngest readvance is dated to shortly after 14,900 14Cyr BP at the Calbuco site. For the northern part of the Golfo Corcovado lobe, numerous radiocarbondates from the Dalcahue site place this readvance at close to 14,805 14Cyr BP. Therefore, we conclude that these four piedmont glacier lobes fluctuated in near synchrony (within the limits of radiocarbon dating) during the youngest major pulse of the LGM. Note that because of numerous new radiocarbon dates reported in Denton et al. (1999), the date for the thin upper peat bed at the railroadbridge site is older than reported in Lowell et al. (1995). Furthermore, the ages of the lower, thick organic bed and underlying organic silt are also older than reported previously in Mercer (1976) and Hoganson and Ashworth (1992). Two older advances into the outer Llanquihue moraine system during the LGM occurred at about 29,400 14Cyr BP and at 26,760 14Cyr BP (Denton et al. 1999) (Fig. 2 and Table 1). The first of these older advances is recorded for only two piedmont lobes (Llanquihue and Seno Reloncavf), and the second for only one lobe (Llanquihue). Nevertheless, we consider it likely thatboth advances are regional, given the nearly simultaneous behavior of several lobes during the youngest two maxima of the LGM. A fifth major point in Fig. 2 involves the structure of the climate changes that, taken together, constitute the last glacial-interglacial transition. An examination of this structurerequires a detailed chronology of the youngest glacial maximum. As described above, the advance to this maximum culminated at 14,550-14,805 '4C yr BP for piedmont glacier lobes in the field area. For the northernpart of the field area, this youngest advance of the LGM terminated behind the outermost late Llanquihue moraine belt. However, in the southern portion of the field area, this final advance was the maximum of late Llanquihue time. We interpret the lack of younger radiocarbon dates associated with sediments deposited at this maximum to mean that ice recession was underway immediately. Pollen analyses of three organic beds deposited Geografiska Annaler * 81 A (1999) · 2 LINKAGEOF PALEOCLIMATE INTERHEMISPHERIC DURINGTHELASTGLACIATION just before the culmination of this youngest advance at the Llanquihue, Bella Vista Bluff, and Punta Penas sites all show cold, wet Subantarctic Parkland conditions (Heusser et al. 1999). For example, the pollen analysis of the organic silt bed at the Bella Vista Bluff site in Puerto Varas, which covers the interval from 14,540 to 15,640 '4C yr BP, indicates continuous cold Subantarctic Parkland conditions right up until 14,540 14Cyr BP, when organic accumulation ceased as a result of the lakelevel rise that heralded readvance of the Lago Llanquihue piedmont lobe. This is consistent with other detailed pollen evidence for cold moorland habitats in the southern Chilean Lake District throughout the latter part of the LGM from the Canal de la Puntilla (Moreno 1997; Moreno et al. 1999) and Fundo Llanquihue (Heusser et al. 1999) sites. The open environment and increased precipitation implied by the moorland pollen taxa suggest a northward shift of the westerlies storm tracks (Moreno et al. 1999). These paleoenvironmental inferences from pollen analysis are consistent with earlier conclusions drawn from fossil beetle data (Hoganson and Ashworth 1992; Ashworth and Hoganson 1993). One of the key fossil-beetle sites is an organic silt layer dated to 15,715 440 14Cyr BP (GX-5275) within the kame terrace at the Bella Vista Park site (Hoganson and Ashworth 1992), situated about 175 m north of the Bella Vista Bluff site. The age and stratigraphicposition indicate that this organic bed at the park site corresponds with the organic silt bed at the bluff site. Another key sample locality is the Puerto Varas railroad bridge site, where a silt and sand unit is capped by a 26-cm-thick layer of organic silt, peat, and wood (see Denton et al. 1999). Mercer (1976) reported a date of 16,270± 360 14C yr BP (RL-1 13) of wood from near the base of the silt and sand unit. Hoganson and Ashworth (1992) gave an age of 14,060±450 14C yr BP (GX5507) for the capping organic layer. Ten samples of fossil beetles come from regular intervals within the silt and sand unit, and two samples from within the capping organic layer (Hoganson and Ashworth 1992). The implication based on these two dates is that the beetle samples come from regular intervals between 16,270 14Cyr BP and 14,060 14C yr BP, the critical last several thousand years of the LGM (Hoganson and Ashworth 1992, table 1 and fig. 4). However, our new dates show that the entire silt and sand unit was deposited at 17,350-17,880 14C yr BP (Denton et al. 1999), and hence that the ten beetle samples from the silt and sand unit refer Geografiska Annaler · 81 A (1999) · 2 only to that time. In addition, wood from the upper few centimeters of the capping organic layer yielded a mean age of 14,613 '4C yr BP (Denton et al. 1999). In view of the fact that the capping organic bed is developed into the top of the silt unit, it is not clear whether the two beetle samples from the organic bed refer to 14,613 '4C yr BP or to 17,35017,880 '4C yr BP. With this revised chronology, the fossil-beetle data are invaluable in showing cold, wet environmental conditions near Lago Llanquihue at 16,000-18,170 14Cyr BP (Canal de Chanchan site near Puerto Octay), 17,350-17,880 14Cyr BP (railroadbridge site in Puerto Varas), and 15,715 14Cyr BP (Bella Vista Park site in Puerto Varas); all of these times occur within the long interval of the LGM. We earlier presented evidence for significant amelioration of climate at the end of the last glaciation. That evidence was the invasion of the lowlands of the southern Chilean Lake District by thermophilic arboreal elements of the North Patagonian Evergreen Forest. This arboreal diversification began with the spread of Myrtaceae, Nothofagus cf. dombeyi, Lomatia, Maytenus and other relatively thermophilic arborealtaxa, which became prominent by 13,900 14Cyr BP at the Canal de la Puntilla site, by 13,500 14Cyr BP at the Fundo Llanquihue site, and by 13,700 '4C yr BP at the Alerce site (Lowell et al. 1995). To determine more closely the beginning of the glacial/interglacial transition, we subsequently investigated in detail several cores in a transect at the Canal de la Puntilla site (Moreno et al. 1999) and a single core with a high sedimentation rate at the Huelmo site (Moreno 1998). In both cases the chronology was established with numerous new AMS radiocarbon dates. This detailed examination shows that the first indication of warming was a rapid rise of Nothofagus at 14,600 14Cyr BP at both sites. Warming continued with the invasion of thermophilic tree species of the North Patagonian Evergreen Forest about 14,100 '4C yr BP at Canal de la Puntilla and 14,200 '4C yr BP at Huelmo. These vegetation changes are taken to be regional because they are recorded nearly simultaneously at two widely separated sites. The chronologies show that both vegetational changes are rapid at each site. Thanks to these detailed and well-dated stratigraphies, we can place age brackets on the climate change that marked the end of glacial conditions in the southern Chilean Lake District. The pollen record from the Bella Vista Bluff section in the terrace at Puerto Varas shows cold conditions until the 117 G.H. DENTON ET AL. culmination of the last glacial advance of the LGM at 14,550-14,805 14Cyr BP (Heusser et al. 1999). The first indication of subsequent climate amelioration is the rise of Nothofagus at 14,600 14Cyr BP at Canal de la Puntilla and at Huelmo (Moreno 1998; Moreno etal. 1999), followed by invasion of thermophilic species at about 14,000 14Cyr BP at many sites in the Lake District (Heusser et al. 1999). Note that because of extensive new radiocarbon dating described in Denton et al. (1999), this age for the beginning of the last termination is slightly earlier than we reported in Lowell et al. (1995). The pollen records from throughout the southern Chilean Lake District and on Isla Grande de Chiloe register a consistent late-glacial pattern of vegetational development (Heusser et al. 1996, 1999; Moreno 1997, 1998; Moreno et al. 1999). The inferences drawn from the detailed pollen records from the Fundo Llanquihue and Canal de la Puntilla sites are supported by a network of cores analysed with less temporal resolution. The interval between 14,000 and 13,000 14Cyr BP is a transition zone in which thermophilic arboreal elements of the North Patagonian Evergreen Forest invaded the lowlands, but in such relatively low numbers that grasslands and elements of Magellanic Moorland remained. The persistence of moorland taxa suggests that the westerlies storm belt remained in the inferred northern glacial position. A closed-canopy North Patagonian Evergreen Forest was then established at 12,700-13,000 14C yr BP. The rapidity and magnitude of this event at 12,700-13,000 14C yr BP are particularly well shown in the influx diagram from Canal de la Puntilla (Moreno et al. 1999), but are also evident in other diagrams. One explanation for the disappearance of moorland taxa is the postulated southward shift of the westerlies storm track. The evergreen forest reached its fullest development about 12,000-12,200 '4C yr BP, and may at that time have contained some elements of the Valdivian Evergreen Forest.After 12,000 '4C yr BP, opening of the forest and the appearance of cold-tolerant elements of the North Patagonian Evergreen Forest (Podocarpus nubigena, Pseudopanax laetevirens) suggest climate cooling that continued until at least 10,500 14Cyr BP, when the Fundo Llanquihue and many other vegetational records are disturbed by the influence of fire. However, farthersouth on Isla Grande de Chiloe, the Taiquem6 record escaped the influence of fire, and here North Patagonian Evergreen Forest communities opened to allow the 118 significant presence of Podocarpus and Pilgerodendron-type during episodic cooling between 11,360 and 10,355 14Cyr BP(Heusser et al. 1999). We mentioned above that the overall deterioration of climate at the coldest times of the LGM was about 6-8°C in mean summer temperaturerelative to the present-day value (Heusser et al. 1999; Moreno et al. 1999). About 3°C of this difference was recovered in the first warming pulses between 14,600 '4Cyr BPand 12,700-13,000 14CyrBP.Another 3°C was recovered between 12,700-13,000 14Cyr BP and 12,200-12,500 '4C yr BP. Our data suggest that warming at the beginning of these intervals was rapid. Finally, we infer that the late-glacial climate reversal after 12,000-12,200 '4C yr BP involved a relatively small decline in temperature (probably < 2-3°C, Heusser et al. 1999), compared to the high values achieved at 12,200-12,500 14C yr BP. New Zealand paleoclimaterecord To test the Chilean paleoclimate record as a barometer of change in the zone of Southern Hemisphere westerlies, rather than simply as a piece of a complex matrix of varying regional climate changes, we examined the available records and we obtained additional radiocarbon dates from New Zealand at the same Southern Hemisphere middle latitudes but on the opposite side of the Pacific Ocean. Although this test is preliminary, we conclude that major climate shifts in New Zealand are similar to those in the Chilean Andes. The basis for this conclusion follows. Grassland and shrubland were dominant during the LGM in most of New Zealand south of about 37°S (the latitude of Auckland in northern North Island) (McGlone 1995), with continuous forest persisting only in the extreme north (Fig. 3). Only small patches of Nothofagus, Libocedrus, and podocarp trees remained in South Island. Forest stands may have persisted in hilly areas of North Island south of 37°S, but grassland and shrubland dominated the lower, rolling terrain. Such extensive deforestation probably reflects severe frosts from outbreaks of cold subantarcticair masses over most of New Zealand during the LGM (McGlone 1988). The oceanic reconstruction from Deep Sea Drilling Project (DSDP) Site 594 on the south flank of Chatham Rise 300 km east of South Island is consistent with severe terrestrial conditions, because it shows cold subantarctic water close to South Island during the LGM (Nelson et al. 1993). Geografiska Annaler - 81 A (1999) 2 · INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION 170° 168° I 172° I I 174° 176° 178° I I I Explanation Ice today [-j I Ice at last glacial maximum |Alpine vegetation .r Grassland - shrubland t I! Tall, lowland - montane lowlands coniferous broadleaf forest Fig. 3. The extent of mountain glaciers in the Southern Alps is from Hollin and Schilling (1981) and the distribution of vegetation is from McGlone et al. (1993), both during the LGM at approximately 18,000 14C yr BP. Also listed are the key stratigraphic sites mentioned in the text. The radiocarbondates associated with these sites and not listed in the text or in Denton and Hendy (1994) are as follows. At the Kamaka site, an organic-rich bed is overlain by laminated lacustrine deposits and then by coarse outwash. The laminated deposits reflect glacier expansion. They have yielded the following succession of AMS radiocarbon dates of small enclosed wood and plant fragments, from bottom to top: 22,420±230 14C yr BP (AA22271), 22,460±240 14Cyr BP (AA22275), 22,360±190 14C yr BP (ETH-16387),22,620+200 '4CyrBP (ETH-16388),22,160±310 4C yrBP (AA-22275), 22,490±330 14Cyr BP (AA-22279), 22,100+790 14Cyr BP (AA-22282), 22,570±200 14Cyr BP (AA-22283), 22,860+200 14Cyr BP (AA-22285), 22,730±200 14Cyr BP (AA-22286), 22,680+220 14Cyr BP (AA-22290), 22,650+240 14Cyr BP (AA-22291), 21,990±220 14Cyr BP (ETH-16390), and 22,380+190 14C yr BP (ETH-16391. + |Scattered forest areas | t|7 Present-day coastline Coastline at last glacial maximum I Radiocarbon or exposure sample site Pollen site e Raupo Kamaka Kumara--" Abut Head Omoeroa Bluff ~-- -Pukaki o DSDP Site 594 9 0 Scale 100 200 300 km °.. 192 4s° 17n- 172° 174° 176° 178° Drafted by Richard D. Kelly Jr., Augusta, Maine, 1997. In anycase, the harshterrestrialclimateduringthe LGMpromotedinstabilityof the deforestedlandscape, with widespreadmechanical weathering, slopeerosion,andloess deposition;extensivealluvial plains developedbetween the SouthernAlps andthe east coast of SouthIsland. Figure 4 displays the glacier advances in the SouthernAlps thatoccurredwithinthis LGMenvironment.Duringthe maximumof the lateOtiran glaciation(LGM),equilibrium-linealtitudesnear Lake Pukaki were depressed 875 ±50 m below present-dayvalues during deposition of the Mt John(Kumara-22) morainesand750 + 50 m during Geografiska Annaler * 81 A (1999) · 2 River deposition of the Tekapo(Kumara-33)moraines (Porter 1975). Radiocarbondates from several sites indicate multiple advances into late Otiran morainebeltsthataretakento be theequivalentsof the outerof the two late Otiranmorainebelts near LakePukaki(Suggate1978).An organicbedoverlain by till occurswithinthe outerOtiranmoraine belt at Mt Hercules on the western flank of the SouthernAlps. Three small wood samples from within this bed yielded new ages of 23,870 ± 330 14C yr BP (A-6188), 23,560 370 14C yr BP (A6591), and 23,510 350 '4C yr BP (A-6592). The glacial advancerepresentedby the overlyingtill 119 G.H. DENTON ET AL. Fig. 4. Paleoclimate records for New Zealand. The left panel shows the radiocarbon-dated glacial deposits indicative of ice advances into the outer Otiranmoraine belts of the LGM. These are plotted with regardto the associated drop in equilibrium line altitude associated with correlative moraine belts near Lake Pukaki (Porter 1975). The existence and chronology of the Younger-Dryas-age glacier readvance is from Denton and Hendy (1994), Lowell et al. (1995), and Ivy-Ochs et al. (1999). The right three panels show the record for core DSDP Site 594 (Nelson et al. 1993, Heusser and van der Geer 1994). The marine pollen reflects changing vegetation in New Zealand. The CaCO3 record is a reflection of the amount of silt input from New Zealand. The benthonic 6180 signal is largely a reflection of Northern Hemisphere ice-sheet volume. On the right is shown the numbers of marine isotope stages from Nelson et al. (1993). likely occurred shortly after this time. Farthernorth in the Grey River Valley, a stratigraphic section at Kamaka registers an advance of the northern of the two major piedmont lobes of the TaramakuGlacier system at the LGM. Here an organic silt bed lies beneath laminated glacial lacustrine sediments and outwash that herald advance of this lobe to the outer moraine system (Suggate 1965). New AMS radiocarbon dates of 14 small wood pieces from the lacustrine sediments yielded a mean value of 22,400 14Cyr BP, consistent with an earlier date of 22,300+350 14Cyr BP (NZ-116) from the organic silt bed (Suggate 1965). A prominent maximum of the western of the two piedmont lobes of the former Taramaku Glacier system occurred near Kumara at close to 17,700 14Cyr BP. This maximum was the greatest of late Otiran time for this lobe. The age comes from five new AMS radiocarbon dates of small samples of grass, wood, bark, and beetles from near the top of an organic silt bed that underlies till