ru t; NAUTILUS 120 2):52-65, 2006
A review of Typhisopsis fousseaume, 1880, and Typhisala
mint
Roland Bloiiart1
C a r o le M. H e r t z 2
lijsliU.it atya I d e s Si m u t e s n a h m lit js
Santa Bari iara Museum <>l Natural Motors
2550 Puesta del Sol Boat I
Santa Badium ( A 03105 I KV
d e B elgique
line Vau tier, 20
1000 B nivelles, BELGIUM
ABSTRACT
T y p h i s o p s i s loitsseannie. ISSU and T y p h i s a l a |ousscai.unc..
ISSI. are n •described and the included species reMewrd. T ip
p hisojisis ( ' o ro a a ta s BrinkTip. I S.>3 is eimsiderei I as (lu ■seniitr
svnnnvin ui /' t f u t u l n / t u s Minds, LS-13:■ M u r e x s i p h o n i / t ins
Lesson PS-44, and T y p h i s i m n i y n a Dull 1002 In addition
Ti j phi s oj t si s t 'tm>l skt >gl m u Iac n e w s p e c i e s is d e seri lied I n .....
w e s t e r n P a n a m a Its r a n g e e x t e n d s to S o n o r a M e x ic o . J im new
s p e c i e s is c o m p a r e d with T y p h i s o p s i s r o n m o t u s
Typhisala
¿ninths
V. \ d a n i s . t 8 5 5 a n d I c l a r k i K e e n and C a m p b e l l
I004.L
1
T im to u r s p e c i e s Nave diUt.-rent ''loll m o r p h o l o g y e s p e c i a l h
t h e n in n h e r . h e ig h t, and In v a d í 11 o! t h e spiral c o r d s
in
ll ie m o r -
phnlogv ol the axial l'ihs and (tie siplion a l t a n a i f l i c pl'oloc o n e l i s ol (he loui s p e c i e s are illu str a le d loi t h e Ins! l i m e
Le« i n i \ p e s are d e s i g n a t e d loi' W a r e \ s i p h o n i / a us a n d loi Ty­
p h i s a l a '¿noa Its
iN T R onut ;t i o \
V h w
m onths
titication
ago,
tile
VI
lirsl
sight
i typhisopsis
species:
1 11i w* \ e i
in
the
was described
previaisK
I'OSSj
received
Iro n i C osta
a
lol
R i c a loi
was
separated
t'ttm n a h is
and
T yphisala
ol
idcn
into
tw o
grandis.
a l t e r n i i a* ■ c a n d i d e x a m i n a t i c m a n d d i s c i i ssi< u l
occurring
• ¿ r f i m l is
that
antho]
lol
began a c o m p l e t e
cies
senior
T yphis" species
s p e c i m e n s ol
w e
C o ssm a iin . 1003. D A t t i l i o a n d Hertz £1088) in c l u d e d
th e species re la te d to T y p h i s in th e lainih I v p h i d a e [as
in Sacco ( I OU I 1 a n d C a n a r d { 1963')] a n d split. it into two
sn h lu in ih os: T\ phi liae a n d T r i p t e r o U q i l u n u e D Vttilio
a nd l i m i / 19SS. T h e i r s t ud\ was hastai on m o rp h o lo g ical d i f ten n e r s ol s h e l l a n d r a d u l a r stru c tu re . T h e v a i i d i h
ol I r i p t c m t x phi liae is n o t q u e s t i o n e d h e r e i n . S u b s e ­
q u e n t a u th o r s , h o w r u T . eb o sc lo retain Tript.erotxpili­
dae, as well as lAphinat as a s n h f a m i h ol M u r ic id a e
instead o f I \ p in d a e ( Vnkes, lOSOa lOSOb. 100 B; N o u a i t.
1901. 1004, 2Ui ï2: Aksalào a n d Sai Uis. 2003). D N A work
c i i r r e n t h u n d e r wax e l s e w h e r e l( )li\erio a n d H o n ar R in
prep.) for Tt/phiia l i t i s labiatus i.( li stolon a nd |an 1S32)
'Typhis s o u r r h y i fl a n d V. Adams. 1.S5S.) as well as
I n l i n e I ) \ V w ork ma\ lielp c la rih th e taxonomx in this
group. Rat 1 ilar t c n u i u o l o g v i..11ow s I jm lari i 1 0 0 1 ) bast <1
on D A t t i l i o a n d H e r t z ( lOSSi.
Vbbrevinlions used in ibis s t in k are BM N II): N a t u ­
ral I lisleu \ M u s e u m , L o n d o n (usait Britain ( VS: C a li­
fornia Vcadeim of Sciences San ITuneisvo ( Jalifornia,
USA: I BSN B. Institut royal des Sciences n a tu r e lle s d e
Belgique, Bruxelles. Belgium: LACM: N atural I l i s t o n
M u s e u m ol Los Vngeles C o m ile. (.’ali lo r n i a. i SA;
M N I I N M ust u m national tl Misione naturelle. Paris,
I1'rance
M / t le M u s e o d e ’/ o o l o g i a . San [osé, C a s t a
Rica S R M N I I : Santa B a rb a ra M usen m ni N atural I lis­
t o n California l ’SA S D N H M : San D ie g o N atura l I lis­
t o n Musta un. California, USA: SD PTC: Stan lor d Universitx Paleontological J\ pe C o l l o d i o n , Stanloi'd, USA;
t ' S N M N ational M u s e u m o l Natural IMsion M a s h in g Ion D (
I S V; ( j 11 ( áu oie and 1 îles I Ie riz ( ni IclI ion
San D ie g o California US V CS: Carol Skogh md C ollé e
lion. Phoenix Vrizonu I S V- RII Roland Mouart C o l ­
lection L a n d e n Belgium: RZ: Rielo / a n d a l i ( ¡ollection
\ b a c o Bai
las spec.: live-taken sp ec im en : dd spec.:
cmptx sia lis d e a d collected.
by
w as
J)
it
eastern
Ironi
Vtlilio
T t/pltisopsis
IM edic,
a sub-adult
idcntiiied
as
a tiw ari
lí
is
D
here
lom i
('o m n a lu s
V n o t h e r spe
specim en.
1 0 -Sj i a n d
it ■-( ■\ a 111i 111w 1.
ol t h e w h o l e u n nip
evaluation
T tjphisala
ol
Vf f i l i o
and
d escrib ed
as
licit/
a
now
spreit s \4 T yp h iso p sis.
k e rn
1041
3D / P i )_ \ n k e s
C erinum
I 0 SS.'
( 1 9 (i9 ?
R adw in
and
J)
Vtlilio
a n d o t h e r a u t h o r s c l a s s i f i e d all t h e
/ i / p / a s d i k e s p e c i e s in Ih e M u r i c i d a e
sol ila m i h
1
1
\\ pinnae
SYSTËMATK s
S ubiam iK Fxphinae ( Mssmaim
I ¡i
j rj '11 V.xsnctalc
\ va 11-1. «I■
19ÍJ3
D escription:
Shells w il.li a p e r l u r e e n ti r e usiialh willi
p r o j e i l i n g p e r i s t o m e and n o sulcus; n s u a l k loui oi m o r e
IL I louait and (
M I I r r t / . 2006
Vítrices p e r w liorh e it h e r wing-like 07 with axial swellings;
m u uried v e u t r a l k se aled anal tu b es originating I rum
id a p a al spiral enrd t>nk e u r r e n t anal t u b e Innetinnal
oliler tu b e s sealed; siphnuni canai ve ntr alh sealed with
1<4t side o v e rla p p in g right side.