with a non-erosive conformable contact. These dates are 17,380+ 130 14CyrBP(ETH-13398),17,720±120 '4C yrBP 120 (ETH-11104), 17,900±140 14C yr BP (ETH13399), 18,160±140 14Cyr BP(ETH-13400), and 18,360 + 140 14C yr BP(ETH-13402). Two other radiocarbon samples previously collected within the organic silt bed yielded ages of 18,450 ±300 14C yr BP (NZ-4408) and 17,250+250 14C yr BP (NZ4407) (Moar 1980). The terrace at Raupo (Kumara-22terraceof Suggate and Moar 1970) in the Grey Valley northeast of Kamaka reveals an interstadial organic silt bed with an aggregate thickness of 60 cm that separates two outwash units. Previous radiocarbon ages of samples from within this bed are 18,600+290 14C yr BP (NZ-891) and 18,750+180 14Cyr BP (NZ737) (Suggate and Moar 1970). Additional radiocarbon samples that we collected yielded ages of 19,740± 150 14C yr BP (A-6550) for the base of the organic silt bed, 18,940+170 14Cyr BP (A-6551) for the middle, and 18,780 140 14CyrBP(A-6552) for the top. Renewed outwash deposition in the Grey Valley after 18,780 '4C yr BP probably represents glacier advance into the outermost late Otiran moraine system. Also, glacier ice must have exGeografiska Annaler - 81 A (1999) · 2 DURINGTHELASTGLACIATION LINKAGEOF PALEOCLIMATE INTERHEMISPHERIC tended into this moraine system when outwash was deposited below the organic silt bed. Near Abut Head on the Tasman seacoast, an advance into the outer moraine belt resulted in the deposition of a large moraine on a thin organic bed now exposed in a sea cliff. Radiocarbon samples from the undisturbed uppermost part of this bed give the following new ages for this advance: 16,615+95 14CyrsBP(A-9063), 16,575 +8 14CyrBP (A-9064), 16,920±100 14C yr BP (A-9065), and 16,525 ±90 14Cyr BP (A-9066). The advance to the widespread Kumara-33 moraines (or their equivalents), nested just behind the outer Otiranmoraine belt, is dated only at Omoeroa Bluff, cut into a moraine ridge on the Tasman Sea coast near Franz Josef Glacier. Here two radiocarbon samples of a thin peat bed buried by drift of this youngest advance yielded results of 13,950±140 4C yr BP (NZ-479) (Wardle 1978) and 15,300± 120 14C yr BP (Suggate 1990). Behind these moraines (or their equivalents) in most valleys lie basins, now or formerly filled with lake or sea water, which reflect rapid collapse of ice tongues. Ice cleared these basins prior to 13,500 14C yr BP in the Tasman Valley near Lake Pukaki and 13,400 14Cyr BP in the Paringa River valley (Suggate 1968). From this preliminary chronology, Suggate (1965) and Suggate and Moar (1970) estimated that rapid ice recession close to 14,000 14C yr BP marked the end of the LGM. As discussed below, however, palynological data from the Waikato lowlands in North Island place the first decisive warming of the last glacial/interglacial transition a bit earlier, at shortly after 14,700 14C yr BP, based on the radiocarbon ages of numerous distinctive volcanic tephras registered in pollen records (Newnham et al. 1989). Late-glacial readvances are recorded in many valleys of the Southern Alps. Readvance of Franz Josef Glacier toward the Waiho Loop moraine occurred at 11,050± 14 14C yr BP (Denton and Hendy 1994, 1995). A late-glacial moraine remnant in upper Cropp River Valley was dated to 10,250 14Cyr BP by Basher and McSaveney (1989). We have obtained an error-weighted mean age of 10,055+±29 14Cyr BPfor five additional wood samples from this remnant (Lowell et al. 1995). Because Mabin (1995) cautioned against accepting the concept of a regional Younger-Dryas-age readvance on this limited evidence, exposure dates were obtained on boulders from a prominent late-glacial moraine in Arthur'sPass at the head of a tributaryof the former TaramakuGlacier system. The dating results sugGeografiska Annaler * 81 A (1999) · 2 gest that the moraine is Younger Dryas in age (IvyOchs et al. 1999). This moraine represents snowline lowering of about 260-360 m relative to the Little Ice Age value (Ivy-Ochs et al. 1999). Figure 4 also depicts the pollen, carbonate, and 6180 records from marine sediment cores from DSDP Site 594 on the south flank of Chatham Rise 300 km east of South Island (Nelson et al. 1993; Heusser and van der Geer 1994). These records are linked to the changing paleoenvironment of South Island, because DSDP Site 594 is situated offshore and downwind from the Southern Alps, from alluvial fans of the CanterburyPlains, and from major rivers that drain the Southern Alps. The pollen record shows the changing percentages of trees and shrubs as reflected in pollen carried offshore by the prevailing westerly winds and by rivers draining the east coast of South Island. The carbonate record is taken to represent increases in silt input (and a corresponding dilution of CaCO3) during cold intervals from terrestrialerosion and outwash aggradation, with the resulting generation of loess and offshore sediment transport. The pollen and carbonate records are closely correlated with each other, presumably because the extent of forest cover went hand in hand with landscape stability and hence sediment production on South Island. Both of these terrestrial indicators seem to correspond with the benthonic 6180 record. In turn, the benthonic 6180 record is determined by changes in both temperatureand ice volume, with the latterbeing dominant. Because the Antarctic Ice Sheet showed only modest volume change through the last glacial cycle, the benthonic 6180 signal largely follows Northern Hemisphere ice-volume changes. The beginning of reforestation at the end of the LGM is documented in North Island in the Waikato lowlands. Grass and shrubs dominated these lowlands at the LGM (McGlone et al. 1993). Reforestation of the Waikato lowlands by a podocarp-hardwood community was rapid after deposition of the Rerewhakaaitu tephra at 14,700 14Cyr BP (Newnham et al. 1989). Podocarp-hardwood forest had spread across much of North Island by about 14,000 14Cyr BP, followed by progressive reforestation of the rest of New Zealand (McGlone et al. 1993). It is importantto note that the vegetation just prior to 14,700 14Cyr BP was essentially the same as that earlier in the LGM at 18,000 14Cyr BP depicted in Fig. 3 (McGlone etal. 1993). The massive changes in the forest cover of North Island at the end of the LGM, shortly after 14,700 14C yr BP, were synchronous with the rapid rise of Nothofa121 G.H. DENTON ET AL. gus, followed by the invasion of thermophilic trees into the Chilean Lake District, at the end of the LGM. Landscape stabilization accompanied reforestation in central North Island, as documented by the relative preservation of highly erodible tephras. The landscape was unstable throughout the LGM at elevations above 300 m (Leamy et al. 1973; McGlone et al. 1984; Kennedy 1988). The Rerewhakaaitu tephra, dated to 14,700 14Cyr BP (Froggatt and Lowe 1990), is the earliest to be preserved on highly eroded LGM landscapes above 300 m. By 13,080 14C yr BP (Rotorua Tephra; Froggatt and Lowe 1990), extensive areas above 500 m were stable. By 11,850 14C yr BP (Waiohau Tephra; Froggatt and Lowe 1990), landscapes were stable up to 900 m elevation (McGlone 1995). Middle-latitudeSouthern Hemisphere data in a global context In order to place the middle-latitude Southern Hemisphere data from Chile and New Zealand into a global perspective, we first present background about the asymmetric shape of the 100,000-yr glacial cycles of late Quaternarytime, as our data bear on the mechanisms that terminatedthe last such cycle. We also review critical elements of the classic North Atlantic/European paleoclimate record because it is so important for our interhemispheric comparison. We then go on to address the main implications of our data in the context of this background. We conclude by posing a series of questions raised by these implications, and suggest an initiative to answer these questions. Background Late Quaternary 100,000-yr glacial cycles. Explaining the asymmetric 100,000-yr glacial cycles of late Quaternarytime is one of the most difficult challenges of paleoclimate research (Imbrie et al. 1993). The transition toward these 100,000-yr cycles began about 950,000 years ago, but took more than 300,000 years to complete (Imbrie et al. 1993). By 600,000-650,000 years ago a clear asymmetric 100,000-yr cycle had emerged. The subsequent glacial extremes of MIS 16, 12, 10, 6, and 2 reflect the storage of excess ice in huge Northern Hemisphere ice sheets just prior to terminations that abruptly ended the buildup phases of these asymmetric cycles (Raymo 1997). The overall asymmetric shape of the last 122 100,000-yr ice-volume cycle, depicted in Fig. 5, features a prolonged growth phase terminated by a much shorter collapse phase. The existence of such a prolonged growth phase implies to us that the icesheet system (and probably the entire climate system) became largely detached from pervasive orbital forcing early in the glacial cycle (except for the long-term oscillations superimposed on the prolonged growth phase) (see also Imbrie et al. 1993). Also superimposed on the long buildup phase, at least in the North Atlantic region, are shorter asymmetric oscillations known as Bond cycles (Bond et al. 1993; Broecker 1994; Rasmussen et al. 1997). Each such cycle represents a period of climate cooling that ended with a massive Heinrich discharge of icebergs (Table 1), in turnfollowed by abrupt increase of North Atlantic sea-surface temperatures. Each of these abrupt North Atlantic warmings failed to terminate the long growth phase of the 100,000-yr cycle. Instead, climate reverted to cold conditions and ice-sheet buildup continued until the decisive Heinrich 1 ice collapse (Table 1), which was coincident with the beginning of the irreversible decline in ice-sheet volume registered in TR163-31B (Fig. 5). A striking feature of the late Quaternarymarine 6180 record is that the terminations of glacial 100,000-yr cycles had similar magnitudes during times of both high-amplitude and low-amplitude insolation changes. For example, increases both in northern summer insolation and in tropical equinox insolation were considerably smaller during the last termination than during the penultimate termination, and yet both terminations had the same amplitude. Because of the effect of long eccentricity cycles on the amplitude of precession, the same situation occurred about 400,000 years ago during TerminationV between MIS 12 and 11. This is consistent with the postulate that terminations represent ocean-atmosphere reorganizations between preferred glacial and interglacial modes of the climate system (Broecker and Denton 1990). And yet terminations recur at approximately the 100,000-yr intervals set by eccentricity (which controls the amplitude of precession). The implication is that during terminations the increased effect of insolation somehow triggers a fundamental reorganization of a non-linear system but does not control the magnitude of the reorganization. Asymmetric 100,000-yr climate cycles of late Quaternarytime vary in length. In fact, the time intervals between terminations range from 84,000 to 120,000 years (Raymo 1997). Moreover, terminaGeografiska Annaler .81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION Core TR163-31B (3°37'S; 83058'W) 610 (%o) 5.5 6.0 V I lT r t t ? T 5.0 I 4.5 1- t 4.0 I t t t V-19-30 (3023'S; 82°21 'W) Sea Level (m below present) 3.5 o t 3.0 150 100 t , I 50 I f 0 . o0 · * . ** 00 * o .C^S B oi° **o to *.*. *^S o %-12,860t250 c- 15,080:200 16,120t260 *- o° * 6 o < 17,200±220 * * 0 *- ** ~ - 19,580+300 <20,960±370 .--20,890O360 * 9 * * 0 C I 14,020:1180 - 14,340t220 < o 21,480±280 -21,450t250 "I */ 0 o. % ----23,930O320 25,690±460 N4 * P. wuellerstorfi * Uvigerina Orfd by R. D. Kety Jr. a - ----28,690600 % a 917 Fig . 5. The right panel, adapted from Shackleton (1987), shows a sea-level estimate based on the 5 'O records of V 19-30 in the equatorial eastern Pacific Ocean (3°23'S, 83°21'W) and RC-17-177 in the equatorial western Pacific Ocean (1°45'N, 159°27'E). To construct this curve, a correction for changing deep-water temperatures was made by calculating the isotopic difference between V- 19-30 (benthonic) and RC 17-77 (planktonic). This difference was then subtracted from the benthonic V- 19-30 record. The results were scaled to the New Guinea sea-level data to yield an estimate from isotopic data of the global sea-level record. Even though it is certainly not a perfect representation of sea level, the resulting curve nevertheless highlights the long, gradual decline of sea level (increase of continental ice volume), followed by an abrupttermination, that marked the latest 100,000-yr glacial cycle. The left panel shows the timing of the last termination in detail. The figure displays the isotope data set from equatorial eastern Pacific core TR163-31B (3°37'S, 83°58'W) from Shackleton et al. (1988), with original isotope data points kindly provided by NJ. Shackleton. The radiocarbon dates have a marine reservoir correction of 580 14Cyears. The results from two species of benthonic foraminifers are plotted. They are taken to be a reasonable reflection of changes in continental ice volume. They show that the fundamental change between rising and declining isotope values, which marks the beginning of the last termination, occurred close to 14,500 14Cyr BP (not adjusted for unknown mixing time of the ocean at that date). tions that lead to full deglacial conditions similar to those of the present day seem to occur only after accumulation of considerable excess ice in huge Northern Hemisphere ice sheets (Raymo 1997). In other words, the major prerequisite for a sharp, complete termination seems not to be excessive orbital forcing, but rather excessive ice volume. North Atlantic/European late-glacial climate. Mangerud et al. (1974) defined a set of chrons for Geografiska Annaler .81 A (1999) - 2 the subdivision of late-glacial time. The Boiling chron was placed at 12,000-13,000 14Cyr BP, the Older Dryas chron at 11,800-12,000 14C yr BP, the Aller6d chron at 11,000-11,800 14Cyr BP, and the Younger Dryas chron at 10,000-11,000 14Cyr. BP. One of our major points given below is that the climate trigger for the last termination occurred during the millennia before the B6iling chron. Therefore, for the purposes of this paper, we use the term Oldest Dryas chron to encompass the interval be123 G.H. DENTON ET AL. a. Southern Alps, New Zealand (42°-44°30'S; 169°- 172°W) MountainGlaciers ELADepression (m) 1000 , . , , , 500. I . . . b. c. Eastern NorthAtlanticOcean SU81-18 (37°46'N;10°11'W) Chile Southern LakeDistrict (40030'-42°25'S; 72o25'-73045'W) MeanSummerTemperature 6180 (%o) 8°C 10°C 12°C 14°C 16°C 0I I I I i 2 I 1 0 c C - Preboreal 0 2 ___0 ,--- Z" =z-: .-11,e1n - _S.-12-W CL 32,73313--1^t3.t . 13,40t10. ,n i i __a a O, T Z:. 0,-, '- 1- 2 uYounger Dryas 1i23,3e1701 OlderDry B11lingOldest Dryas i o n 0 l i ..i.. 0 - < -- . I . . . i t N o ,) Drafted by R. D. Kelly Jr. 1997 d. e. f. Switzerland MountainGlaciers Gerzensee 06°(46°- 48°N; 10°E) (46°50'N;07°33'E) ELA Depression (m) 1000 8180 (%o) 0 -10 500 -8 -6 Q. _Prboreal Younger Dry_ X- Aller8d I tW Older Dryas' _rc --B1111ng -U'~u_ (U " - Oldest Dryas . I. . -- . . - I. - I -.... , <-Glacier advance- Fig. 6. Comparison of terrestrialrecords from middle latitudes of the Southern Hemisphere with paleoclimate records from the North Atlantic region. Panels (a) and (b) are from Figs 2 and 4. Panel (c) is adapted from Bard etal. (1987). It shows the oxygen-isotope values and AMS radiocarbon ages for the planktonic species Globigerina bulloides in core SU8 1-18 in the eastern North Atlantic Ocean off Portugal. The AMS radiocarbon dates have a standardmarine reservoir correction of 400 14C years. The curve reflects a combination of ice volume and temperaturechanges. It shows the same structureof paleoclimate signals as the records from middle latitudes of the Southern Hemisphere in panels (a), (b), and (d). Particularly important is that the last termination, defined on the basis of the isotope record, begins at 14,500 14C yr BP (Bard et al. 1987). This is the same result found in the detailed isotope record from core TR163-3 1B in Fig. 7 from Shackleton et al. (1988). It correlates with the initial abrupt warming in the southern Chilean Lake District (panel (b)) and in New Zealand (Newnham et al. 1989). Panel (d) is from Ariztegui et al. (1997) and represents a curve of glacier activity in the Argentine Andes that has similarities with the Greenland stable isotope record in panel (e) from Johnsen et al. (1992). Panel (f) shows the Gerzensee lacustrine 6180 record from the forelands of the Swiss Alps (Eicher and Siegenthaler 1976), which likewise exhibits a correspondence with the Greenland record. Also shown in panel (/) are the approximate equilibrium-line elevations on mountain glaciers during the Zurich Stade just before massive recession began about 14,600 14C yr BP (Schliichter 1988; Ivy-Ochs et al. 1996), for the time of maximum Bolling warmth (Maisch 1982), and for Egesen time (Maisch 1995; Ivy-Ochs et al. 1996). The European chronozones shown on panels (c) and (f) are from Mangerud et al. (1974) as modified in the text. An important point here is that the Swiss Alps experienced massive deglaciation during Oldest Dryas time correlative with deglaciation in the southern Chilean Lake District (panel (b)). 124 Geografiska Annaler · 81 A (1999) · 2 DURINGTHELASTGLACIATION LINKAGEOF PALEOCLIMATE INTERHEMISPHERIC tween 13,000 and 15,000 14C yr BP, much in the sense of Welten (1982). This means that we place the boundary of the Upper Pleniglacial and the Late Glacial subages of Hammen et al. (1967) at 15,000 14Cyr BP, not at 13,000 14Cyr BP. The Late Glacial subage would then include the Oldest Dryas, B6lling, Older Dryas, Aller6d, and Younger Dryas chrons. However, when we discuss the climate signature of central Europe, we use the system of pollen zonation of Firbas (1949, 1954), which Lotter et al. (1992) considered as strict biozones. The basic shape for the North Atlantic/European late-glacial climate signal is manifested in British Isles beetle remains (Atkinson et al. 1987), along with 6180 switches in both Greenland ice and Swiss lacustrine marl (Dansgaard et al. 1984) (Fig. 6). This shape consists of an abrupt change near or shortly after the Oldest Dryas/B611ing transition from cold conditions to an interval 200 years in length that was close to interglacial warmth. This warm interval was followed by progressive cooling, with oscillations, throughlate Boiling, Older Dryas, and Allerod time, culminating in the Younger Dryas cold episode (Oeschger 1991). There is remarkable coherence to this regional climate signal, such that even the superimposed short-lived Gerzensee, Older Dryas, and early Preboreal 6180 oscillations can be traced easily across this portion of the globe. The only major discrepancy is that, whereas the 6180 and beetle records show an early B611ing-warmpeak followed by a cooling trend, most pollen-recordsexhibit continuous vegetation development from herbs and shrubs to a succession of forest trees. The only widespread reversal in the pollen record took place in the Younger Dryas and to a much less degree in the Older Dryas (Watts 1980). The discrepancy between the magnitude of early Bolling warmth in Europe implied by vegetation as opposed to isotope and insect records may be more apparent than real. An example comes from Lobsigensee on the Swiss Plateau (Ammann and Lotter 1989; Ammann 1989a, b; Elias and Wilkinson 1983, 1985). The Oldest Dryas part of the Lobsigensee records features a boreal and boreal-montane insect assemblage, along with Betula nana. During the early B611ing warming, the boreal insect assemblage is replaced by a plant-independent temperate assemblage that reflects a mean July temperature close to interglacial values, just as in Great Britain. A shift in water plants supports such marked warming. The early Bolling climate was warm enough to support broad-leafed deciduous trees such as oak or hazel, which do not appearuntil Geografiska Annaler · 81 A (1999) · 2 3000 years later because of migrational lags. It is interesting to note that the 6180 record shows lateglacial climatic deterioration beginning in latest Boiling time and culminating in a Younger Dryas reversal. The vegetation record shows only a small increase in non-arboreal pollen in Younger Dryas time, reflecting some openings in the forest cover. The alpine moraine record shows widespread Egesen moraines of Younger Dryas age (Ivy-Ochs et al. 1999). In sharp contrast to the situation in Great Britain,Younger Dryas cooling is not reflected in the insect record at Lobsigensee. The decisive warming that occurred throughout the North Atlantic region near the Oldest Dryas/ Boiling transition was abrupt. The 6180 transition occurred within a century or less in Greenland ice cores (Oeschger et al. 1985) and in Swiss sediment cores (Siegenthaler et al. 1984). In the British Isles a cold and continental climate ended suddenly, with a 10°C rise of mean annual temperature in an interval of 300 to 800 years. By 12,500 14Cyr BP British climate was as warm as that of today (Atkinson et al. 1987). At Ballybetagh in Ireland, re-establishment of vegetation began about 12,600 14Cyr BP;the warmest interval of B611ing-Older Dryas-Aller6d time was at 11,900-12,400 14Cyr BPand was followed by climate deterioration leading to the Younger Dryas reversal (Cwynar and Watts 1989). In central Europe north of the Alps, a treeless steppe-tundra with some dwarf birch yielded swiftly to reforestation near the biozone Ia/Ib (Oldest Dryas/ B6lling) boundary (Lotter et al. 1992). This change is shown by an abrupt rise in the arboreal/ non-arboreal ratio in numerous pollen profiles; it was followed by the spread of trees across Europe during biozones lb, Ic, and II (B6iling, Older Dryas, Aller6d). For example, vegetation in the Swiss and adjacent French Alps changed markedly in parallel with b180 shifts in precipitation near the biozone Ia/Ib (Oldest Dryas/B61iing) boundary (Eicher and Siegenthaler 1976; Eicher et al. 1981). Here an increase in juniper coincides with the initial 6180 rise. At some sites reforestation by birch began during this early juniper phase, but at most sites it occurred shortly after the juniper rise. This birch reforestation is by far the most dramatic late-glacial event recorded in pollen diagrams on the northern margin of the Alps. Subsequently, a rise of pine marks the beginning of biozone II (Aller6d); open pine forest with some birch and juniper characterizes biozone III (Younger Dryas); and finally a transition from open to 125 G.H. DENTON ET AL. closed pine forest marks the biozone III/IV (Younger Dryas/Preboreal) boundary (Lotter 1991; Lotter et al. 1992). A similar situation prevailed in the Pyrenees adjacent to the Atlantic Ocean and the Mediterranean Sea (Jalut et al. 1992). Here the Oldest Dryas/ Bolling boundary is marked by a rapid increase in birch, a beginning of pine expansion and, at some localities, the spread of juniper.Again the rapid rise of arboreal pollen, which represents the most pronounced change in the vegetation records, signals the real change from an open glacial landscape to an extensive woodland. Subsequently, a rise in pine characterizes the Allerod biozone, whereas the subsequent Younger Dryas biozone is notable for expansion of Artemesia and juniper, with concurrent decline of birch and pine. The circulation regime of the surface and deep waters of the North Atlantic Ocean also changed fundamentally near the Oldest Dryas/Bolling transition. The polar front retreatedrapidly (Ruddiman and McIntyre 1981; Bard et al. 1987); the planktonic foraminifers in core Troll 3.1 show that warm surface water leaked into the Norwegian Sea about 13,400 14Cyr BP and reached near-modern values by about 13,100 14C yr BP, signaling a major change in surface circulation (Lehman and Keigwin 1992a). The circulation regime of the deep ocean showed renewed strong thermohaline production of North Atlantic Deep Water (NADW), including Lower North Atlantic Deep Water (LNADW) formed by convection in the Nordic Seas. Carbonisotope measurements of foraminifers from highsedimentation core RC11-83 from Cape Rise off South Africa show that the first significant contribution of NADW flow to the Southern Ocean occurred between 12,700 and 13,100 14C yr BP (Charles and Fairbanks 1992). Because it is closely tied to surface conditions in the Nordic Seas, rapid turnover of LNADW most likely began with the warm surface water incursion documented in the Troll 3.1 core (Lehman and Keigwin 1992a). This is consistent with Cd/Ca and 613Cratios from core EN-1200-GGCI near Bermuda Rise in the path of deep-water flow (Keigwin et al. 1991). The rapid increase in northwardheat flux caused by this fundamental change in surface and deep circulation must have contributed to the dramatic warming in Europe near the Oldest Dryas/B611ing transition through the demonstrated linkage between North Atlantic sea-surface temperature and European climate (Rind et al. 1986; Lehman and 126 Keigwin 1992a). The fact that this is the earliest such dramatic warming over Europe is consistent with the notion that LNADW, as it now forms, was cut off during the LGM and Oldest Dryas time. Thus the flooding of warm and salty surface water into the Nordic Seas and the consequent initiation of the Nordic Heat Pump of Imbrie et al. (1992), with its extensive import of ocean heat (Lehman and Keigwin 1992b), was probably the most importantevent in the switch to an interglacial climate mode. A less dramatic earlier warming pulse occurred in Europe within Oldest Dryas time. The primary evidence comes from using records of the prominent late-glacial climate signal that swept across Europe near the Oldest Dryas/B611ingtransition to unravel the deglacial chronology of the European Alps. This prominent Oldest Dryas/B611ing signal is coherent from Greenland to Switzerland. The southeastern end of this transect is anchored by the paleoclimate record at Gerzensee in Switzerland that shows the characteristic 6180 and vegetation shifts at the biozone Ia/Ib (Oldest Dryas/B611ing) boundary (Eicher and Siegenthaler 1976) (Fig. 6). For us, the importantpoint is that not only Gerzensee but numerous other lakes and bogs with similar evidence for the Oldest Dryas/B61iing warming event are located well within the boundaries of the expanded glacier system of the European Alps at the LGM, a situation previously pointed out by Schltichter (1988). It is particularly pertinent that many such sites occur in major valleys and passes. At each site, the Oldest Dryas/Bolling transition is located above the base of the core, and hence occurred subsequent to deglaciation. The major point is that extensive deglaciation of the European Alps had already occurred before the Oldest Dryas/ Bolling warming that is so prominent in paleoclimate records across the North Atlantic Ocean and Europe. In fact, the areal distribution of the core sites in Switzerland indicates that, by the time of this Oldest Dryas/B61iing transition, mountain glaciers were already confined to upper reaches of deep alpine valleys or to inter-valley mountain massifs. The radiocarbon age of the Oldest Dryas/ Bolling transition can only be placed roughly at 12,700-13,000 14Cyr BP, because of an atmospheric 14C plateau of several hundred years duration (Zbinden et al. 1989). If it is assumed that this transition was simultaneous within the Swiss and Austrian Alps, as it seems to have been across Europe, then this means that considerable mountain deglaciation antedated 12,700-13,000 14Cyr BP. Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION We interpret this early deglaciation of the Alps as evidence for an Oldest Dryas rise in mountain snowlines in southern Europe within the Oldest Dryas chron. We attributethis snowline rise to climate warming. A contrary suggestion has been made for mountain glaciers of the French Vosges, near the site of the Grande Pile bog (Seret et al. 1990, 1992). It is inferred from indirect evidence that these glaciers were likely to have achieved their maximum in middle Pleniglacial time when slightly warmer climate was presumably accompanied by increased snowfall; glaciers were supposedly contracted during cold late Pleniglacial conditions because they were starved of precipitation. That such a scenario could not apply to the Austrian and Swiss Alps is demonstrated by stratigraphic sections near Innsbruck and Ztirich, respectively (Patzelt and Resch 1986; Schltichter et al. 1987). At both sites till representing glacier advance to the maximum position of the last glaciation is bracketed by radiocarbon ages of about 29,000 and 14,000 14C yr BP; therefore the maximum extent of glaciers coincided with late Pleniglacial cold and dry conditions, not with the more humid conditions of middle Pleniglacial time. Supporting this contention is the lack of radiocarbon evidence in the Swiss and Austrian Alps for ice recession during late Pleniglacial time. What was the timing of the Oldest Dryas deglaciation of the Swiss and Austrian Alps, and what was its duration? The initiation of this recession is defined at Lake Zurich. Here deglaciation from the Zurich Stadial moraines, which are situated only 20 km behind the maximum ice-margin position of the LGM (Schltichter et al. 1987; Schliichter and Rothlisberger 1995), occurred close to 14,600 14C yr BP (Lister 1988). Applying to a core from Lake Zuirich, this date comes from a twig in non-disturbed deposits immediately above ice-rafted and tectonized sediments deposited as glacial ice cleared the lake. Hence, we place the beginning of snowline rise and climate amelioration at shortly before 14,600 14Cyr BP. Widespread deglaciation deep into the mountains driven by this snowline rise was complete before the onset of warming near the Oldest Dryas/B6lling transition. From the amount of mountain snowline rise, we can estimate the fraction of the total full glacialinterglacial change encompassed in this Oldest Dryas warming of the European Alps. Full-glacial mountain snowline depression relative to the AD 1850 position (Maisch 1992) was about 1100 m (Furrer 1991 and references therein). Snowline Geografiska Annaler * 81 A (1999) · 2 lowering during the Zurich Stadial was slightly less. By the end of the widespread Oldest Dryas deglaciation, the remaining alpine ice was confined to the upper reaches of the deep valleys and to the inter-valley massifs. Glacier termini most probably stood near the Gschnitz or Clavadel moraines, as shown by the position of these moraines relative to radiocarbon-dated pollen cores. The Clavadel moraines register a snowline lowering of about 430 m relative to the AD 1850 position, and the Gschnitz about 670 m (Furreret al. 1987). Hence the snowline rise during Oldest Dryas time was at least 330 m and perhaps as much as 670 m. This amounts to about one-third (and perhaps more) of the full glacial-interglacial snowline rise. By comparison with pollen records on the north flank of the Alps at Gerzensee and Faulenseemoos (Eicher and Siegenthaler 1976), Tourbiere de Chirens (Eicher et al. 1981), Rotsee (Lotter 1991), and Soppensee (Lotter et al. 1992), the climate amelioration that caused the snowline rise during Oldest Dryas time was not sufficient to allow reforestation north of the Alps. Instead, relatively severe conditions persisted, and as a result the vegetation remained open. In the French and Swiss Alps this open Oldest Dryas landscape was dominated by a grass assemblage and some shrubs, along with alpine and steppe herbs. However, all of the abovementioned sites lie within the limits of expanded alpine ice at the LGM. Therefore, we must look to other sites outside the last glacial limits for evidence of initiation of the Oldest Dryas environment. In the Pyrenees for example, palynologic data indicate two phases of vegetation development prior to the Oldest Dryas/B61iing transition (Jalut et al. 1992). The first phase reflected cold, open steppe and semi-desert conditions that persisted through the LGM until about 15,000 14Cyr BP. The second phase began about 15,000 14Cyr BP with a large increase in pollen concentration, a spread of juniper, and perhaps an initial establishment of birch and pine in some places. This second phase was succeeded by the typical rise of birch at the Oldest Dryas/Bolling transition. Hence, in the Pyrenees the Oldest Dryas climate warming episode began about 15,000 14C yr BP. In southern France at both Les Echets and Le Bouchet, the vegetation record indicates that the arid, cold late Pleniglacial ended about 15,000 14Cyr BP with the rise of Artemisia, Chenopodiaceae, and Caryophyllaceae accompanied by the spread of steppe, which marks the first warming. These botanical events signal the late Pleniglacial/Oldest Dryas 127 G.H. DENTON ET AL. transition. Within the sparse available radiocarbon control, then, this transition is equivalent to the beginning of ice recession from Lake Zurich. Farther north in the Netherlands, a sharp rise in Artemisia about 14,000 14Cyr BP during the Oldest Dryas was followed by the immigration of large birch trees in the Blling (Hammen and Vogel 1966). Recession of the Scandinavian-Barents Sea icesheet complex suggests that Oldest Dryas snowline rise and associated climatic amelioration was not restricted to the Alps but was widespread throughout Europe (Andersen 1981). By the time of the Oldest Dryas/B611ing transition, the southern margin of the Scandinavian Ice Sheet stood at the Luga (dated between 12,650 and 13,200 14C yr BP)-North Lithuanian-Wolin-Halland ice-marginal position, and thus had retreated about 250 km from its maximum LGM position (Andersen 1981; Lundquist and Saarnisto 1995). Almost 175 km of this recession was from the Vepsovo/Pomeranian and Krestay/Kalinin ice-marginal positions, which have estimated ages of 15,000 14Cyr BPand 14,500 14Cyr BP, respectively (Andersen 1981). This recession took place during the Rauniss interstade of still-severe climate, radiocarbon-dated between 13,250 14Cyr BP and 14,300 14Cyr BP (Serebrjannyj et al. 1970; Raukas 1976; Velichko and Faustova 1986). Hence European alpine glaciers and the Scandinavian Ice Sheet both showed considerable recession in the 1600 14C-yrinterval preceding the Oldest Dryas/B61iing warming. In the case of the Scandinavian