(helius Typhisopsis (o u sse a u m e
Page 53
DAttilio; 1OS i : 8 4-35. ligs 7—11; DAN ilm and Hertz
I9S.S: 21, hg. Ino.
I ¡iphis j o t i a t a l o s
kai e! i n I07S: curd 1595.
I a p h i s I tfphis(tpsis ¿ n i i i f l i s — k e e n . 1071 5-10 Sig 1052 not
i't/j/hisdhi p r o i u l i s
V Vdutiis. 1855,11
\ o t Typhis {'Typhisopsis' o o n m a i u s . — k e e n
1071: .5 10 fie,
1951
ISSO
IN p e S p e c i e s : T tjp h h GOWUútíis B r o d e r i p 1S33; e a s t ­
e r n L a d fie. b\ original d e si g n a h o n
D escription:
si icll with lour varices; ap e rtu ru l varix o(
adult will i varical Hangt*. Iri Iled a b a p e r t u r a l h Varices
lieavv r o u n d e d rope like. S h o u l d e r spines b road-b ased ,
triangular Ila! ta p e r in g inwards lo a sh a rp po titl. Parti
lion c o n n e c t i n g uperlurul s h o u l d e r spine and Iasi teleoe u u c h whorl \ n a l tu b es si tu a te d n e a r p r e c e d i n g varix,
Niphnnal canai b r o a d , v e n tr a lh scaled Lett side ol ra nal
broadK o v e rla p p in g right s i d e .
Typhisopsis i'o n m a tu s >.lireuK‘rip, 1633)
(F ig u re s 1-15. 16. 1 1 - Mi 5 6 - 5 7 , 6 1 - 6 2 , 6 7'
Vdams
lelt ■ nut lO o d e r ip
1853 \
T y p h i s a l a ¿ m a t h s ■\
18 5 5 1
D escription:
Shell up lo 4 6 i n u iii length. Biennie,
l i e u w spin; high ac ute p r o t o c o u c h o il - w h i t e , ol 2.75
r o u n d e d whorls, lirsl p r o t o c o n c h whorl pu stuloso n ■
inainiug whorls smooth: l e i n u u a l lip lliiu raised, curved
l i g n u s II Me. Lirsl loui teleecoiioli whorls s m o o t h ,
g l o s s v . with no microst i ilplu le; inlrilacnk c o v e r i n g pi uultimale and Mitoi w h o i ! with nucrn sc ulp tu re con sisting
of regular g o u g e dike pits I F i g u r e 57). Shell with six
w c a k l v s h o u l d e r e d , n o d o s e t e l e n c o n c h w h o r l s . Las I
w h o i i broad s u l m v u j o d e r a le h injpressí t] and mostlv
o b s c u r e d Vvi.il sc u lp t u re ol lour broad, rounded, ropelike v a r ic e s , e a c h b e a r i n' g a b r o a d , liai, triang|_TIular.
s I u nigh inw a r d - b e u l . s h o u l d e r spine at mt( T.sect ion xx i11
P2. O n Iasi w horl ape rtura! varix broadlx e x p a n d e d e d g e
r e f l e c t e d dorsal k
billed alni p e rtu rallv with p o ste rio r
e d g e e x te n d i n g into a s h a r p dorsallv incurved spine at
shoulder: anal siphon tube! adjacent to p r e c e d i n g \ a n \
a n d a p p r e s s e d to partition c o n n e c t i n g the varix ol pre vi­
ous whorl. Final anal siphon long and dorsallv directed.
Spiral s c u l p t u r e ou t h r e e earliest varices ol h e a w
raised spiral co rd s {L3-LG¡ b e c o m i n g less p r o m i n e n t at
the a n t e r i o r end i n t e n a r i c a l spiral scu lp tu re hunt o r ab
.sent. Siphona! canal long, b ro a d T a b l e L, v e n t r a l h
1
T y p h i s e e n / notus lir o d e r ip
I 888: IT s S o w e r h v . I S 4 ], pi 2()U.
figs 3 A
T y ph is q u a d r a tu s H ind s. 1843; IS, Hinds, 1844: JM, pho. f.igso. 4.
\ } t t n V s i p h o n i f e r u s L e s s o n , Is I t 16.8
T y p h i s m a r t y r i a Dull. 6)1)2: 5 5 9 ; D a lh 1 0 0 8 , p i . 15. l i g . l l .
T y p h i s Ttfphi s o p s i* d H V i u t l t t s — K r e n . 1944: Mg 17 k i m .
1971. 5 4 0 Mg 1051 righi l e p e ligni e o f ï (puit I ra tus ]
A b b ott. 1974. 192 ( 2 0 1 7 ) ,
Tt/j<his<>psis r , n o n a t u s
l> Vttilio. 1 9 7 5 : 5 7 text lig.: I) Vttilio
1976: 28. text t'igv Ludwin and IT.Utiliu, 1.970: 2 1 2 pi 32.
figS. 1 0 - 1 2 : Vbbott a n d 1J a n e e 19 S 2 . 157. M \l lig..
-.•m
F ig u re L
M nrpholngv ul the spii J c o r d s ui Tiff ¡fusopsis i ■#o <uatio s
Brodenp
I855
T U E N A U T I L U S Vol. 12(1. No. 2
Page 5 4
T a b l e 1. Tfenilinology u s e d to d e s c r i b e th e spiral c o r d s (based
on Merle 1999, 201CÏ)
F
s
PI
Brima tv cord
Secondary cord
Shoulder cord
P2-PG
Prim arv cords on the convex part ol the
s I—s6
te le n e t >nch win ul
S e c o n d a r y c o r d s o n t h e c o n v e x part o t th e
leleoconcli whorl
p.xaiuple si - secundan cord between PI and P2:
s2
secondary n >rd Iíe tween P2 and P8. etc
AMP
udapertural primary cord ou tin siplional canal
MP
median priman cord on Ilie siplional canal
\B1'
abapertural priman cord on the siplional canal
ads
adapert 11raI secondary conl ou the siphona] canai
ms
median seeondarv c o r d on the siphonid canal
ai>s
abapertiiral secundan cord on the siplional canal
scaled alo n g ils length with 5 cords ( p r o b a h h \ D P ads.
MP. VHP nhsx t a p e r i n g with long, ba c k w a rd s lacing
sealed l u b e at distal e n d Sutural line strnnglv shifted to
11h ■right. \ p e rt i ire w hile, m o d e r a t e l v large, sm o o th , su bcircular, e n ti re, lo r m i n g an erect p e r i s t o m e a p p r e s s e d
posteriorly InlritaeaK ehnlkv. o l b w h i t e , d e e p k a n d irreguhirk pitted. Shell colo r w h ite to c r e a m with bro w n
sn lin sio ns on varices, tips ot s h o u l d e r spines a n d lip ol
siphona! canai: two to live b ro w n spots on e d g e ol a p e r ­
tura! lip. s o m e t i m e s none, on occasional s p e c i m e n s th e s e
spots alsi >on le a d in g side1ol varices. í ) p e r c u h i m c o r n e u s ,
light brow lí with 13 - 15 c o n c e n t ric lamellae and te r m in a l
nock ais. 11,mInia I ig u r e 10 ) with rachidi iii to o th I >eai‘ing
a long centr al c u s p a nd on e a c h side 3 o r I slighfk
sh o r te r, n a r r o w e r lateral d e n ti c le s ol v aried s t r e n g th a n d
le n g t h (>r i. rratically 1 is* M. a n d a 1iroai I lateral cusp, slightly
s h o r t e r than central cusp. Paierai teeth sickle sh a p e d ,
I >r<md.