Ice Sheet, this recession ended with readvance to the Luga moraine; in the case of the alpine system, the recession ended with ice margins close to the Gschnitz or Clavadel moraines in the inner Alps. The Oldest Dryas climate amelioration devastated the glacier system of the European Alps because of its susceptibility to a moderate snowline rise, while the massive Scandinavian Ice Sheet lost marginal ice but still remained largely intact. The marine-based, western margin of the coalesced Barents Sea and Scandinavian Ice Sheets also showed Oldest Dryas recession. One piece of evidence comes from a low-b180 spike in cores from off the continental margin (Jones 1991; Weinelt et al. 1991). The first age estimate of this spike was 14,500 14Cyr BP from low-sedimentation-rate core PS-21295 in the Fram Strait off the Barents Sea continental shelf (Jones and Keigwin 1988). But younger ages for this spike of 13,60014,000 14Cyr BPcome from cores with higher sedimentation rates farthersouth in the Norwegian Sea 128 (Sarnthein et al. 1992). This first pronounced 6b80 decrease in the Norwegian Sea is taken to represent a great iceberg outburst caused by collapse of marine-based ice sheets from the Norwegian and Barents continental shelves (Jones 1991; Sarnthein et al. 1992; Landvik et al. 1998). Likewise, a pronounced 6180 depletion is recorded at 13,10014,300 14Cyr BP just northwest of the Faeroe Islands during the Heinrich 1 event (Rasmussen et al. 1997); the magnitude of this spike implies significant iceberg contribution from the Faeroe and Shetland Islands, Scotland, and Scandinavia. This is consistent with the conclusion of Fronval et al. (1996) from marine-sediment cores off the western margin of the Scandinavian Ice Sheet. A near-basal radiocarbon date from the Troll 3.1 core is also consistent with this premise, because it suggests ice recession from the outer Norwegian continental shelf at 14,700-15,000 14C yr BP (Lehman et al. 1991). These early conclusions are supported by new radiocarbondates from marine sediment cores that show recession of grounded portions of ice sheets from the Barents and the Norwegian continental shelves at 14,500-14,800 14Cyr BP (Bischof 1994; Haflidason et al. 1995; Svendsen et al. 1996). This recession of marine margins corresponds with retreat of terrestrial Scandinavian ice during the Rauniss interstade, as well as with Oldest Dryas recession of mountain glaciers in the Swiss Alps. Taken together, these data imply that the Oldest Dryas amelioration affected the European land mass over a wide latitude range and marked the onset of the last termination. It should be noted that the Oldest Dryas climate amelioration in Europe was not nearly as dramatic as the subsequent warming near the Oldest Dryas/ Bolling transition. Paleoclimate records from the Rauniss interstade of Oldest Dryas age in northern Europe and from correlative Oldest Dryas pollenbearing sediments in France and Switzerland indicate that the climate still remained severe enough to preclude reforestation north of the Alps. Even at the end of Oldest Dryas time, Europe was still marked by a cold tundra and steppe environment, reflecting cold North Atlantic sea-surface temperatures. This is consistent with the fact that NADW of the current mode, with a strong LNADW component produced in the Nordic Seas from the inflow of warm, salty water, was not evident in Oldest Dryas time (Sarnthein et al. 1992). In contrast, the dramatic warming near the Oldest Dryas/B611ing transition was tied to a major thermohaline switch, with the initiation of strong LNADW production Geografiska Annaler · 81 A (1999) - 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION Drafted by R. D. Kelly Jr. 1998 Fig. 7. Southern Chilean Lake District paleoclimate record (from Fig. 2) transferredto a calendar-year time scale and compared with Greenland and Antarctic ice-core stable isotope records. The ice-core records are from Sowers and Bender (1995) and Steig et al. (1998). The ice cores are placed in a common calendar-year time scale by linking the Antarctic 6 80 record of ice to the Greenland chronology by matching the signatures of trapped gasses in the ice cores (after correcting for the difference in ice and gas ages). and greatly increased poleward heat flux in the North Atlantic Ocean (Lehman and Keigwin 1992a, b; Sarnthein et al. 1994). Global implications of Chilean Andes and New Zealand data Figure 5 shows a detailed 6180 record of benthonic foraminifers from core TR163-31B from the eastern equatorial Pacific Ocean, taken to be representative largely of Northern Hemisphere ice volume. Fig. 6 compares the paleoclimate records from the Chilean Andes and Southern Alps of New Zealand with those from the North Atlantic region. Fig. 7 displays the Southern Hemisphere middle-latitude paleoclimate records compared with the Antarctic and Greenland stable isotope signals in the Taylor Dome, Byrd, and GISP2 ice cores. The following implications are derived from these figures and from the text. Relative to present-day values, snowline and/or treeline depression at the LGM was about the same in the Southern Alps (875 m) and the ChilGeografiska Annaler . 81 A (1999) · 2 ean Andes (1000 m) as in many mountain areas in the Northern Hemisphere. Moreover, the timing of the LGM was similar in both polar hemispheres (with the possible exception of Antarctica). The implication is of planetary cooling at the LGM that was about equivalent in both polar hemispheres. * The last glacial-interglacial transition began abruptly in both hemispheres (at least outside of Antarctica) with a warming pulse at close to 14,600 14Cyr BP (17,300 cal. yr BP) within Oldest Dryas time. In Chile the last glacier expansion of the LGM culminated at 14,550-14,805 14Cyr BP, and was followed by massive recession. Note that, because of new radiocarbon dates given in Denton et al. (1999b), this age is slightly older than reported in Lowell et al. (1995). Nothofagus increased significantly at the Canal de la Puntilla and Huelmo sites (Moreno 1998; Moreno et al. 1999) about 14,600 14Cyr BP. At a number of sites in Valle Central over nearly 2° of latitude, pollen diagrams show at about 14,000 14Cyr BP the first strong influx of thermophilic elements of the 129 G.H. DENTON ET AL. North Patagonian Evergreen Forest since before 49,892 14C yr BP (Heusser et al. 1999). In the Southern Alps of New Zealand, significant glacier recession had occurred prior to 13,500 14C yr BP after an advance to near-maximum positions between 14,000 and 15,000 14Cyr BP. In northern New Zealand reforestation of the Waikato lowlands began shortly after 14,700 14Cyr BP (Newnham et al. 1989) and much of the open glacial landscape of North Island was rapidly reforested by podocarp and hardwood trees by 14,000 14C-yrBP. The 6180 curve of benthonic foraminifers from core TR163-3 lB in the eastern tropical Pacific Ocean shows that a steady increase in benthonic 6180 gave way to a unilateral decrease about 14,500 14Cyr BP (this 6180 shift is not adjusted for unknown mixing time of the ocean at that date) (Fig. 5). We take this change to mark the beginning of the termination of glaciation in the Northern Hemisphere, because it implies a fundamental change from volume increase to volume decline of Northern Hemisphere ice sheets. In core SU8118 in the eastern North Atlantic Ocean off Portugal, the termination also began at about 14,500 14C yr BP as defined by the planktonic 6180 record (Fig. 6). As detailed above, the southern margin of the Scandinavian Ice Sheet retreated significantly during Oldest Dryas time and grounded portions of ice sheets receded from both the Barents and the Norwegian continental shelves between 14,500 and 14,800 14Cyr BP. Also as mentioned above, the alpine glacier system in the Swiss Alps contracted from the forelands to the inner valleys between 14,600 14Cyr BP and 12,700-13,000 14Cyr BP. In addition, by the end of Oldest Dryas time, the southern margin of the Laurentide Ice Sheet in North America had already retreated to the position of the Port Huron moraine system (Mayewski et al. 1981) and the southern Cordilleran Ice Sheet had already undergone considerable recession (Booth 1987). Another major step of the last glacial/interglacial transition was a decisive warming pulse at 12,700-13,000 14Cyr BP. In the Chilean Andes this step is marked by the rapid spread of a closed-canopy North Patagonian Evergreen Forest throughout the lowlands of the Chilean Lake District and Isle Grande de Chiloe. Development of this forest culminated at 12,00012,200 14C yr BP, when climate conditions were close to full interglacial values. Plant species in- 130 dicative of wet moorland environments disappeared, possibly from a southward shift of the westerlies storm belt (Moreno et al. 1999). In the North Atlantic region this second step is marked by the abrupt warming near the Oldest Dryas/B611ing transition shown by isotope, methane, and dust records in the Greenland ice cores, as well as by isotope and vegetation records in Swiss lacustrine sediments. The modem mode of North Atlantic thermohaline circulation resumed, with strong production of LNADW and pronounced warming in the Nordic Seas from the inflow of warm, salty water. The insect records from Great Britain and the Swiss Plateau show that this warming near the Oldest Dryas/B11ling transition culminated in temperatures nearly as high as those of today (Atkinson et al. 1987; Amman 1989a). At the height of B611ingwarming, snowline on Swiss alpine glaciers probably rose to the position it occupied during the AD 1850 highstand of the Little Ice Age (Maisch 1982). A Younger-Dryas-ageclimate reversalcharacterized the North Atlantic and perhaps the middlelatitude Southern Hemisphere paleoclimate records. The Greenland ice-core 6180 signal in Figs 6 and 7 shows the typical form of the North Atlantic climatic deterioration,which began after the peak B6iling warmth and culminated in the Younger Dryas cold reversal. In the Chilean Andes, pollen diagrams show climate reversal (expansion of Podocarpus nubigena followed later by decline of mesic North Patagonian taxa) beginning at 12,000-12,200 14Cyr BP after peak late-glacial warmth(Heusser et al. 1999; Moreno et al. 1999). Unfortunately,fire disturbanceof the vegetation is indicated at many sites during Younger Dryas time, complicating paleoclimate interpretation. However, the Taiquemo pollen record escaped the influence of fire and shows episodic cooling between 11,360 and 10,355 14C yr BP (Heusser et al. 1999). In this regard,the sediment record from proglacial Lago Mascardi in Argentina, located near Mt. Tronador only 115 km east of our key pollen sites at Canal de la Puntilla and Fundo Llanquihue, shows evidence not only of rapid ice recession beginning at 13,000 14C yr BP and peaking at 12,400 14C yr BP, but also of a subsequent reversal of trend that culminated in a Younger-Dryas-age glacier readvance between 11,400 14C yr BP and 10,200 14C yr BP (Ariztegui et al. 1997) (Figs 2,6). In the Southern Alps of New Zealand, an advance of Franz Josef Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION Glacier occurred at the beginning of Younger Dryas time at 11,050 14C yr BP (Denton and Hendy 1994) (the beginning of Younger Dryas time in the NorthAtlantic region is most accurately dated to 11,030 14Cyr BP in Switzerland; Hajdas et al. 1995). AnotherYounger-Dryas-ageglacier advance is dated to 10,100 14Cyr BPin the upper Cropp River Valley (Basher and McSaveney 1989; Lowell et al. 1995). Exposure dates indicate that the prominent Lake Misery moraine complex in Arthur'sPass is YoungerDryas in age (Ivy-Ochs et al. 1999). These lattermoraines represent snowline lowering of about 260-360 m below the Little Ice Age position, compared with a similar value of 200-300 m for the Egesen moraines of Younger Dryas age in the Swiss Alps (Kerschner 1978, 1985; Maisch 1982, 1987). Thus there is mounting evidence for YoungerDryas-age readvanceof SouthernAlps glaciers in New Zealand, although caution in accepting such a conclusion is urged by Mabin (1995), McGlone (1995), and Singer et al. (1998). The Chilean Andes glacial geologic record features pronounced maxima during the LGM at 29,400 14C yr BP, 26,700 14C yr BP, 22,29522,570 14Cyr BP, and 14,550-14,805 14Cyr BP (Fig. 2 and Table 1). The vegetation record from Taiquem6 shows a series of striking maxima of grass pollen during middle and late Llanquihue time. The duration of these Gramineae events are listed in Table 1 (Heusser et al. 1999) as 44,520-47,110 14C yr BP for T-11; 32,10535,764 14Cyr BP for T-9; 24,895-26,019 14Cyr BP forT-7; 21,430-22,774 14Cyr BP for T-5; and 13,040-15,200 14Cyr BP for T-3. Also shown in Table 1 are the durations (not the peaks) of the Heinrich lithic events over 20° of latitude in the North Atlantic Ocean (Elliot et al. 1998). Heinrich events H- 1, H-2, H-3, H-4, and H-5 show a reasonably close match to Gramineae events in Chile. With the exception of the situation at 29,400 14Cyr BP, the Chilean glacier maxima fall within or close to the Heinrich-Gramineae events. This reinforces our earlier conclusion that there is a broad correspondence between North Atlantic Heinrich events and Chilean Andes glacier maxima (Lowell et al. 1995). However, the far greater chronologic and stratigraphicdetail now available (Denton et al. 1999; Moreno et al. 1999; Heusser et al. 1999) shows an importantnew insight. The last glacier maximum (14,550-14,805 14C yr BP) in the Chilean Andes occurs at the beginning of the Geografiska Annaler * 81 A (1999) · 2 long H-1 event in the North Atlantic Ocean (Table 1). It is notable that much of the H-1 lithic event correlates with glacier recession in the Andes. This relationship is reinforced by detailed pollen records showing that a marked rise of Nothofagus and then the invasion of thermophilous tree species into the Chilean Lake District (Moreno et al. 1999; Heusser et al. 1999) was also coeval with much of the H- 1 lithic event in the North Atlantic Ocean. Moreover, recession of terrestrial Scandinavian ice and mountain glaciers in the European Alps was coincident with the younger part of the H-1 lithic layer. The implication is that the H- 1 lithic event encompasses not only ice-sheet expansion onto North Atlantic continental shelves during global cooling, but also the subsequent unstable collapse of marine-based ice. That this advancecollapse sequence may be characteristic of Heinrich events is suggested by the age of the major Chilean glacier advance of 22,29522,570 14C yr BP near the beginning of the H-2 lithic event. The New Zealand Southern Alps record features the last two of the major glacier maxima in the Chilean Andes. In addition, New Zealand glaciers show maxima at 17,500 14C yr BP and 16,200 14Cyr BP. Maxima may also have been achieved just before and just after deposition of the Raupo interstadial bed, now dated between 19,740 and 18,780 14Cyr BP, as well as during Younger Dryas time. A potential weakness of our trans-Pacific comparison is that some of the glacier fluctuations are not yet recorded in both data sets. One could therefore argue that the climate signals do not match during these parts of the record. But we think it more likely that these differences merely reflect the fact that glacial moraine sequences are inherently incomplete. Questions What caused the atmospheric cooling at the LGM evident in the snowline and/or treeline records from the Chilean Andes and Southern Alps? We argue that this cooling is approximately synchronous and of the same magnitude in both hemispheres, thus pointing to the primary role of overall atmospheric cooling (from greenhouse gas content or reflectivity) rather than simply redistribution of heat on the planet from changes in ocean circulation. 131 G.H. DENTON ET AL. What caused the atmospheric warming evident in the first two major steps (Oldest Dryas and Oldest Dryas/Billing) of the last termination? These two steps, registered in both hemispheres, abruptly terminated the LGM, together bringing the atmosphere from full-glacial to nearly full-interglacial temperature in about 1600 14Cyr. The synchrony and magnitude of these two steps in the two hemispheres, at least outside of Antarctica, implicates greenhouse gas as the direct cause of atmospheric warming ratherthan simply switches in ocean heat transfer. The Byrd ice-core record in Antarctica, calibrated with the Greenland ice-core chronology by trapped gas, shows only a minor change in atmospheric CO2 during these steps (Neftel et al. 1988; Staffelbach et al. 1991; Sowers and Bender 1995; Blunier et al. 1997). If the new ice-core chronologies are correct, the implication is that jumps in atmospheric CO2 were not responsible for the abrupt warming steps that terminated the last ice age simultaneously in both hemispheres. This leaves changes in the inventory of atmospheric water vapor as the most likely factor to have caused the two major atmospheric warming steps during the last termination. It follows that a decreased inventory of atmospheric water vapor was the most important cause of LGM planetary cooling (Broecker 1994). An alternate explanation is that Northern Hemisphere ice sheets responded to seasonality forcing and, in turn, drove the last termination through their global thermal impact. The radiocarbon chronology of the two important initial steps of the termination makes this explanation highly unlikely. These two steps, which encompassed 1600 14Cyr, culminated in B6iling time with atmospheric temperatures approaching interglacial values from Greenland to the Swiss Plateau, as well as in the Chilean Andes. Reference to Fig. 5 shows that ice volume was then still near the maximum value of the LGM. From this phasing of events it thus seems most probable that, ratherthan causing this crucial warming step of the termination, the ice sheets responded to it by a greatly increased ablation rate registered in the sea-level record as meltwater pulse IA (Fairbanks 1989). What caused the changes in water-vapor production implied by paleoclimate data during the initial Oldest Dryas step of the termination? It is unlikely that the thermal impact of a decrease in ice-sheet albedo or elevation is implicated, because glacier retreat started simultaneously in both hemispheres 132 (see also Broecker and Denton 1990). A major North Atlantic thermohaline switch is also an unlikely trigger, as the Heinrich 1 ice-rafting event of Oldest Dryas age actually suppressed overturning in the North Atlantic Ocean to its lowest level (Sarnthein et al. 1994). Likewise, our data suggest that any southward shift of the Southern Hemisphere westerlies that might implicate important sea-ice changes in the Southern Ocean was delayed until near the Oldest Dryas/ B611ing transition (Moreno et al. 1999). The effect of rising summer insolation in the Northern Hemisphere is commonly cited as a trigger for the ocean-atmosphere reorganization of the last termination (Imbrie et al. 1992, 1993). For example, Broecker and Denton (1990) pointed out that terminations commonly occur during rises toward maxima in seasonality at middle and high latitudes in the Northern Hemisphere. But our data imply that any such Northern Hemisphere effect was translated to middle latitudes of the Southern Hemisphere by an atmospheric process. In the absence of major changes in Northern Hemisphere ice sheets or of major switches in North Atlantic thermohaline circulation, it is difficult to identify a high-latitude mechanism that was triggered by rising summer insolation during Oldest Dryas time. This difficulty is compounded by the fact that during Oldest Dryas time the Greenland ice-core 680 record is dissimilar to paleoclimate records elsewhere in the Northern Hemisphere and in Chile, probably because of a regional climate signal induced by the Heinrich 1 iceberg influx into the North Atlantic Ocean. This suggests to us that increased water-vaporproduction in the tropics is the most likely source of the initial Oldest Dryas step of the last termination, perhaps from forcing by half-precession insolation, whose amplitude is controlled by eccentricity (McIntyre and Molfino 1996; Berger and Loutre 1997). What caused the changes in water-vapor production inferred to have produced decisive warming near the Oldest Dryas/Billing transition? A key point is that near-interglacial warmth was achieved in early Blling time in both hemispheres (from Greenland to the Swiss Plateau in the North Atlantic region, as well as in the Chilean Andes). It is very unlikely that this event was driven by the direct thermal impact of the Northern Hemisphere ice sheets, because reference to Fig. 5 shows that they were then still close to their maximum LGM volume. Instead the chronologies reviewed here afGeografiska Annaler · 81 A (1999) - 2 INTERHEMISPHERIC LINKAGEOF PALEOCLIMATE DURINGTHELASTGLACIATION ford strong circumstantial evidence that this decisive warming was caused by an abrupt switch of thermohaline circulation to the modern mode of operation, with renewed downwelling in the Nordic Seas. Because near-interglacial warmth was also achieved in middle latitudes of the Southern Hemisphere, the decisive switch of thermohaline circulation must also have triggered increased production of atmospheric water vapor to near-interglacial values. As well as having a strong influence on high-latitude Northern Hemisphere climate (Lehman and Keigwin 1992b), renewed NADW production may also have caused reduction of Southern Ocean sea-ice extent and the southward shift of the westerlies by the standard explanation for coupling the hemispheres by thermohaline circulation (Weyl 1968). Such speculation is consistent with the Taylor Dome but not with the Byrd and Vostok ice-core records from Antarctica near the Oldest Dryas/B611ingtransition, as discussed in the next section. What caused the resumption of vigorous North Atlantic thermohaline circulation with a strong LNADWcomponent near the Oldest Dryas/B6lling transition? The radiocarbon chronologies given here are consistent with the idea that the initial Oldest Dryas warming step triggered this decisive event of the last termination. But how? The fact that the termination began when ice sheets achieved their maximum volume as recorded by the 6180 signal in benthonic foraminifers (Fig. 5) (Shackleton et al. 1988) suggests that the existence of large ice sheets was a necessary condition for the initiation of the last termination. The Heinrich 1 event is also implicated during the most recent termination, because it occurs right at the time of the break in the fundamental benthonic 8180 trends in core T16331B in Fig. 5. To explore this last implication, we turn to a discussion of Heinrich events in the North Atlantic Ocean. Massive, short-lived discharges of icebergs into the North Atlantic Ocean occurred each 700010,000 years during the gradual buildup phase of the last 100,000-yr cycle (Heinrich 1988). These outbursts left prominent Heinrich layers of icerafted debris with sharp lower boundaries deposited rapidly on the sea floor along the southern margin of the glacial-age North Atlantic Ocean (Broecker et al. 1992; Bond et al. 1992, 1993; Bond and Lotti 1995; Manighetti and McCave 1995; Manighetti et al. 1995). Heinrich deposits are marked by high percentages of ice-rafted detriGeografiska Annaler · 81 A (1999) · 2 tus and low concentrations of foraminifers. They become thinner from west to east across the Atlantic Ocean, and commonly are rich in detrital carbonate. These characteristics point to a source in eastern Canada or western Greenland, with Hudson Strait a prime candidate, for most of the debris in the main iceberg track. Ice-rafted grains from the penultimate Heinrich layer deposited about 20,500 14C yr BP show a lead isotopic composition also consistent with derivation of ice-rafted debris from eastern Canada (Gwiazda et al. 1996). Revel et al. (1996) argued that Scandinavian, British, and Icelandic ice caps contributed to the flux along the eastern part of the main iceberg track, as well as to the north of the main track. They thus suggested that all ice caps surroundingthe North Atlantic, not just the Laurentide Ice Sheet, experienced major discharge of icebergs at the time of Heinrich events. This is consistent with the conclusions drawn from the 6180 record of a core taken northwest of the Faeroe Islands (Rasmussen etal. 1997). From two important cores at the eastern end of the maximum iceberg track, Bond and Lotti (1995) likewise found evidence of synchronous discharges of icebergs from several ice sheets during Heinrich events. From a high-resolution marine-sediment record from the Irminger Basin, which registers ice-rafted debris from the Greenland and Norwegian Seas, Elliot et al. (1998) concluded that lithic layers corresponding to Heinrich events were deposited by enhanced calving from Nordic ice caps, as well as the Laurentide Ice Sheet, over 20° of latitude in the North Atlantic Ocean. A comparison of Greenland and North Atlantic paleoclimate records reveals that Heinrich events occurred at or near the culmination of Bond cooling hemicycles superimposed on the overall buildup phase of the last 100,000-yr cycle (Bond et al. 1993; Rasmussen et al. 1997). Following each Heinrich event, the sea-surface temperatures warmed as the North Atlantic Ocean made a short but failed excursion toward the interglacial mode of circulation. The situation is even more complex, because millennial-scale Dansgaard-Oeschger oscillations are superimposed on each cooling hemicycle (Bond et al. 1993). The cause of the huge discharges of icebergs during Heinrich events is not clear. Thus such a discharge could be a trigger for climate change, a response to climate forcing, or both (Broecker et al. 1993; Broecker 1994, 1995a, b). There are at least three possible explanations for these massive iceberg discharges into the North Atlantic Ocean. One 133 G.H. DENTON ET AL. is from internally triggered surges of the Laurentide Ice Sheet (Broecker et al. 1992; MacAyeal 1993). This surge mechanism has the advantage of explaining rapid deposition of Heinrich layers. One apparent disadvantage is that clear geologic evidence of down-trough flow of an ice stream draining the heart of the Laurentide Ice Sheet has not been found in Hudson Strait (Miller et al. 1993). Another disadvantage is that several ice sheets, not just Laurentide ice, apparently had an increased iceberg flux at the time of Heinrich discharge events (Bond and Lotti 1995; Fronval et al. 1996; Revel etal. 1996; Rasmussen etal. 1997; Elliot et al. 1998). A second potential origin for the Heinrich icebergs is simply increased calving from expanded ice sheets on North Atlantic continental shelves during maxima of Bond cycles. The main disadvantage of this mechanism is that it cannot easily account for the catastrophic outbursts of icebergs implied by the sharp lower boundaries of the Heinrich ice-rafted debris that buried organic horizons on the ocean floor within a few hundred years and accumulated so rapidly that it is only little disturbed by bioturbation (Manighetti and McCave 1995; Manighetti et al. 1995). A third potential source for a massive iceberg outburst is from collapse of unstable grounded marine ice on North Atlantic continental shelves, with associated ice-sheet downdraw by accelerated discharge of feeder ice streams. For example, Fronval et al. (1996) recently argued that those portions of ice sheets grounded on extensive shelf areas would be unstable, subject to collapse by marine downdraw. This suggestion is very similar to the mechanism previously proposed by Ruddiman and McIntyre (1981) to explain in many North Atlantic marine sediment cores an interval nearly barren of planktonic foraminifers. This interval, now known to be equivalent to the Heinrich 1 event, was interpreted to result from the most massive influx of icebergs and meltwater to the North Atlantic Ocean during the entire 10,000-yr deglaciation. From a consideration of the mechanics of the last deglaciation by Denton and Hughes (1981), this massive early iceberg influx was attributed to the collapse of unstable marine-based ice sheets on continental shelves, with interior downdraw of feeder ice streams (Ruddiman and McIntyre 1981). An important focus of marine downdraw was Hudson Strait (Denton and Hughes 1981), a potential source of detrital carbonate. However, a difficulty again involves the lack of 134 evidence for ice flow eastward through Hudson Strait (Miller et al. 1993). By the scenario of unstable marine collapse, each cooling Bond hemicycle superimposed on the long buildup phase of the 100,000-yr cycle involved expansion of marine-based ice onto continental shelves to reach a maximum at the beginning of a Heinrich event. Each cooling hemicycle then ended with collapse of destabilized ice from these shelves during a Heinrich event. Although marine ice-sheet collapses could be triggered internally, it is unlikely that all ice caps would then collapse simultaneously. Rather, as pointed out by Revel et al. (1996), the fact that not only the Laurentide Ice Sheet, but also the Greenland Ice Sheet, the Icelandic ice cap, the Scandinavian Ice Sheet, and the Irish-Scottish ice cap, were all sources of enhanced calving during the time of Heinrich events points to common external forcing. A warming pulse and/or sea-level rise could initiate a near simultaneous collapse of extensive marine-based sectors of ice sheets on North Atlantic continental shelves, thus explaining iceberg outbursts from all ice caps during the time of Heinrich events. We prefer the thirdoption because it is consistent with several aspects of our Chilean paleoclimate record. The chronologic data in Table 1 suggest that the last five Heinrich lithic events correspond with Chilean grass maxima. The implication is that the Bond hemicycles represent global cooling that not only drives large ice sheets onto North Atlantic continental shelves, where they are potentially unstable, but also causes Chilean mountain-glacier advance. The last mountain-glacier maximum in Chile occurs just at the beginning of the H-1 lithic event. The ensuing collapse of mountain glaciers, along with the invasion of the southern Chilean Lake District by Nothofagus and thermophilic tree species, is coeval with deposition of much of the H1 lithic layer. This situation suggests that major recession of Chilean mountain glaciers was coeval with the unstable collapse of grounded ice from North Atlantic continental shelves that produced most of the icebergs responsible for the H-1 lithic layer. The pollen record shows that climate warming was the trigger for the Chilean glacier collapse. Retreat of terrestrialScandinavian ice and of European alpine glaciers suggests that warming may also have triggered the marine ice-sheet collapse from North Atlantic continental shelves. We also prefer the third option because it features an external trigger of marine collapse mechanisms that would act nearly simultaneously on all Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCI MATE DURING THE ILASTGILACIATION expanded marine margins. It has long been suggested that marine-based sectors of large ice sheets could be susceptible to very rapid grounding-line recession from hydrostatic instability triggered by sea-level rise or climate warming (Weertman 1957, 1961, 1974; Denton and Hughes 1981; Denton et al. 1986; Hughes 1987). A potential climate trigger for the youngest Heinrich collapse is documented in the Oldest Dryas paleoclimate record in both hemispheres. As just mentioned, the terrestrial margin of the Scandinavian Ice Sheet, as well as the glacier system of the European Alps, receded extensively during Oldest Dryas time, coincident with the youngest Heinrich collapse. As also just mentioned, our South American data show that Oldest Dryas climate warming coincided with much of the youngest Heinrich lithic layer. This warming in southern South America is the most extensive since sometime prior to MIS 4 (Heusser et al. 1999). It is mirroredby similar warming in New Zealand (Newnham et al. 1989). Thus the Oldest Dryas climate warming that caused recession of terrestrial glaciers in both hemispheres could also have triggered unstable collapse of marine icesheet sectors on North Atlantic continental shelves. Such a collapse would have kept the North Atlantic cold, both from the influx of icebergs and from the consequent suppression of thermohaline downwelling and hence northwardocean heat transport. This could explain a very curious aspect of the Greenland ice-core 18sOrecord, namely the striking absence of any abrupt warming signal until near the Oldest Dryas/Boiling transition. In this way, persistence of cold conditions in Greenland during Oldest Dryas time could reflect a massive influx of icebergs into the North Atlantic Ocean. In contrast to the stable isotope record, the first rise of methane in the Greenland ice cores, probably indicative of climate change in the tropics, occurred about 17,000 cal. yr BP (Chappellaz et al. 1993) and hence was correlative with the initial Oldest Dryas warming pulse recognized in the southern Chilean Lake District, New Zealand, and Europe. The differences in two kinds of paleoclimate signals from core SU81-18 from the eastern North Atlantic Ocean off Portugal support the interpretation that regional cooling from the Heinrich 1 ice collapse dominated the Greenland isotope record in Oldest Dryas time. The 5180 signal of the planktonic species Globigerina bulloides in SU81-18 shows a marked depletion beginning at 14,500 14C yr BP (Fig. 6) (Bard et al. 1987), compatible with the timing of the Heinrich 1 ice collapse. And yet Geografiska Annaler · 81 A (1999) · 2 sea-surface temperatures in SU81-18 reconstructed from micropaleontological transfer functions show a significant drop in Oldest Dryas time. Again, this is consistent with decisive cooling of the North Atlantic Ocean from the flood of Heinrich 1 icebergs and the consequent collapse or nearcollapse of the glacial mode of thermohaline overturn (Sarnthein et al. 1994). Yet another reason for preferring the third option is that deglaciation of marine-based sectors of both the Scandinavian (Lehman et al. 1991; Haflidason et al. 1995) and Barents Sea (Bischof 1994; Svendsen et al. 1996) Ice Sheets coincided with the Heinrich 1 ice-rafting event in the North Atlantic Ocean. In core