1
sand b o t t o m w ith rocks. 5 spec M’S); G u a v m a v Sonora,
d r e d g e d . 2 4 m, Is p ee ., ( R U
I spec, d r e d g e d ( S D N H M
4 3 8 1 2 ): 55 m I dd specs i M N H N ); B a h ia Bacochil nunpo,
Sonora, d r e d g e d 12 m 1 spec. ( M N U N ); d r e d g e d 18—27
m 2 spec, i [ [ U: 150 spec. S B M N H); S o n o r a 1 spec.
( M N U N i. 2 s ;>ee. (RH3; oil _ Bahia San C aria s, Sonora,
19 -3 8 m, I spee, d BSN B 1G 26.158); d r e d g e d 1 8 -2 2 m,
9 8 spec. i( '[ Il ): d r e d g e d J 5 - 3 0 m , 14 spec. (CS ): 4 spec.,
31 m i . S D N H M 9 6 8 3 7 ) : d r e d g e d 106 m, 3 s p e c .
( S D M I M 71926); nii T eta s d e C a b r a s , d r e d g e d 6 6 - 1 0 0
m. 7 spec. (S D N 1 1 M 92766); T e n a ea tit a . [alisen, 2 spec.
( S B M N II ): Manzanillo. C o li m a , n e a r h a r b o r e n t rance,
15-22 m, I Spec, ( R H ) ; d r e d g e d 16—3 6 lïï; 3 spec. (CS):
' l o d g e d 81 iii I s p e e S D M I M); d r e d g e d IÖQ m.
3 s p e c . i S D N 11 M ) ; d i e d g e d 6 6 - [ 0 0 ni. 7 s p e c .
■'SDMI M h 16 spec. S B M N 1 I Salina C r u z , Oaxaca, 5
spec. (SBMN TI): ( o sta Rica. P lavas del C o co , G u a n a ­
c aste d r e d g e d 24 ui, 4 spec. 6 S): d r e d g e d 2 1 - 3 6 m, i
s p e c , ( C j H ) ; d r e d g e d 2 4 - 3 6 m, 2 s p e c . ( S D N H M
96771): I d d spec. { S D N H M 91566); I spec. ( S B M N H);
16 spec, i S B M N I i ); P a n a m a . 21 spec, i S B M N H ) ; Isla
S a n ta (hataiina d r e d g e d 5-8 m. 5 spec, i lil i ): Boca d e la
Monda. 7 ,J2 7 ' N . SO 5 K \ V in w h ite sand, 2 spec. (R H );
SK Bia Ran cheria. ( mlfo de < b i r i q u í . d r e d g e d 15436 m,
I spec. S D N H M 96775); Isla G d b ac o . olí ( a i l l o de
M on (.iju, \ e raguas, in si 11x sam I ami rubble. 6 - 9 in. 1
spec, i M N 1 I N ) : Ballia M ont i jo. \ 'e raguas, 19 m, 1 spec,
IHI); V o u a s d e O u e b r o d r e d g e d 1 9 -3 8 i i i , I spec,
( l d i ) : Isla V en ad o. I dd spec, ( S D M I M 64236); K c u a ­
dor. Isias Galápag os, Isla San Salvador, close to Bahia
fames. 8 6 - 8 5 m. coli D 41. Shaskv, I spec. (SBMNTI
366662).
D istribution:
San Felipe. Baja ( kililornia and ( iabo
T e poca. Sonora, Mexico, (o ( am \ ai jini K c u a d o r a n d I si as
t -akipagi i.s, 5 -42 m.
Remarks:
T y p e M aterial:
Typhis c o rn u a tu s E c u a d o r , Sakmgo.
li o lo K p e B M ( N I I ) 1966447 'Typhis <ptti<h titus: G o sta
Pica. ( aillo ile Nicoya, a n d Kc uador, Bahia de G u a y ­
aquil. d r e d g e d trom a muddy b o t t o m 7 lii IS Inis ( 13—53
m i o l i o d e Nicoya is h e r e d e s i g n a te d as Ivpe locality ),
h o l o t v p e B M NII
I S 11 B . , >7v. Mu arx s i p h e w i fi rus:
Mc 1\ i(s i . V a p u l e o . ( ¡ucrr ero, 1 [ectotxpi NINI IN du re
se le c te d .1 Typhis u u iily r ia ( a i l l o d e 1 ¡alilornia, oti San
Pedro Martii 26 m. sand, holotvpe L'SNM I3Ü62Ô
( H h r r M aterial E xam ined:
M e x i c o . Bahia M a g d a ­
lena Baja C a l i f o r n i a Stiiy I spec : S B M N 11 y San Felipe,
Baja Galii
. 1 spec. ( S B M N I I Ballia d e kis \u geli s.
[hija ( a lí Iori ña d o ■<Igc 'i I 2 7 iii I spi •c. S I )N 1 ! M 2 8 2 16
ligured in Bar.lwiu and I) \llilio. 1976]); olí tsla San
Marcos Baja t mlilornia Sur, dive rs, 12 m on Sptutthfhis
princcjjs, I spec. i ( 3S ); Isla ( ’a r m e n . Baja Kali lori ira. I dd
spec ( S D N H M J; Bia D a u z a n l e Baja G alil n rn ia Sur,
d r u d g e d io- 4 0 ui. 3 spec, t S . d r e d g e d 3 0 —45 u l . o
spec
C [II Bahia í d o u e e p c i ó m B a j a c a l i f o r n i a , I spec,
S I ) N 11M í abo I [‘p o c a S o n o ra d r e d g e d 26 SU ni.
1
1
S u b a d u ll s p e c i m e n s o( i c o m m i t us ( F i g ­
ures I I 12. 62 . as in the tv p r , bave au incompletely
Io n ia 'd a p e r t n r a l Iip w hich g i v e s t h e shell a i iarn iw e r and
m o r e s l e n d e r oiifliue. Ute 1irst lo ur tí le o c o n c h whorls
(to a length of approximately 15 mm) of /. coronatus a r e
si
th a n d shiny yvitlioi.it the microsculpture evident on
m ature shells. In juyenile specim ens the siphona! canal is
also narrower. ha\ mg a more centrally situated sutural line.
t )l d ie two svntvpes ol \ l a n \ siph<uti¡cnts . M N I IN .
o n e i.s i'tfphisdht ¿rai alis, the o t h e r is c<. mi specific with
Ttjpniwpsis coromfus. As lirsl revisers, a n d in arder t o
m aintain t h e n a m e T ¿ n a tilis as it was illustrated by
re c e n t a u th o r s , w e d e s i g n a t e that la t te r s p e c i m e n of
Murex sip h o n ife ru s as le c to ly p e I Figure s 6 -7 ). T w o nl
IIic three - sy ntvpes oj Typhisala ¿ret m l i.s \ 9 lams, 1855)
a r e s pet: m lens ul Typhisopsis cornwalli.s (see u n d e r 11oil
s p e c ie s [or f u r t h e r e x p l a n a t i o n ) I he h o l o l v p e oí /
ip t a d n ih t s PAI N I
has the siplional canai b r o k e n but
the overall sh a p e ol t h e shell is q u ite ihe s a m e as ihe
intact s p e c i m e n illustrated h\ llinils IS 14: pi. 8, jigs
5 - 4 ' No o t h e r s p e c i m e n was located in t h e t y p e m a t e ­
rial.
[
1 1
R I Io n a h a n d (
\
^
'¡ *
M I It at/.. 2066
i - ,
Page 55
,
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..
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f» bf í rl: . fraii1
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&
F i g u re s 2—15. Typhisopsis cútonfittiis (Bmderîp, ls.S.'P1 2—3. Ecuador. SaPuigo, kofotype? BM(NPÎ) 196644?. 26 n
photo P
Craie HMNH-. 4-5 . typfiis quetdmtm Hinds 1s 1.3. Costa Pira. Coîfe de Nicoya (herein designated as ivpc locality) l'iofctypé
KM NII I s 11.6.7.35. 17 5 mm (photo P Kräh BMNH). 6—7. Murex siphon'ifenis P cssim . 154 I. Mexico Acapulco. G u en ero
lecti itvpe M \ I f \ lu je 4 i ‘sj^ iiaii'4 2 7 . 9 m m q u i n t o D Hrabanl MXIIN
S—9.
iuariyita I. >; 11 161)2 h o ln hp r t S \ M
1
1 3 6 6 2 9 . ( a rlfo iii < aiii'.iniia. oil San Pi-dn i M u l u 2 7 6 m m 10. Tt /j ih ho p vi s n Jor/r/v/uv T r i a s d<-< a i n a f a u \ mas S im n ia
J 9 4 m m . < | Il p h o t o P S a ilr g h ia u
p r o l o c o m l i illustrated F ig u r e s | 3 —|5), 1 1 - 1 2 . P a n a m a Santa t ,alaim a 5 S m LII
13-15. W.'St Panam a. Hm „i i P la IP inda 7 27 \ So 51 \ \ in while sand lili 2 3 . 5 6 m m
V!'
'I '
: i
T H E NAUTILUS
I'a ge 56
Vol. 126. No. 2
Typhisopsis ca ro lsko g lu n d a e now species
( F i g u r e s 1 7 -2 5 . 4 7 —19. 59* 63)
¿ n nuits. O p e n ‘il In rn l i g h t b r o w n w i t h 12—14 c o n c e n t r i c
l a m e l l a e a n d l e r m m a l nucleus, R a d u l a not e x a m i n e d
Typhisala ¿road is .— D A ttilio, 1987:32, ligs. 1-6: IVAttilin and
H ertz. ]95S 72-73, Jigs. IOS a - e [not Typhisala ¿ m tulis
J y [it1 M a t e r i a l :
H o l o t v p e : S D N 11 M 9677.8 C o s t a
Rica Flavas del i neo. <m a n a e a s t e , oti m u d b o tt o m . 2 4 ,87 iFi Paratypus: (Iusta Rica Flavas del Coco, G u a n a ­
caste, on m u d b o t t o m , 2 4 - 3 7 m. j p a r a type ( S D N H M
93558); on m u d b o t t o m . 2 4 - 3 7 m. 1 p a r a t v p e ( C [ t l ) ;
d r e d g e d 1 2 - 2 5 tu, 8 sp. C5 x 2 spec., I S—27 m in coarse
sand, b r o k e n s h e II b o t t o m (S B M N T I 3 5 9 4 3 ] ) ; J p a r a l x p e
i S B M M I 8594361: 5 spec., 9 - 3 6 m ( S B M N H 359 435); I
p a ra tv p e T S N M 1654295a I p a r a t v p e ( L A C M 3666);
Isla N e g r ito s, C o l l o d e Nicoya, 8 paratvpes, 2 1 - 2 7 ire
c oars e sand b r o k e n shell ( S B M N H >59430); P a n a m a
A re n a s d e Q u e b r ó . Veraguas, 1 9 - 3 7 m, 1 p a r a t v p e i RH);
Boca de la H o n d a , 7 2 7 ' N 50 5 1 ' \V. in w h i t e sand. 1
p a r a t v p e ( RM N 11 26 656871): I p a r a t v p e (M Z C R I ( Tl
6 I 5 8 1: I p a r a t v p e {MNTIN Moll 6 991); 2 p a r a l v p e s
( R i l ) ; n i i Isl a G o b e r n a d o r a , intertidal, I p a r a h p e ( III I
ol I Isla \ ei iado.. Bai na Fai uii na in muddy sand at low tide.
I p a r a t v p e i I J í s N B IG 26.517/566); Bahia C h i n i [ui, 74
in. 1 p a r a l x p e ( I R S N B 1( - 25.466/567); C o n t a d o r a , L e j ­
ías Archipelago, Bahia d e P a n a m a , t paraty pus. 15-3 0 m
(S B M N T I 3 5 9 43 2 ); I p a r a t x p e ( S B M M I 359429); E c u a ­
d o r , Isla Santiago, Isias G ala p a g o s I p a r a t x p e tS B M N H
359433).
A. Adii n s 1855 i |.
D escription:
Shell up to AKA nui] in le ng th , slend er,
heavy. Spire high. IO - ISo ol total shell length. P r o t o ­
col ich uft-while, coili cal with 2.7 l'on ík led p r o toen u el i
whorls iirsl whorl pustnlo.se, r e m a i n i n g whorls s u m o ;h
e n d i n g with de lic a te thin, yveaklv ere c t a n d c u r v e d t e r ­
minal lip: (F ig u re s 4 7 —19' te l e o c o n c h with u p to 5 or 6
b r o a d , strongly s h o u l d e r e d whorls. S u t u r e im p r e s s e d ,
partly o b s c u r e d : axial s c u l p tu r e of lirst t h r e e te l e o c o n c h
w horls w ith s h a r p varices; r e m a i n i n g te l e o c o n c h w horls
consi sting
til lo u r strong, b ro a d , r o u n d e d varices with
i_y
r e m a in s ni s h a r p la m in a visible on v a r i c e s ol Iasi whorl,
caeli varix w ith a b r o a d , flat, long, x t r o n g k i n w a r d s
p o i n t e d triangula]'. Nat s p i n e a ! adapicat extremity t o u c h ­
ing prex ions whorl. \u a l t u b e a d ja ce n t to le a d in g e d g e <4
varix a n d a p p r e s s e d to p r e v i o u s p a r t i t i o n , V a r i c e s
b r o a d e r a n d swollen at .shouldei peripheryApertura!
varix with a moderate ly b ro a d, r e c u r v e d , si n u ou s flange,
a slightk broad, long, s o m e w h a t c u r v e d sp in e at p o s t e r i o r
e d g e , s p i n e a t t a c h e d to p re v io u s whorl by an axially
gro o ved partition densely s c u l p t u r e d spiralix: lip e d g e
r o n n d h cu rv ing into sipl
canai approximately 1/4
from t h e eral.