V-2381 in the North Atlantic, Heinrich ice rafting began about 14,800 14C yr BP, culminated at close to 14,100-14,300 14Cyr BP, and was in rapid decline after 13,700 14Cyr BP (Bond and Lotti 1995). All this is consistent with the initial reduction of Northern Hemisphere ice volume as deduced from the benthonic 6180 record in eastern Pacific core TR163-31B (Fig. 5) (Shackleton et al. 1988). The consequence of selecting this third option is that the magnitude of each Heinrich collapse could well be linked to a combination of ice-sheet size and of the intensity of the warming trigger. The final Heinrich collapse that peaked at 14,100 14Cyr BP could then be the largest, because by this time the ice sheets had reached their maximum volume of the last 100,000-yr cycle (Fig. 5). The key to the last termination is that ice sheets had to grow sufficiently large to produce a Heinrich collapse massive enough to cripple the glacial mode of the Atlantic salinity conveyor. The conveyor circulation then reorganized into its interglacial mode of operation. The probable reason for this reorganization is that the massive collapse during the time of the Heinrich 1 event flushed out so much marine-based ice peripheral to the North Atlantic Ocean that, at the end of the collapse, the iceberg flux dropped dramatically. Through the long buildup phase, the ice-sheet behavior during Bond cycles could be an important regulator of North Atlantic near-surface salinity. The steady background iceberg influx during buildup could keep North Atlantic salinity low. A Heinrich collapse could produce the lowest salinity at the peak of each cycle. North Atlantic salinity could then jump after each Heinrich collapse, thus triggering thermohaline switches and a rise of North Atlantic sea-surface temperatures. The largest salinity jump would follow the massive Heinrich 1 event, because it flushed out the most ice. At the same time the sum135 G.H. DENTON ET AL. mer radiation field from orbital forcing was approaching an interglacial value, which might favor evaporation, and hence net export of water vapor, from the northern North Atlantic Ocean. Perhaps these two factors together trip the North Atlantic into its interglacial circulation mode, with a strong LNADW component. By this scenario, the resumption of the interglacial mode of North Atlantic thermohaline circulation occurs because of, not in spite of, the growth of huge ice sheets peripheral to the North Atlantic Ocean. The unstable behavior of marine shelf portions of these ice sheets is the key ingredient. The reason that ice sheets have to grow to such huge dimensions is to produce an unstable collapse large enough to deplete the grounded marine ice reservoir and its feeder ice streams. Once the resulting massive iceberg influx ceases, the North Atlantic becomes largely free of icebergs for the first time since early in the long buildup phase of the 100,000-yr asymmetric cycle. North Atlantic salinity can then rapidly increase and trigger renewed vigorous downwelling in the Nordic Seas. This explains the curious observation that terminations occur just when ice sheets achieve their largest volume. It also is consistent with the observation that a prerequisite for a sharp, complete termination seems to be excessive ice volume in huge Northern Hemisphere ice sheets (Raymo 1997). What caused the asymmetric 100,000-yr glacial cycles of late Quaternary time? Terminations are a dominant feature of these cycles. From radiocarbon-dated paleoclimate sequences, we argued above that renewal of the modern mode of NADW formation, with a strong LNADW component, was the key event of the last termination. From this we assume that the current mode of thermohaline circulation is the essential feature of late Quaternary interglaciations. We now discuss the implications of this assumption. The modern mode of thermohaline overturn in the northern North Atlantic Ocean switched on abruptly near the Oldest Dryas/B61iing transition. This event was accompanied by rapid reforestation of Europe, including the northward spread of thermophilic trees, presumably because of the demonstratedlink between North Atlantic heat import and European climate. Even more-extensive forest cover occurred in northern Europe during Eemian (MIS 5e) time (Behre 1989), consistent with vigorous formation of LNADW from the inflow of warm, salty surface water. This is in accord with a 136 paleoceanographic reconstruction of the Norwegian and Greenland Seas (Kellogg 1980). Deep-water temperatures of MIS 5e were similar to those of the Holocene (Labeyrie et al. 1987). Deep-water temperature in the Pacific (Chappell and Shackleton 1986; Labeyrie et al. 1987) and southern Indian (Labeyrie et al. 1987) Oceans dropped to glacial values about 115,000 years ago during the MIS 5e/5d transition. Deep Atlantic waters reached glacial values in two major steps, one at about 115,000 years ago and the other at about 75,000 years ago during the MIS 5/4 transition (Labeyrie et al. 1987). After these cooling steps, the deep ocean waters remained at glacial temperatures throughout MIS 3 and MIS 2 (Chappell and Shackleton 1986; Labeyrie et al. 1987). The paleoenvironment of northern Europe (Beaulieu and Reille 1984, 1989, 1990, 1992; Behre 1989; Guiot 1990; Seret et al. 1992) suggests that these cooling steps of deep-water temperature could well reflect the progressive shutdown of warm LNADW early in the last glacial cycle. At the close of the Eemian, a cooling episode that left an environment markedonly by shrubtundrain northern Europe during the Herning stadial (Behre 1989) implies sharp curtailment of warm LNADW formation as the primary cause of the deep-water temperaturedecline about 115,000 years ago. During the subsequent Br6rup (MIS 5c) and Odderade (MIS 5a) European interstades, each lasting as much as 10,000 years, northern Europe was reforested, with coniferous forests in the north giving way to deciduous forests in the south (Behre 1989). The inferred north-to-south climate gradient was distinctly steeper than that of the Eemian (Behre 1989). During MIS 5a the Norwegian and Greenland Seas were seasonally ice covered, with the formation of LNADW in smaller amounts than today (Kellogg 1980), in accord with the Odderade paleovegetation reconstruction. The onset of cold Pleniglacial conditions in northern Europe immediately after the Odderade interstade (Behre 1989) is consistent with the shutdown of warm LNADW as the cause of the second major cooling step in North Atlantic deep water at the MIS 5/4 transition about 75,000 years ago. Warming sufficient to cause reforestation in northern Europe did not occur during the cold Pleniglacial climate conditions of MIS 3 and MIS 2, even at the peaks of Greenland ice-core interstades 8 (Denekamp in Europe), 12 (Hengelo in Europe), 14 (Glinde in Europe), and 16 (Oerel in Europe) (Kolstrup and Wijmstra 1977; Behre 1989; Guiot Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATEDURING THE LAST GLACIATION 1990; Pons et al. 1992; Schulz et al. 1998). Rather, these interstades were marked only by reversion from tundra to open, treeless shrub tundra (Behre 1989). There probably was some thermohaline downwelling near the Faeroe Islands in these interstades (Rasmussen et al. 1997), but the lack of reforestation in northernEurope suggests that it must have been very weak. Continuous cold, treeless conditions in Europe during MIS 3 and MIS 2 are consistent with a lack of LNADW formation from warm, salty water and, consequently, with continuous cold deep-water temperatures. These considerations suggest that the modem mode of North Atlantic thermohaline circulation switched off in two steps early in the last glacial cycle when ice sheets were still small. It did not switch back on again until near the Oldest Dryas/B61iing transition, when ice sheets were near their maximum LGM volume. The result is that early in the last glacial cycle the climate system became largely detached from orbital forcing except for oscillations superimposed on the long buildup to the LGM (see also Imbrie et al. 1993) (Fig. 5). The formation of LNADW could have been switched off early in the last cycle by orbital forcing of ice-sheet growth and the consequent iceberg influx into the critical areas of downwelling in the Nordic Seas. But subsequent orbital forcing in the opposite sense was not sufficient to switch it back on. Instead, an extraordinary mechanism must have operated in the climate system in order for the North Atlantic salinity conveyor to switch abruptly into a vigorous interglacial mode of operation at what seems to be a very unlikely time, namely, just when Northern Hemisphere ice sheets were close to their maximum LGM volume. We argue above that, rather than precluding a circulation switch to an interglacial mode, the growth of these ice sheets to their maximum volume was actually a prerequisite for the decisive ocean-atmosphere reorganization near the Oldest Dryas/B611ing transition, because it set up the necessary conditions for the huge Heinrich 1 ice-sheet collapse into the North Atlantic Ocean, which we think is the key precursor event. The marine-based portions of ice sheets grounded on North Atlantic continental shelves exhibited unstable collapse behavior at the culmination of each of the shorter Bond cycles superimposed on the long growth phase of the last asymmetric 100,000-yr cycle. This long and fluctuating growth phase ended abruptly when ice sheets became large enough to produce an unstable collapse sufficiently Geografiska Annaler 81 A (1999) 2 massive not only to cripple the glacial mode of North Atlantic thermohaline circulation, but also to flush out grounded continental-shelf ice and feeder ice streams adjacent to it. As a result, the influx of icebergs into the North Atlantic declined greatly once the collapse was over. In the absence of an iceberg influx and of conveyor circulation, the salinity of the northern North Atlantic Ocean increased rapidly, probably from net export of water vapor, until a vigorous interglacial mode of LNADW formation was initiated. By this scenario, the length of an individual 100,000-yr cycle is set by two main factors that work in combination to produce the massive marine ice-sheet collapse that resets the North Atlantic salinity conveyor. The first factor is the length of time required to accumulate the excess ice volume necessary for a large collapse. The second is the timing of the warming trigger for this collapse, presumably set by the effect of eccentricity on the amplitude of precession and half-precession insolation. The timing of the warming trigger, and hence of the collapse, need not be unique. For example, if the Heinrich 1 collapse had been too small to reset the salinity conveyor, then a later collapse could have triggered the termination at about 12,000 years ago (Northern Hemisphere summer insolation maximum). In this way terminations can occur at any of several times within intervals of high eccentricity. Thus the role of orbital forcing of terminations is to cause the climate warming that triggers a massive ice-sheet collapse and then, from high summer insolation, to impose a precipitation/ evaporation ratio that promotes this interglacial circulation mode by the net export of water vapor from the northern North Atlantic Ocean. This is compatible with the argument given earlier that, during terminations, insolation forcing simply acts as a trigger for the fundamental reorganization of a non-linear system, but does not control the amplitude of the resulting reorganization. From the paleoclimate record, we argue above that the critical step of the last glacial/interglacial transition was the resumption of the modern mode of NADW formation, with a vigorous LNADW component. We also argue that it took the entire buildup phase of each asymmetric glacial cycle to produce ice sheets large enough to reset the modern mode of global thermohaline circulation through a massive Heinrich marine-ice-sheet collapse. The notion that it is so difficult to turnon the interglacial mode of thermohaline circulation suggests that 100,000-yr glacial cycles emerged because the 137 G.H.DENTONETAL. modem interglacial mode of thermohaline circulation became easier to switch off and harder to switch on than it was prior to the 100,000-yr regime. But what caused the development of such an asymmetric behavior of the interglacial mode of thermohaline circulation? For a possible answer we turn to the topography of the Greenland-Scotland submarine ridge across the North Atlantic Ocean. Wright and Miller (1996) postulated that the changes in the height of the Greenland-Scotland ridge (from varying activity of the Icelandic mantle plume) regulated the Neogene flow of northward-flowing warm surface water into the Nordic Seas and hence controlled NADW circulation. Rise of this ridge from mantle-plume activity is postulated to have brought mid-Pliocene Arctic warmth to a close (Wright and Miller 1996). A further rise of this ridge between 950,000 and 600,000 years ago could have caused the asymmetric behavior of North Atlantic thermohaline circulation that we suggest was characteristic of late Quaternary time. This would have initiated asymmetric 100,000-yr climate cycles because of the necessity for the buildup of large ice sheets to reset the interglacial mode of thermohaline circulation by unstable collapse from continental shelves. What caused the climate cooling reversal thatfollowed peak Bolling warmth and culminated in the YoungerDryas cold pulse? Any explanation must address the 1000 14C-yr length of the Younger Dryas cold pulse and its occurrence at very near the peak of maximum Northern Hemisphere summer insolation. Also, it should clarify why sea level continued to rise throughout this climate deterioration and even during the culminating Younger Dryas cold pulse (Fairbanks 1989; Bard et al. 1996). If our paleoclimate reconstructions from Chilean pollen records are correct, then peak Bollingage warmth was short-lived, and was followed by a climate reversal that persisted through much of late-B61ling and Aller6d time in both hemispheres. Also, if our reconstructions from the Chilean Andes and Southern Alps, along with the reconstruction of Ariztegui et al. (1997) from the Argentine Andes, are correct, then a Younger-Dryas-age cold event at the end of this long deterioration may have marked both hemispheres. The radiocarbon dates of the Younger Dryas readvance of Franz Josef Glacier in the Southern Alps of New Zealand (11,050 14C yr BP; Denton and Hendy 1995) and of the Younger Dryas isotope shift in Switzerland 138 (11,030 14C yr BP; Hajdas et al. 1995) imply rapid transmittal of an atmospheric cooling signal at the beginning of Younger Dryas time. The nearly equivalent amount of snowline lowering in the European Alps and the Southern Alps suggests that the Younger Dryas event represented a reversal of about 35% toward full-glacial conditions (IvyOchs et al. 1999). This suggestion must be considered tentative, however, until more Southern Hemisphere data on Younger-Dryas-age climate events are available (for alternative view see Markgraf 1991,1993; Mabin 1995; McGlone 1995; Singer et al. 1998). A leading explanation for the basic shape of the late-glacial climate signal is that the North Atlantic salinity conveyor turned on at maximum strength near the Oldest Dryas/B611ing transition and subsequently weakened by dilution of salt from melting of continental ice sheets caused by the peak Boiling warmth. The increasingly sluggish thermohaline overturn culminated in Younger Dryas shutdown-or near shutdown-of LNADW production, probably triggered by an additional pulse of icebergs and/or meltwater (Broecker 1990, 1992; Broecker et al. 1990; Fanning and Weaver 1997). New modeling studies of the stability of North Atlantic thermohaline circulation show that cooling and wind-stress feedbacks destabilize those modes of circulation without deepwater formation (Schiller et al. 1997). In the modeling experiments deep-water formation resumes soon after meltwater input ceases. The implication is that only a meltwater pulse lasting 1000 14C yr could have produced the North Atlantic Younger Dryas event. Two sources have been suggested. One is diversion of North American glacial lakes from the Mississippi drainage into the Gulf of St Lawrence drainage (Rooth 1982; Broecker et al. 1988, 1989), or the Baltic Ice Lake into the North Atlantic (Bj6rck et al. 1996). Potential problems are that freshening of the Champlain Sea apparentlydid not occur until about 500 14Cyr after the beginning of the Younger Dryas (Rodrigues and Villas 1994) and that a distinct 6180 minimum has not been found off Nova Scotia (Keigwin and Jones 1995). A second potential source of freshwater is an iceberg influx from the Arctic due to the collapse of floating ice shelves and of ice sheets grounded on high-latitude continental shelves (Mercer 1969). The available uplift curves from Svalbard, Franz Josef Land, and the Queen Elizabeth Islands are at least permissive of this hypothesis (Blake 1975, 1993; Forman et al. 1996, 1997). The collapse of marine ice sheets Geografiska Annaler * 81 A (1999) - 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION couldeasily encompass1000 14Cyr andcouldexplain continued sea-level rise during Younger Dryastime. In any case, our New Zealanddata suggest a SouthPacificatmosphericsignalrightatthebeginning of YoungerDryas time. If a NorthAtlantic thermohalineswitch is the ultimatecause of the YoungerDryas reversal,then the atmosphericeffects of this switchmusthavebeentransmittedrapidly,bothto the tropics(Robertset al. 1993;Hughen et al. 1996; Clappertonet al. 1997) and to the South Pacific region (Denton and Hendy 1994). The new trapped-gaschronologiesof theByrdicecore recordin Antarcticasuggest that changes of atmosphericCO2contentwerenottheoriginof this atmosphericeffect (SowersandBender1995;Blunier et al. 1997). This again leaves a linkage between thermohalinecirculationand the tropical productionof watervaporas themostlikely source of this atmosphericsignal. Did fluctuations in the strength of NADW cause out-of-phase climate variations in the Northern and Southern Hemispheres during the last glacia- tion? Crowley (1992) suggested that changes in oceanicheattransferacrossthe equatorcausedby variationsin the productionof NADW would alternatelyfavornetheatgainin one andthentheother hemisphere.This is becausethe amountof heat transportedto Antarcticaby NADW would be morethancompensatedfor by the heattransferred across the equatorby northward-flowing shallow currentsin the Atlantic Ocean. During times of strong NADW production,the SouthernHemisphere would lose heat. The reverse situation wouldoccurduringtimesof weakproduction.The middle-latitudepaleoclimaterecords given here suggest thatthis mechanismdid not dominateatmosphericclimateoscillationsduringthe last glacial/interglacialtransition.Middle-latitudeChilean climatewarmedin OldestDryastime when the productionof NADWreachedits lowest value,but What is the role of millennial-scale oscillations in glaciersalso recededin the NorthernHemisphere producing abrupt climate changes? A pervasive at the same time. Moreover,middle-latitudeChilmillennial-scaleoscillation that persists through eanclimatewarmedalmostto full-interglacialvalmajorclimate transitionshas been recognizedin ues coincidentwitha switchto themodernmodeof Holocene,late-glacial,andglacialrecords(Denton NADW formationnear the Oldest Dryas/B611ing and Karlen 1973; Karlen and Denton 1976; transition. O'Brienetal. 