\ p e r t u rai varix with si r o n g spiral c on Is b e g i n n i n g a! ils
o u t e r lip a n d ox e r - r i d d e n by s t r o n g axial I i inbriation
o u t e r r e c u r v e d e d g e of a l >apertural side squainose. Spiral
s c u l p t u r e ol raised, heavy p r i m a n ' co rd s ou varices. Last
whorl with lo c u rd s ( P I . F2. s2. F3, s3, PI- s Í, Vo, so. Lo.
lolloxved b\ r o u n d e d co rd s ou siphonal canai, probably
A Ö P ; ads. MF. ms a n d ABP1. forming a broad ex p an s io n
at adapiea! e x tr em it y of a p ertu ra ! varix LÍ with sealed,
r o u n d e d anal lulli1 n e a r p r e c e d i n g varix. L2 with Hat.
tria ngular spine al i n t e r s e c t ion with varices. I h l e n arica!
sei lipi lire [aiul o r a b s e n t u n d e r niagnitical ion (2 0x).
Sealed r o u n d e d anal t u b e a p p r e s s e d to partition ou p r e ­
c e d i n g v arix torn ling an angle ol a p p ro x i m a te l y 56 90
w ith axis of shell. Spiral co rd s e n d i n g as short, strongly
backward c u rv e d spines at e d g e of a p e r t u r a l varix
Aperture si ihr i r a ilar. lori m u g a c o n t i n u o u s p e ri s to m e ,
creel at ouh r lip and a n t e r i o r po rt io n of colii niellai lip
a n d a p p r e s s e d posteriorly. Siphona! canai c o m p a r a i k e h
s h or t, b r o a d , x eu Mu llv s e a l e d . S ul m a i li n e strongly
shilled 1u die righi left side ot canal broadlv o v e r l a p p in g
right side. Intact s p e c i m e n s e n d i n g with canai dorsallv
reflected and c u r v e d to the right at extremity
Fresii, not overly cleaned specimens, xxith a simple,
chalkv
■j off -\v luii- iiitritacahx showing
CJ occasional axial
striae Protect niel i of J-white with whit* p u s t u l e s on first
y hor I T e l e o c o n c h color win Je lo en 'uii i w iih brow u chi
d incidí i spines and anal l u b e s 2 .5 b r o w n spots on e d g e
ul oui i apt 11m .J lip and o l l e n visiblr on lead i ng e d g e ol
previous Varices O n e b r o w n spot midway inle rxaricalh
on i a< 1 varix t ■yee pf die iperiiiml v a rix as in / y p h i s a h i
1
O t h e r M aterial E x a m i n e d :
Mexico. Bahia San C a r ­
I spec. ( R í l ) .
los. Sonora, 31 m
T >Vc I j o c a l i t v : C o s t a Rica, ( 4 a m a r a s t e . Play as del
Coco, ou m u d In. 46 »iii , 2 1—3 . iii .
D istribution:
So n ora Mexico to Isla \ c n a d o Ballia de
P an a m a , P a n a m a from t h e intertidal Lo 74 m a nd Isias
( àilapagos. E c u a d o r .
E tym ology:
T his species is n a m e d in h o n o r of Carol
S k o g l u n d ot Phoenix. Arizona, a u t h o r a n d specialist in
P a n a m i c mollusks. w h o iirst c o ll e c te d the type material
ol f hi s species a n d d o n a t e d th e h o lo tv p e a n d a p a ra tv p e
lo i h e S D N H M
S h e lias a ls o d o n a t e d a d d i t i o n a l
parafypes nj this species Irom lier private c o l l o d i o n to
d i e L 8 N M a n d lo ih e [ ACM.
Tifphisnpsis c/¡m!skí>¿hnt<htr di i Iers Irom T
in hav in g a b io a d e i sipfional canai with an
e x p a n d e d va ri cal llange e x t e n d i n g almos! hi t h e tip of
f l i e siphona] canal: flange sinuous midway ti» t h e a n t e r i o r
e n d r o u n d k i n n i n g ink» t h e siplional canai a p p r o x i ­
mately 1 T Irom t h e e n d a n d c o n s t r i c t e d a t i h i 1 base. It
has a relatively s m o o t h shell su r1 fa" c e u n d e r a chalky,
R em arks:
jjw .
'
“
■,
'“
""
■■ ■ '
p" ■- ■’ ■■ "
°
r■
' if '
■ intritacak ( . an r i p a r e d lo ila' s q u a m u & e . pitte d irlI r i t a c a R iu T r o r o ïw fu ü . T h e axial ribs a r e usually
b r o a d e r at t h e s h o u l d e r and ther e are m o r o n u m e r o u s
a n d slightly n a r r o w e r spiial c o n i s o u fhe last l e l e o c u i i r h
whorl 15 i n ('o ni r a s i to I 1 12 i u / c i t n m a h t s T h e va­
ne i . s a p p e a l I n o a t l e j m e o c e n a 6 / s a n d J h e spiral c o r d s are
l e s s p r o n o m ii.vd i u I lie ill ay s p e c i e s ,
s u 11pi *
lile h o lu lv p r ol Tifphixopsis c a n o sh o p ltn u la c xx ;js il
11 1 lí marl and (
l ‘agi ■ >j
M í lertz. 2 6 0 6
- r .i i
I igu res 16-26.
S p n ( i n s ol T y p h i s o p s i s und T y p h i s a l a
H>. Tifphrsopsis co i o n a f us lir o d m tp 15.88 \ ] . - \ i c n < n i mm M an za n illo
ui ar JLLirl Ji o’ i a tr a n c o 1 > - 2 2 m. m i l KH. 2 4 . 3 5 m m 1 7 - 2 5 , T y p h i s o p s i s c a r o l s k o ¿ h t m h n now q u-rios I 7 - 1 S. ( a >sta I'lica H a p u k i
( i k . r i ( mai m( .mti.- un n m d h o l t e n i
: i >55 N I 7 I m m p l m t o k Harveii [.
2 1 -.7 m lu »loh ¡io 5 J ) N J I M 9 6 7 7 4 17 S m m pï ml n K R undel16. I'.nohfH S D N I I M
20—22* \ \ i 'Stem I'anai'na Boca du' In I lu n d a 7 2 . ' \ N 6 ”>1‘ \ \ i i i w h i t e sand [u r a lv p n l i l i
10J mm.
W e stu m Panama Boí i á e Ia Honda 7 27' \ su ~*\' \ \ in w h t e sandL paratvpe M N îîN Moll 6991 I s u hum 2 L
Mexico, S «j • L Ballia 5,m Gárlqs, 31 jn. OU. 26 .... 25. Paral \ pi ■NHMMI 859436. type? locality !6 6
(photo P Sacie glmtij
p ro to c o l tch il in^rraf c d \ ign ros 16 IS 2 6 —29* T y p h i s a l a c l a r k i k e t a and i ¡impi mil 1664 2 6 - 2 7 . Bai im l a n a i iii Iski
n. .d.1 1
- 1
Im.1' a ■■ : ( \ S l Z 6 6 4 G 6 7 rx-31 PTC ' 17 2 3 22 S i n
p h o t o ( . Me t z
2 S - 2 9 . Paratvpt
B M NII
22 ...... p l i n i o i ' < mh BAI NI I
Lage 38
T H E N A U T I L U S . Vol. 120. No. 2
In st r a t e d in D A t t i l i o 1957; ligs. I a n d 2 [kTt|> and
D' Al t i l l o a n d I l e r t /. 1 1985: f'ig. [OS. a - e ) , as a dwarf Tip
phisala t i m m i is, b u t T yp h iso p sis ca ro lsko ¿ h m d a i differs
1Vínii 'Typhisala ¿ r a n tlis iu having a smalle t , n a r r o w e r
shell ai maturity a n d (e w e r a n d b r o a d e r spiral c o rd s on
the last t e l e o c o n c h w h o rl 15 in T yp h istyisis ert rols k o ­
g lu n d a e in co n tr a st io 2 0 - 2 3 on T yp h isa la grandis). Tip
phisojisis ca ro lsk o g lu n d a e also has b r o a d e r , sh o r te r, flat
\ arica! s h o u l d e r spines. typical ol T y p h iso p sis a n d n a r ­
ro w e r less s h a rp axial ribs with i h e flange on t h e a p e r ­
tura! varix nn»re stronglv c o n s t r i c t e d at th e base ol (he
siphona! canal lhan i i i Typhisala ¿candis
T y p h is o p s is ctirotsfioglm uhie d i l l e r s from Typhisala
<hnki iu having b ro a d r o u n d e d varices, ra t h e r than the
s h a r p varices in 7’. clarki in having anal tu b es a d ja c e n t to
( l i e l e a d i n gvn e d g
n e o f v a r i c e s r a t h e r th an niidwuv b e t w e e n
varices, and in h ayin g b r o w n spots on t h e a p e r t u r e e d g e
and on prev ious varices, which a r e lacking in Typhisala
clarki T h e new species is siinilai to ¿rtnnlis and clarki in
having ¿ l i i e x p a n d e d , si r o u g h fim b riate varieal llange and
a s m o o t h shell u n d e r ¿i si m p le challo intrilacalx,
(demis Ttpihisala | o u s s c a u m c , 1851
Tv p c sp ecies:
T yph is ¿ can dis A Vdan is, 1855. e a s t e r n
Pacific, bvrf original
designati
on.
\3
“
D escrip tio n :
Shell with four varices: apertural varix ot
adult with b r o a d apert mul varieal flange extending to
a l m o s t t he tip of siplional canal, fimbriate abaperUiralK :
o t h e r varices sharp, narrow . S h o u l d e r s p in e s narrow:
s p i n e s ol t h r e e a b a p i e a l t e l e o c o n c h w h o r l s n a rr ow ,
twisted c u r v e d inwards partition c o n n c c l i n g aporia irai
s p in e and last telenet >ncl i w Iiorl. \ u a l 11ibes si I uatio! neai
pix c e d i n g varix, p r e s s e d against [»receding partition S i ­
phona! canal broad, ve nt ral h st illed; left sitlc ot canai
o v e r h i p p i n g righi side.
Typhisala clarki Keen a n d C a m p b e l l . 196 4
Ligures 2 6 - 2 9 , 8 6 - 3 4 , 5 0 - 5 2 . 55 6 4 .
I yphïs (Typhisopsis) clarki Kirti and Campbell. 1964: 45. pi. 4.
figs. 15. 19 23.
Typhis Typhisopsis d a rk
Keen 1974:5-11). tig 1056; Ab­
bott, 1974: 192 (2016).
Typhis ('kirki. Crnmiell. 147 1 Jon 108. 5 ligs K anina
1986: card 2568,
Typhisala clarki
l> Amin», 1475 .58. text fig ; Radwin and
DAttilio, 1976: 2 11. pi 31, fig, 5. G em m cll and Ö'Attiïio
1474
88 4 8 , figs
I 8: I) \ t t i l i n ¿uitt f h r l z
1488
24
Iriei »concii consisting ol to u r high s h a r p narrow, l a m e l ­
late varices, each with a slighllv o p e n , long, twisted in
w a r d c u r v e d spine adapicallv, each varix c o m p o s e d ol
two a p p r e s s e d laminae, o n e o u t h e r e c e d i n g e d g e ex­
t e n d i n g f u r t h e r than ihe o n e ou h a d i n g edge, the two
t e r m i n g t h e slightly o p e n sp in e at shoulde r. Vual siphon
( t u b e ) fo rm in g an angle of approximately 5 5 - 9 6 with
axis ol shell, pla ced approximately midvvav b e t w e e n va
rices a n d a p p r e s s e d to p re v io u s partition, last t u b e onlv
c o m p l e t e , l o n g , o p e n , A p e r t u r a ! varix f l a n g e - l i k e ,
strongly fimbriate, dorsallv re fle cte d as shallow, o p e n
s p i n d e l s at o u t e r e d g e . Aperta irai s p i n i1 broad, triangular.
weak.lv or stronglv
dorsallvm c u r v e d , c o n n e c t e d to last te J
lencouch w ho rl bv a b ro a d , high, lamellate, partition. All
o t h e r spiral c o r d s e n d i n g lis short, wcaklv o r stronglv
b a c k w a r d cu rv ed , o p e n s p i n d e l s at e d g e of a p e r t u r a l
varix. Spiral s c u l p t u r e faint, w h e r e p r e s e n t . 12-1.4 in te rvarical lines visible on sonic s p e c i m e n s u n d e r magnification Mix) c o r r e s p o n d i n g lo o p e n spinelets at o u t e r npert urn l edge- < hi Iresh s p e c i m e n s , not o v e r-c le a n e d , i n ­
tii taca I\ s c u l p t u r e of si m ple nxriil striae p r e s e n t , FI with
sealed, ta p e rin g , anal tube. L2 o r n a m e n t e d with long,
u p w a r d s c u r v e d sp in e at in tersectio n willi varix.
Aperture small, ovate, fo r m in g a c o n t i n u o u s peristo m e , e r e c t at u n t e r lip and a n t e r i o r p o rt io n ol coi m n e l ­
la r lip Siphona! canai long, b ro a d , ven I raiIv sealed, ta ­
p e r i n g to a s l e n d e r , dorsallv c u r v e d t u b e at distal end.
Sutural line weaklv shifted to right, left side ol canal
w e itidy o v e r l a p p in g right side, ( o l o r of protocol ich offw h i t e w ith w h i t e p u s t u l e s on lir.st whorl: t e l e o c o n c h
b ro w n , c r e a m o r vcllovv isli-vv bite s o m e t i m e s with all
d i n e colors on o n e shell), ha v in g d a r k e r b ro w n anal
tu b e s, s h o u l d e r ¿uva shoulder spines a nd I ip ol siplional
cana! Apertural flange often w h ile ex en on b ro w n shells,
( >n Iresh s p e c i m e n s , not o v e r - c l e a n e d , intrilacalx s c u l p ­
t u r e oí sim p le axial s i n a e p re s e n t . ( J p e r c u l m n c o rn e u s ,
light b ro w n with 12 o r 18 c o n c e n t r i c lamellae a nd t e r ­
minal nu cleus. R a d u l a ( F i g u r e 65] with racki it liai i to oth
h e a r i n g a lo n g central cusp, and on each side 2 s h o r t e r
n a r r o w e r , l a t e r a l d e n t i c l e s of a p p r o x i m a t e l y s a m e
s t r o n g i h a n d le ngth a n d a b r o a d Literal cusp, slighth
s h o r t e r than c e n tr a l cusp. Lateral tee th sickle-s haped,
b n >ad.
O
I ype M aterial:
I lolotv pe: Manama. Isla \ criado. Bahia
P a n a m a , Panama ( A S I / 664667 ( l o r m c r lv S U P T C
9724): 1 p a r a t v p e ( ASIZ (Formerly S U F T U 9725); 1
p a r a t v p e B M ( Ñ H ) 1964433.
fig.