1995;Bondetal. 1997).Infact,Bond et al. (1997) suggestedthatmillennial-scaleoscillationsarethe pacemakerof rapidclimatechange. Southern Ocean sediment cores and Antarctic In this regard,the timingof abruptinitialwarming ice cores in theChileanrecordmayhavebeenpacedandam- Ourradiocarbonchronologyof pollenprofilesand plifiedby themillennial-scaleoscillationssuperim- morainesshows that the estimatesof 875 m for posedon theLGM.Forexample,thelastmajormil- mountainsnowlineloweringin the SouthernAlps lennial-scaleglacierpulse reacheda maximumat of New Zealand(Porter1975)andof about1000m 14,550-14,805 14Cyr BP, just priorto the abrupt for snowline (Porter1981) and treeline lowering warmingin OldestDryastimethattriggeredthelast (Morenoet al. 1999; Heusseret al. 1999) in the termination.Althoughnot documentedin Chile, a southernChileanLake Districtpertainto glacier similar situation occurredlater in the Northern maximawithinthe long, cold intervalof the LGM. Hemisphere,where the PortHuron(GreatLakes) Thesevaluesof snowlineloweringareverysimilar and Luga (north-centralEurope) ice-sheet read- to those in many mountainranges elsewhere on vances immediatelypreceded the abruptOldest Earth(BroeckerandDenton 1990). Such uniform Dryas/B611ing warming(e.g. Denton and Hughes loweringof mountainsnowlinesstronglysuggests 1981).These relationshipssuggestthatmillennial- globalcoolingof the samemagnitudein bothhemscale pulsesmay be importantin settingthe timing ispheresduringthe LGM. From our middle-latiof abruptwarmingsteps. For example, the cold tude SouthernHemispheredata,we arguethatthe pulse at 14,550-14,805 14Cyr BP could have de- LGM was terminatedby several abruptclimate layed the onset of the last termination.Then the changes that are registeredin both hemispheres. subsequentwarm phase of the millennial-scale Most importantarethose duringthe OldestDryas transition.Topulse could have reinforcedabruptOldest Dryas and nearthe Oldest Dryas/B611ing climatewarming.Thus the timingand abruptness getherthese changesbroughtthe planetfromfullof the Oldest Dryas/B11lingwarmingcould have glacial to near-interglacialtemperaturesin about been affectedby a millennial-scaleoscillation. 1600 14Cyr. Geografiska Annaler * 81 A (1999) · 2 139 G.H.DENTONETAL. We argue that neither the global cooling of the LGM, nor the abruptglobal warming steps that terminated the last glaciation, can be explained simply by redistribution of heat on the planet from ocean circulation changes. Rather,either the reflectivity of the planet or the greenhouse gas content of the atmosphere had to change. Because of the basic similarities of the Chilean and New Zealand paleoclimate records with the classic North Atlantic/ European record, we stress changes in atmospheric greenhouse gas content (probably predominantly the water vapor inventory, somehow linked to thermohaline switches), as the major mechanism of interhemispheric climate linkage. During the decisive ocean-atmosphere reorganization near the Oldest Dryas/B611ing transition, a switch in thermohaline circulation must have been connected with a change in the production of water vapor. During the precursor Oldest Dryas warming a change in atmospheric water-vapor content was not obviously connected with a thermohaline circulation switch but was probably linked to precession effects in the tropics. But there is a potential problem with our approach. Namely, it has been suggested for more than two decades that sea-surface temperature change in the Southern Ocean leads ice-volume change in the Northern Hemisphere into and out of interglaciations (Hays et al. 1976; Hays 1978). This inference comes from offsets in individual marine sediment cores of the 6180 values of benthonic and planktonic foraminifers and the sea-surface temperature estimated from diatoms, foraminifers, and radiolaria (Howard and Prell 1984, 1992; Labeyrie etal. 1986,1996; Labracherie etal. 1989; Pichon etal. 1992). For example, sea-surface temperature is taken to lead ice-volume change by several thousand years during the transition into the current interglaciation. In the absence of benthonic 6180 change, the early Holocene peak in Southern Ocean sea-surface temperatures is taken to indicate a lead of Southern compared to the Northern Hemisphere climate in the transition out of the current interglaciation (Hays 1978). To complicate the situation, the stable isotope record calibrated with new trapped-gas chronologies from the Byrd ice core in Antarctica does not show the abrupt changes during the last termination so evident in the North Atlantic (Fig. 7). Instead, a gradual change in the stable-isotope signal in the Byrd ice core began several thousand years before the first abrupt warming registered in Greenland ice cores at the Oldest Dryas/Bolling 140 transition (Sowers and Bender 1995; Blunier et al. 1998). Also, late-glacial stable isotope oscillations recorded in the Byrd and Vostok ice cores during Bolling/Aller6d and Younger Dryas times were out of phase with their Greenland counterparts(Fig. 7) (Sowers and Bender 1995; Blunier et al. 1997, 1998). These results are taken to be consistent with Southern Ocean marine cores in showing a Southern Hemisphere lead through important climate transitions. This Antarctic lead seemingly precludes any hypothesis that climate changes are a response to Northern Hemisphere events (Blunier et al. 1998). The asynchronism strongly suggests, moreover, that the interhemispheric connection cannot be through the atmosphere. Rather, it favors models that call for overall ocean heat extraction from the Southern Hemisphere during times of vigorous North Atlantic thermohaline circulation (Crowley 1992; Stocker et al. 1992). Because they suggest that any Antarctic asynchrony is not hemisphere-wide, our paleoclimate data from Chile and New Zealand are not in accord with an ocean heat-transfermechanism that applies to the entire hemisphere. Moreover, a heat-transfer mechanism by itself cannot explain the overall warming of both hemispheres during the last deglaciation. One way around this dilemma is to postulate that the asynchrony is confined to regions south of the Antarctic Circumpolar Current, thus pointing to a mechanism that applies only to Antarctica rather than to the Southern Hemisphere as a whole. In this regard, Broecker (1998) attributed out-of-phase late-glacial climate oscillations between Greenland and Antarctica to a bipolar seesaw behavior of thermohaline circulation. By this mechanism, decreased (increased) downwelling of NADW in the North Atlantic Ocean was replaced by increased (decreased) downwelling in the Southern Ocean. The resulting out-of-phase import of ocean heat into the North Atlantic and Southern Oceans is what would have caused out-of-phase late-glacial climate signals in the Greenland and Antarctic ice cores. Broecker (1998) and Broecker and Henderson (1999) suggested an early trigger of thermohaline overturn in the Southern Ocean from local summer insolation during at least the last two terminations. A possible greaterAntarctic lead during the penultimate than during the last termination may reflect stronger early insolation forcing (Broecker and Henderson 1999). But there are also complications with this approach. One is that almost all the important marine sediment cores showing a lead of Southern Ocean Geografiska Annaler * 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION Fig. 8. Index map of the Southern Ocean showing the location of major oceanic boundaries, including the Subtropical (STF), Subantarctic (SAF), Polar (PF), and southern Antarctic Circumpolar Current (SACCF) Fronts. Deep-sea sediment cores MD88-770 (46°01'S, (41°36'S, 96°28'E), RC11-83 9°48'E), MD84-527 (43°49'S, 51°19'E), and DSDP 594 (45°32'S, 174°57'E) as well as the Taylor Dome (77°47'S, 2400 m), Byrd (80°01S, 1530 m), and Vostok (78°28'S, 3488 m) ice cores, are also shown, as well as many of the pertinent Southern Ocean cores discussed in the literature. Frontal boundaries and terminology are after Orsi et al. (1995). lie northof the PolarFrontin Subantemperatures tarctic waters (Fig. 8). Hence these cores come from nearly the same SouthernHemispherelatitudes as our terrestrialdata,which we interpretas showingpaleoclimatechanges similarto those in the North Atlantic/Europeansector of the planet transition.A secduringthelastglacial/interglacial ond is thatthe newTaylorDome ice core (77°47'S, 2400 m) (Mayewskiet al. 1996;Steig et al. 1998), situatedin Antarcticaabouthalfwaybetween the Byrd(80°0I'S, 1530m) andVostok(78°28'S,3488 m) ice cores (Fig. 8), showsa late-glacialpaleoclimaterecordsimilarto thatin Greenlandandin the middlelatitudesof the SouthernHemisphere.For example,unlike the Byrd or Vostokrecords,that from the TaylorDome shows an abruptwarming near the Oldest Dryas/B611ingtransitionand a YoungerDryas reversal (Mayewski et al. 1996; White and Steig 1998; Lehman1998; Steig et al. 1998). Thustherearenow two fundamentallydifferent paleoclimatesignalsfromAntarcticice cores, one in phaseandthe otherout of phasewith NorthAtlantic and middle-latitudeSouthernHemisphere Geografiska Annaler · 81 A (1999) · 2 records.This has led to yet a thirdexplanationof Antarcticice-corepaleoclimaterecords.Whiteand Steig (1998) andSteig andWhite(1998) suggested thatthe differencesin ice-corerecordsreflectheterogeneityin the upwellingpatternsof NADW in the SouthernOcean. Antarctic CircumpolarTransect A resolution of the differing interpretationsof SouthernHemispherepaleoclimaterecordsis not at hand.A majorproblemis thewidespreadlackof theextensivedatingbasenecessaryto placethedetails of deglaciationon a hemisphericand interhemisphericscale.Weproposethattheissuecanbe clarifiedby obtainingdetailedmarineand terrestrial paleoclimaterecordsalong a proposedAntarctic CircumpolarTransect(ACT) between 42° and 54°S (Fig. 8). The essential feature of this transectalong the outer marginof the Southern Ocean aroundAntarcticawould be detailedAMS radiocarbon chronologies. Thus the proposed transectis designed to lie largely within or just northof Subantarcticwaters(where foraminifers 141 G.H. DENTON ET AL. are common for radiocarbon dating) and to pass across mountainous areas of South America, New Zealand, Tasmania, and South Georgia (where wood and gyttja are common for radiocarbon dating). Comparison of the resulting vegetation, mountain-glacier, benthonic 8'80, planktonic b180, and sea-surface-temperature records should clarify the issue of leads and lags of Southern Hemisphere paleoclimate along the outer margin of the Southern Ocean relative to both high-latitude Antarctica and the Northern Hemisphere. The results should also clarify the issue of a bipolar seesaw of thermohaline circulation, as well as the amplitude of paleoclimate changes in different sectors of the Southern Ocean. The currentsituation along the marine portion of ACT is that three marine sediment cores have an AMS radiocarbon chronology. Fig. 9 shows the dated isotope record for RC11-83 (41°36'S, 9°481E) (Charles et al. 1996). This particular core does not show an obvious lead of planktonic over benthonic 6180 at the initiation of the last deglaciation, placed before 15,000 14Cyr BP largely on the basis of a single date in this interval. The transitory oscillation or plateau of planktonic 6180 during deglaciation is thought to be correlative with the Antarctic Cold Reversal (Billing in age, Blunier et al. 1997) in the Vostok ice-core record (Charles et al. 1996). Such a correspondence could be expected because the subantarctic waters near RC11-83 are the ultimate source of most precipitation in the region of Vostok (Koster et al. 1992). However, the 6180 minimum at about 12,360 14Cyr BP in RC1183 is coincident with the full development of a closed-canopy North Patagonian Evergreen Forest at 12,200 14Cyr BP in the southern Lake District of Chile. Also the subsequent isotope signal is compatible with the reversal in climate trend evident in pollen records in the southern Lake District (Heusser et al. 1999; Moreno et al. 1999). Thus it is difficult to understand how the late-glacial planktonic 5180 signal from RC 11-83 is taken to be correlative with the Vostok record and to show a 1000-yr lead of Southern relative to Northern Hemisphere climate. Marine sediment core MD88-770 (46°01'S, 96°28'E), calibrated with 16 acceptable AMS radiocarbon dates, shows a small lead of planktonic 5180 and summer sea-surface temperatures over benthonic 6180 at the initiation of the last deglaciation (Fig. 9) (Labeyrie et al. 1996). Bracketing radiocarbon dates suggests that surface water began to warm at sometime between 16,780 and 13,500 142 14Cyr BP, whereas benthonic 6180 began to change shortly before 12,150 14C yr BP. The reason for the large delay in response of benthonic b180 relative to the change in marine sediment core TR 163-31 B (Fig. 7) is not known. In any case, interglacial values of sea-surface temperature were reached at MD88-770 by 12,150 14Cyr BP. Marine sediment core MD84-527 (43°49'S, 51 ° 19'E), calibrated by 15 AMS radiocarbondates, together with core MD84-551 (55°00'S, 73° 17'E), calibrated by three AMS dates, shows that the deglacial 6180 decline in the Southern Ocean began after 16,510 and prior to 13,000 14Cyr BP (Fig. 9) (Bard etal. 1990). Between 13,000 and 12,000 14C yr BP, the sea-surface temperatures reached interglacial values (Labracherie et al. 1989; Bard et al. 1990). There followed transitory oscillations, but none is recorded in all indicators and so it is not known if they are global or regional in origin (Bard etal. 1990). The potential value of ACT can be illustrated by examples from the few paleoclimate records now available in the pertinent latitudinal band of the Southern Hemisphere. * The Byrd and Vostok stable isotope records from Antarctica are not proxies for hemispherewide paleotemperature changes during the last glacial-interglacial transition, because they differ significantly from the Taylor Dome, New Zealand, and southern Chilean Lake District paleoclimate signals. * Although Antarctic warming inferred from the stable isotope record at the Byrd ice core seems to have preceded the first abrupt warming in Greenland ice-core records (near the Oldest Dryas/B61ling transition), it does not necessarily follow that Southern Hemisphere climate as a whole led Northern Hemisphere climate into the last glacial/interglacial transition. This is because the Chilean and New Zealand paleoclimate records both show abrupt warming at about 14,600 14Cyr BP that correlates with similar abruptwarming in the Northern Hemisphere and is quite different in timing and character from the gradual early warming documented in the Byrd ice core. Moreover, the abrupt warming in the Greenland ice cores near the Oldest Dryas/B61oingtransition occurred at least 1600 14Cyr after deglaciation began elsewhere in the Northern Hemisphere. * Although the chronology of the last deglaciation in the Southern Ocean remains confusing, all Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION a. MD88-770 SummerSST (°C) 2 - 4 6 8 10I 5 Foraminiferal 6180 (%o) 4 3 2 5700 -<~v g cm- 0. 40 0. , oO-C Diatoms -- ~-\ 8~~~~~~--2 ;/:= ~-> Foraminifers Foraminifers ~<~*~~ - 'f>|- % -,._._ ,' 2150 1,530 13500 216,780 Foraminifers v-" . < -^ -< Diatoms ~ Benthonic Y ~32,790 23 21,610 20,400 1,290 25,160 °25,010 2730,500 ( - -'-37,220 44,620 C. MD84-527 N. pachyderma 1. 6180 (%0) b. RC11-83 E -O 0. *0 0 Draftedby R. D.KellyJr. 1997 - - -0 28,620 - - 30,580 Fig. 9. Foraminiferal records from the three well-dated deep-sea cores in the Southern Ocean. Isotope records from core MD88-770 show a small lead of planktonic 6180 and summer SST over benthonic )180. Interglacial SST values were reached by 12,150 14Cyr BP (Labeyrie et al. 1996). Oxygen-isotope records from RC11-83 do not show an obvious lead of planktonic over benthonic 6180 at the beginning of deglaciation; however, a plateau in planktonic 6180 during deglaciation is thought by Charles etal. (1996) to correlate with the Antarctic Cold Reversal seen in the Vostok Ice Core. Based on gas chronologic comparisons of Byrd and GISP2 ice cores, the Antarctic Cold Reversal is thought to have occurred during Boiling time (Sowers and Bender 1995). In any case, 6180 values typical of interglacial temperatures were reached by 12,360 14Cyr BP (Charles et al. 1996). The third well-dated record comes from MD84527 where i180 values of Neogloboquadrina pachyderma indicate that full interglacial conditions were reached by 12,130 14Cyr BP (Bard et al. 1990; Labracherie et al. 1989). Geografiska Annaler * 81 A (1999) · 2 143 G.H. DENTON ET AL three of the extensively AMS-dated marine sediment cores show near-Holocene values of seasurface temperatures, or else significant depletion of planktonic 6180, at close to 12,00012,500 14Cyr BP.This is in accord with the situation in the southern Chilean Lake District terrestrial pollen records (Fig. 5) (Heusser et al. 1999; Moreno et al. 1999) and the Argentine Andes glacial record (Ariztegui et al. 1997). This warm interval corresponds with peak Bolling warmth in the North Atlantic region (Fig. 5), and it is out of phase with the climate oscillations in the Byrd and Vostok, but not in the Taylor Dome, ice cores in Antarctica (Sowers and Bender 1995; Blunier et al. 1997; Steig etal. 1998). Figure 10 suggests that the Southern Hemisphere summer insolation peak at about 23,000 cal. yr BP(which is nearly out of phase with the precession-forced summer minimum in the Northern Hemisphere) did not cause an early warming response in the New Zealand and Chilean paleoclimate records during Termination I. In contrast, detailed pollen analysis of cores and sections from the Canal de la Puntilla, Llanquihue, Fundo Llanquihue, Bella Vista Bluff, Dalcahue, and Mayol sites together show that the cold, wet conditions of the LGM continued through the interval of high southern insolation right up to the time of the youngest glacial peak at 14,550-14,805 14Cyr BP(Denton et al. 1999; Heusser et al. 1999; Moreno et al. 1999). LGM conditions came to a close only after the culmination of this youngest advance. The same situation holds for New Zealand. In like fashion, reference to Fig. 4 shows that significant early warming is not obvious in the New Zealand paleoclimate record during Termination II. Summary Combined glacial geologic and palynologic data from the Chilean Andes and Southern Alps highlight the following features of middle latitude Southern Hemisphere paleoclimate through the LGM and the last termination. The moraine chronology suggests that full-glacial or near-full-glacial climate conditions prevailed from about 29,400 to 14,550 14Cyr BPin the southern Lake District-Isla Grande de Chiloe region. However, pollen records from Isla Grande de Chilo6 suggest that full-glacial con144 Southern Lake District, Chile (40°30'- 42°25'S; 72o25'- 73045'W) Mean Summer Temperature 0 0) a) (U 10 Insolation (W/m2) -- ·- ------ - ---- Drafted by R. D. Kelly Jr. 1998 Fig. 10. Comparison of our new paleoclimate record for the Chilean Lake District and the New Zealand Southern Alps derived from glacial geologic and palynological data with summer solar insolation at 45°N and 45°S over the last 40,000 years (Berger 1978). A rise in Southern Hemisphere summer insolation beginning 30,000 cal. years ago did not produce signs of early warming in New Zealand and Chilean paleoclimate records. ditions did not set in until approximately 26,000 14C yr BP. * Within this long interval of the LGM, snowline was depressed about 1000 m, relative to presentday values, during glacier advances into the outer Llanquihue-age moraine belt at 29,400, 26,760, 22,295-22,570, and 14,550-14,805 14C yr BP. Additional glacial maxima may have been achieved shortly before 17,800 14C yr BP and again shortly before 15,730 14C yr BP. The coeval lowering of treeline suggests that mean summer temperature was depressed 6-8°C, compared to modern values, during the LGM. * In the Southern Alps, the known glacial advances during the LGM culminated at 22,400, 17,700, 16,200, and 14,000-15,000 14C yr BP. Geografiska Annaler 81 A (1999) · 2 LINKAGEOF PALEOCLIMATE INTERHEMISPHERIC DURINGTHELASTGLACIATION Additional maxima may have occurred shortly before 19,740 14C yr BP, shortly after 18,600 14C yr BP, and in Younger Dryas time. Relative to present-day values, snowline depression during the most extensive advances was about 875 m (Porter 1975). The last glacial/interglacial transition began with a decisive warming at 14,600-14,700 14C yr BP in both New Zealand and the Chilean Andes. Subsequent details are not yet known in New Zealand, except for late-glacial readvances. But a second and decisive warming pulse in the region of the southern Lake District-Isla Grande de Chiloe occurred at 12,700-13,000 14C yr BP. After an interval of near-interglacial warmth, a late-glacial climate reversal set in close to 12,200 14C yr BP and continued at least into the early part of Younger Dryas time. Although strong contrary opinions have been expressed, this reversal of trend may well have culminated in a Younger Dryas event in both the Chilean Andes and the Southern Alps. The similarity in timing and magnitude of middlelatitude snowline lowering with that in the Northern Hemisphere suggests global atmospheric cooling during the LGM. When compared with paleoclimate records from the North Atlantic region, the middle-latitude Southern Hemisphere terrestrial data imply interhemispheric symmetry of the structure and timing of the last glacial/interglacial transition. Particularly prominent are warming pulses beginning at 14,600 14C yr BP (Oldest Dryas) and 12,700-13,00014C yr BP (near the Oldest Dryas-Boiling transition) that together terminated the LGM and brought large regions in both hemispheres to near-interglacial warmth in about 1600 14Cyr. This synchrony of atmospheric temperature changes during both the LGM and the glacialinterglacial transition suggests that shifts of ocean heat across the equator from changing North Atlantic ventilation power (Crowley 1992) was not the major control on interhemispheric atmospheric signals. Rather it implicates changes in greenhouse gas, most likely in the inventory of atmospheric water vapor, as the dominant climate linkage between the hemispheres. A conspicuous anomaly in the Greenland icecore records may afford the clue for the cause of the two climate warming pulses that dominated the last termination. Namely, there is little or no indication in these ice-core records of the Oldest Dryas warmGeografiska Annaler - 81 A (1999) · 2 ing pulse so evident elsewhere. We suggest that this was because of a massive collapse of marine icesheet sectors from North Atlantic continental shelves during the Heinrich 1 event of Oldest Dryas age. We speculate that this massive collapse was the key connection between these two (Oldest Dryas and Oldest Dryas/B61iing) warming pulses, suggested by our Southern Hemisphere data to have been interhemispheric events. The timing and magnitude of the Oldest Dryas warming pulse make it a prime candidate for triggering the massive Heinrich 1 ice collapse, which so decreased North Atlantic salinity that downwelling was suppressed to its lowest level of the last glacial cycle (Sarnthein et al. 1994), thereby depressing North Atlantic sea-surface temperatures and maintaining regional cooling of Greenland. We speculate further that a strong rise in the salinity of the northern North Atlantic Ocean occurred at the end of the collapse, because the reservoir of glacial ice on continental shelves, as well as in feeder ice streams, was so depleted that the influx of icebergs dropped precipitously. Together with rising summer insolation in the Northern Hemisphere, such a strong salinity jump could have switched North Atlantic thermohaline circulation into its interglacial mode, with a vigorous LNDW component formed from inflow of warm and salty water into the Nordic Seas. The paleoclimate record is the prime clue to the fundamental role of this thermohaline circulation switch in the last termination, because it shows coeval warming to near-interglacial values not only in Chile but even in Europe, despite the fact that Northern Hemisphere ice sheets were still close to maximum LGM volumes. In this view, the decisive thermohaline mode switch near the Oldest Dryas/B61iing boundary was the culmination of a series of cycles in which marine-based ice built up on North Atlantic continental shelves and then collapsed during Heinrich events. Such buildup-and-collapse cycles of grounded continental shelf ice could have been a powerful regulator of North Atlantic near-surface salinity. Low salinity from a steady influx of icebergs occurred during each buildup phase, the lowest salinity accompanied each Heinrich iceberg pulse, and a salinity jump occurred after each Heinrich collapse depleted reservoirs of grounded ice marginal to the North Atlantic Ocean. The salinity jump after each Heinrich collapse caused increased conveyor overturn and ocean heat input into the North Atlantic Ocean. But it is postulated that only the Heinrich 1 collapse was massive enough to 145 G.H. DENTON ETAL. drain peripheral ice completely (and hence to cause a salinity jump big enough to initiate the interglacial mode of North Atlantic thermohaline overturn), because by this time the circum-Atlantic ice sheets had reached maximum size (and hence vulnerability to unstable collapse from continental shelves) and also because the Oldest Dryas warming trigger was the most significant, at least as recorded in the Chilean Lake District, since prior to 49,892 14Cyr BP. Because it is registered so strongly in our Southern Hemisphere data, as well as in some European paleoclimate repositories, at a time when Greenland ice-core records preclude a major North Atlantic thermohaline switch, the Oldest Dryas warming pulse is inferred to have originated in the tropics, perhaps from direct forcing of atmospheric water-vapor production by half-precession effect. In contrast, the decisive warming near the Oldest Dryas/Bolling transition was most likely caused by the major thermohaline switch recorded so prominently in North Atlantic paleoclimate repositories. This switch was the most fundamental change in deep circulation of the world ocean during the last glacial cycle. The fact that atmospheric warming was registered as far south as the Chilean Lake District and Isla Grande de Chiloe, and even at Taylor Dome in peripheral East Antarctica (Steig et al. 1998), strongly suggests a coincident change in the greenhouse gas content of the atmosphere, most likely water vapor. Thus our paleoclimate record from the southern Chilean Lake District suggests that the major thermohaline switch near the Oldest Dryas/B611ing transition affected tropical atmosphere/ocean dynamics in such a way that the production of water vapor was reset to near-interglacial values. Because the middle-latitude Southern Hemisphere and the North Atlantic regional paleoclimate records show key similarities, we stress tropical and North Atlantic triggers for major climate changes during the last glacial-interglacial transition. But there is a major problem with this approach. Namely, the Byrd ice core in central West Antarctica, along with some Southern Ocean marine cores, suggest that Southern Hemisphere warming preceded that in the Northern Hemisphere by several thousand years at the beginning of the last termination (Hays et al. 1976; Sowers and Bender 1995). Also the Vostok and Byrd icecore records imply that the B611ing and Younger Dryas oscillations were out of phase between the two hemispheres (Blunier et al. 1997). The South146 em Hemisphere lead at climate transitions has been attributed to varying ocean heat transport across the equator linked to changing strength of North Atlantic Deep Water production (Crowley 1992; Blunier et al. 1998), to local insolation forcing of sea ice on the Southern Ocean (Kim et al. 1998), or to a bipolar seesaw of thermohaline circulation (Broecker 1998). There are two unsolved stratigraphic problems in identifying interhemispheric climate linkages. The first is that a new ice-core record from the peripheral Taylor Dome of the East Antarctic Ice Sheet shows a climate signal very close to that of the Greenland ice cores, implying interhemispheric synchrony in atmospheric signals (Steig et al. 1998), probably facilitated by rapid thermohaline linkage of polar seas (Weyl 1968). The reason for such fundamental differences between the Taylor Dome and Byrd isotope records is unknown. The second problem is that the key subantarcticmarine cores that suggest a Southern Hemisphere lead of sea-surface temperaturesare at nearly the same latitudes as the terrestrialrecords in the southern Chilean Lake District and New Zealand discussed here. Again, the reason for the perceived differences between the two data sets is unknown. A possible resolution of these problems can come from detailed AMS radiocarbon chronologies of marine and terrestrial records from a proposed Antarctic CircumpolarTransect (ACT) at the latitude of subantarctic waters (where abundantforaminifers allow extensive dating). This transect would cross key land masses, including southern South America and New Zealand, to permit detailed comparison of marine and terrestrialrecords. The results from Pacific, Atlantic, and Indian Ocean sectors should show if abrupt climate changes during the last glacial/interglacial transition were synchronous between the hemispheres, if there was a lead in Southern Hemisphere climate, or if there was a mosaic of NADW outcrops at the surface of the Southern Ocean that caused mixed climate signals in Antarctic ice cores. Acknowledgements The research was supported by the Office of Climate Dynamics of the National Science Foundation, the Lamont-Scripps Consortium for Climate Research of the National Oceanic andAtmospheric Administration, the National Geographic Society, the Norwegian Research Council, and the Swiss National Science Fund. P.I. Moreno was supported Geografiska Annaler · 81 A (1999) · 2 INTERHEMISPHERIC LINKAGE OF PALEOCLIMATE DURING THE LAST GLACIATION by the EPSCoRProgramof the NationalScience Foundation.The overallprojectwas carriedout in co-operationwith the Servicio Nacional de Geologia y Mineria, Santiago, Chile. We are very gratefulto W.S.Broeckerforhis continuedsupport andfor manydiscussionson problemsof paleoclimate;he initiatedthis workin 1989 by invitingG. Denton,C. Heusser,andL. Heusseron a field trip to southernSouthAmericafundedby Exxon. We thankGerardBondfor manydiscussionsaboutthe NorthAtlanticpaleoclimaterecord.ArturoHauser Y., JorgeMufiozB., and Hugo MorenoR. helped enormouslywithgeologicaladviceandwithlogistics in the southernChileanLakeDistrict.G. Dentonis gratefulto Nick Shackletonforprovidingthe basic isotope data for TR163-31B reproducedin Fig. 5. WethankW.Beck, G. Bonani,C. Eastoe,S. Gulliksen,I. Hajdas,A. Hogg, T. Jull,R. Kalin,R. Nydal, and M. Stuiverfor providingradiocarbon dates. RichardKelly draftedthe figures, and D. Seymourtyped the manuscript.George Jacobson aidedgreatlyin scientificdiscussions,andalso edited the manuscript.We thankJohnSplettstoesser for editingthe manuscript.The reviewsof Chalmers Clappertonand David Sugden improvedthe manuscriptextensively.This paper is funded in part by a grants/cooperativeagreementfrom the National Oceanic and AtmosphericAdministration. The views expressedhereinare those of the author(s)anddo notnecessarilyreflecttheviews of NOAA or any of its sub-agencies. George H. Denton, Department of Geological Sciences and Institutefor Quaternary Studies, Bryand Global Sciences Center, University of Maine, Orono, Maine 04469, USA. Thomas V. Lowell, Department of Geology, University of Cincinnati, Cincinnati, Ohio 45221, USA. Calvin J. Heusser, 100 Clinton Road, Tuxedo, New York10987, USA. Patricio I. Moreno, Institutefor Quaternary Studies, Bryand Global Sciences Center, University of Maine, Orono, Maine 04469, USA. Bj0rn G. 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