Mexico: Lia. San Felipe*
Baja I altiforma intertidal h in m ud a n d rocks, 28 spec.
( 11I }; intertidal, in m u d b e t w e e n rocks 7 spec, ( U S 1
m te rh d n l.
t.8 m tide. 14 spec, l S D N H M SIU.8S5 2
spec., intertidal.
I 5 in tide ( S D N H M 5 3 45 2 figured
iii
R a d w i n aral |> Attilm 1 4 7 6 a i n t e r t i d a l . 9 s p e c .
[ S D N I I M 5 2 8 6 5 8 31 spec, intertidal ( S B M N 11 : Bahía
l:i t liella. Sonora. intertidal i i i sand ov e r rocks. 7 spec.
< 8 inlertidai
I 5 in ride 2 spec. 8 J ) \ I I \ I 96885);
O ilier M aterial F \iu n in eil:
D escription:
Sin Î1 up to 27 m m in length biconic
lamellate delic ate, Spire moderé)tel) high, 3 9 - 4 2'7 of
total shell length; p r o t o c o n e h of 2.5 oil -w hit e, sm oo th ,
c u ’. . i”, conic. 4 whoris, Ii j s I whorl pn.stulnsc r e m a i n i n g
■.' h o d s M a m o i h
Ligi1n ‘s 56 52 tc lcocom h v\ il h six an
■julal
h o n g i 1 s h o u l d e r e d whorls S u t u r e i m p r e s s e d
; o r11l u b s M M o d b ' s u c c e e d i n g vv hoj‘1 Axial si u l p l u r e i »t
R
Ilouait and (
\ i
II cri/.,
A H)B
I'agí: 59
lim
i
S p e c i e s o f I i/phiwifti 3 0 - 3 4 . Ttfji/ti-.tdti c l a rk i k e e n niei < ¿ m i p b e l l I B M
8 0 —3 2 . I .m una. oft L i i G ç B a c o
Jri'ijg. ■(! m 6 J m u'.lh sand arid m l «I ík B i I. 2 2 S miri 3 3 . Ballia 1'ui mina I ski A ■'i iado und* r stm íes al k m hik- l i l i 2 2 5 mm 8 4 .
M e x ic o San Ff l i p t Baja t 'a lilu n u a ( ' S
2.5 0 m m p h o t o I' S a d e g h iu n
3 5 — 4 3 . T y p h i s a l a ¿ c a n d i s \ Adams.. 1S55 3 5 - 3 6 .
< ¿iii I■
a m i a . Iei:Lot\ p * * R M i \ 11 ) I 9 7 4 4 7 0 i I t e r e s e l e c t e d ) , >7 0 • ¡1111 p l m l n ï ' < 'raki F M N Î I
8 7 —5 9 . M e x i c o S im o n i s, ., t ;I| In
F ig u re s
.30 —4 3 .
F in ita D o b l e 2 0 - 2 2 u l . iii n i b b l e . LII > 3 7 n un 40 i ¿ a n a l i s S B M N II M e x ic o S i me r i . < m u \ m a s . Faina S a u f 11 Ios I m. I S O
lililí p h o t o P S.n I, gluau
11- 13 . ( o s l a Bien. F reviiK v e l 1'ïml. trena I si i Ballena i aiiton d e ( ka e l i d i e r e bel w e e i i 12 2 > iii I I I !
2 4 . u nun.
Pumj 60
T U I ’. N \ Í T I H
S, Vol. !2i) N o 2
44
'
yï’k'ï"'-
50
%., ¿ t s i :'■■■- --V
■-
:
"
i
'
*1 ■■ :
-
ivsgçm
|,s
■- ■;
M 't M Ï
s
'm
53
F ig u r e s
I i-5 5 ,
5 0 - 5 2 . i if¡tlttt¡!
;
v] m -
SHMMI
¡13 l> 9 Iii siltv WIH
l i i Lí I n
^011 | in l
i n u lí 'I
f S B M \ IÍ
Sl( JIH\S
in 111m i
S| ï( l( 1
1 *■i ■ ■l I ■ n ^ *
55
17—10. /
ISí.i
utitulh
i 1rtr'i I m -d
n ; l í : Sj
U I I 1 1,1.1 V e ­
ihi i 1ri ii Lii
111FE'
21 .s lu r
1¡I [i L, 1 N|irr.
JH S N U
S | Jl V .
s;
Ê Ê m
\
\i \ inn
t'tft'ohli* '¿Jtimhii* nr\\ speen-s
I S.Vj
Si il« I », cr
lim n vs M
17 5 0
5.1
V il n i m i u s I n
sla ( r \u id i
.lini II j 1iFiii
m
54
I 'i ' >! ■ «i ■ n r Ii\ 1 1 - Hi. ƒ
¡' otiHUtlttx
■■
’■.■hr ■ K' í n mii ( 11 =. | ■I n I! H.W>4 5 5 - 5 5 .
rfc ”> t i m i j u l l u i B a u r e s
zm
¡lite 11 n Lii
Perl Lis
i hi tidal
0
s
j n ■<
i
S
Isl.t
i
il fl m ‘i M i i i L ira
lí. I íoiiart and <
\I
I I r r t / . 2<M)I5
Page i'll
F i g u r e s 5(M?0. Detail oi spines and intrilaeulv ■
’>(>-.‘>7* Ti /p hi ^o j^ is c o r o n a t u s Brodnip. is.'Vi ÓS. i'i/jihisala r l a r k i Ki *t n i and
( mnphcll I ‘->D4 59, Tif¡)hisojisis i y m i s k o c h t i uta*' ru an species fiO. Ti/plthnlti w a n t lis \ Vdams, 1 5 5 5 ' \ l l l i l l S ca le Pars = 2 iri m
D istribution:
San l e l i p e . Baja i 'aliTornia a nd Bahia la
Cholla. Sonora, Mexico to P a n a m a , i n t e r t i d a l l\ to B m.
i#'
Rem arks:
I lie e le g a n t lam ellae ol the varices, th e
s m o o t h or licarh smooth shell suriner, ni id the lack ol
b r o w n spots on tile a p e r l u r a l lip and previous cálices ol
T. clùrki distinguish it Ironi t h e Other t h r e e species.
two whorls. S u t u r e d e e p h i m p re s s e d o b sc u re d l>\ s u c ­
c e e d i n g whorls. Vxial sciilplure from first te le o c o n c h
whorl to a n t e p e n u l t i m a t e whorl consisling ol lour low.
siiar]c 'varices, h e m m i n g b m a d l v roundi i! at p e r i p h e n of
p e n u l t i m a t e and Iasi whorl with a sh a rp liangi ■exte nding
out Irom tile varix b e c o m i n g íiuibriate ou its leading
edge: each varix with a modérai e h s levi 1er. cu n d n g spine
at its adapieal e \ t r e m i f \ . Ini mediatelv a d jac e n t to die
spine is a d e e p in d e n ta tio n a n t e i i o r h next to which the
anal siphon ■tube1) rests against tin jorvner partition. f i­
nal anal siphon j ui íet iomug l u b e 1 long at approximate])
9 0 to t h e shell axis, \ p e r t u r a l varix b r o a d h e x p a n d e d ,
const ricted at. a n te r io r end: oulei edge, r e i le c tr d and
denselv li m b r ia lc : e u n u e e l e d lo p re v io u s d leucuiich
whorl b\ a b roa d , high, d e n s e l y lamellose partition \p erturai spine short, s h a r p h do rs alh curved,
Spiral s c u l p tu r e ul sh a rp raised p r i m a n cords s e c ­
u n d a n c o r d s visible u n d e r magnification lo . Last
whorl, including siphona! canai with 2 2 - 2 3 cords. PI.
with sealed r o u n d e d ana! tu b e , P2 rima m en ted with
short spine at inters* c! ion with \ a n s Oth< r e in Is carnu it
be qu an tified .Spiral cords ol approxu natel v s a m e m a g ­
ín t u d e . 11 h us ' o b \ ious o n v a n e e s om ling■as short, stronglv
o
b a c k w a r d s c u r v e d sh ort s p i n e af oui er e d g e rij api u l u ra I
1
Ttjplti-stila iin m dis A. Vdams. 1555
Figures 55 —to 5 3 - 5 5 ti() fi5 fifi. 69)
Lessen 1544 105 pauilectotvpe M M I \
ser under Tifplthnjisis r o n n u t t i t s )
Tijplti* grandix A. \ciams, 1555 42. pi 27 jig. 1
A/p/dvepsA grandis
heen. 19U 54 lie. |S \bbott
1074; i 92 (201 S).
/5/p/iiWu -¿rond ¡'s — D \t tilio 197." 57 i text fig.; Hadaii i and
D Attilio. 1970 21!. pi 29. fig I. \hhott and Dance 1952:
15S; 1 text flg
Tnjihis Tiff'Jtisf’ftsi'. ' cun>it(itu,s- Kren 1.97] fig 105] Irjl
not I coronatus Brodmp, 15.30
Sol Ti/phis (Typhisopsis grandis.— Ke^n, 1971:540. fig, 1052
= '/"vp/nsr/eê coronatus
Not Typhisala grrynd/w— D'Attilie. 1957 figs 1-6; D Attilro
and Hertz 1955- jig |05 Li-e ;= Ttljtltisojisi'i rarolskiif lundae iimw species
M un:X
D escription:
Shell up to 37 9 nun i i i length, broad,
li t'u w Spire low or m o d e r a t e n - high angii late: p ro to c o u c h í >f j -whit i ■ of 2.5 r o u n d e d sm o o th whorls, lirst
whorl îlot pustuloso Figures 55 35): h-leoeoneh ol up lo
six stirni gh s h o u l d e r e d whorls. hroadK r o u n d e d on last
■
.
Varix
\p<Tture l o i i n d - o v ate, l o r m i n g a c o n t i n u o u s e r e c t
p e risto m e : o u t e r lip with 5 - 5 brown spols often a p p e a r ­
ing as well ou leading e d g e ol varices, brown spot a b u
a p p e a r i n g o n each inti r\arica! area Siplional canui long
=
T U P N A U T I L U S , V u l I2n. No. 2
P a g e 62
>i.p]umai canai 6 1 -6 2 . i ' f j f t h i s o j i s i ^ ( o n n m t u ^ lirt h lí -rip 1S33) 61. Adult specimen. 62, |uvrmle fi3* fif pit i s o p s is
now -,[>ccirs 6 í. ii/filti^ahi ffark't k- ■ ,m< I < u pï m-11 IU6-1 6 5 -6 6 , A/p/uWr/ g m m /ó \
la s IS55). 65. \«lull
66. 1 11^ »1II j 11 \ f l UII S l _ . l L - lï LlVS
O
nun.
F ig u re s 6 1-66.
1 1
S p O f 13 j J t ; - | I
1 1
1.1road
I i! d<' I' Vt-ritralk m.-,tied Sutural line stronglv
sh il l e d lo [lii rigl it. I< ft sr« It' ol canai bro ud k o v e r l a p p i n g
right s i i k ■. M i e m s c u l p t u n ol thin low c u r v e d l a n e l i a e
<o\ i-Tiiig tin i amai am I w «aikk t >\ t d a p p i n g v arietis. < \ e e p t
apertura! varix. Intritaea h
w ith occasii jitai axial striae,
off-white*
sometim es
thick,
( olor uii Ik\-w Ijih- lo ¡avera 1er light tau t;m o r ali riosi
oil tirets d a r k browni With b r a w n c o l o r e d spines, anal
1 1 1
lubes, a n d s h o u l d e r area. O t h e r b ro w n sp ots b e t w e e n
spiral e< m Is. ihi í niter l i p . >[ apt r t u n \ <.m lí aiding <J g e ot
variens not inti rvaricalk and occasioiinlh on sutural line
ol siplional eanal. < )p eivulu ui c o r n e u s , light brown with
j I 12 c o n c e n trie lamellae a n d terminal nucleus,
Hadula ( F i g u r e 69 with raehidian tooth hi nring a long
( e n tr a i cusp. and on e a c h sido 3—4 short narrow lateral
denticles ol \ a n e d s t r e n g t h and length <x casionnllv lused
I ¡ 1I (.mart am ! (
M l l e l t / M)( Ï6
Rage 63
a n d gravei b o tl o n i, I spec. ( R H ) ; 6 2 - 0 3 in. I spec
MRS MB IO 26 .8 17); 3 7 - 5 2 m, 1 spec. í I R S N B I G
27 087'- Isla S a n ta ( atalina, d r e d g e d 5-8 m ! spe c
RH): b i a Yc&adn, intertidal. L s p e c (( S E c u a d o r: Isias
G ala p a g os Isla San Salvador efose to Ballia ¡ames, 80
85 m, m i l I > Iï Shuiky. 1 spec ( S B M N H 36600 3)
D istrib u tio n ;
P u n t a D o l d e » Skin o ra M e x i c o t o
P a n a m a and the Isias Galapagos, E c u a d o r mU-rlidulb to
6 2 nr
K e n i ¿irks; Q { tl íe t h r e e sviltypes of Tt/phis g f m a /B
h o u s e d in B M ( N J I ) , !\\o are T yph isopsis cornuatus, th e
i ïtlier is Typhisala gr¿mfos In o r d e r to maintain t h e sta­
tus ol /’ f f i m d i s as il was illustrated b\ recent a u th o r s
(except K een. 1U71), o n e svntvpe s p e c i m e n is h e r e des
iguated as the lectotvpe (F ig u re s 35-361 T h e ty p e local­
ity is d o u b t lui because, to o u r k n o w le d g e , no s p e c i m e n
lias i 'M T bei :ip n pt tried from ( ali fottii lí s i u c A h e n M a m
ti m es iu old publicatio ns ( nlilornin is also used to r
h m* a- ( lalifomîa now known as Baja ( fell fo rnia w h ic h is
p a r t ol Mexico. Ii is p r n b a b h t h e case here. S u ba d u lt
forms ol T, grandis (Figures 40—18. 661 differ tr o m the
ad ult form, in h avin g a n a r r o w e r , lo w e r p artit io n, a
r o u n d e d a p e r t u r a l varix w ithout varica! flange, a n a r ­
rower. m o r e acute, siplional canai, with the ventral suIaral line bein g menv central* In adults th e left side of the
canai cxienfeveK overlaps t h e right side Figure 65 ¡.
F ig u re s 67-69* Kariulae (from D A!.[ iIn > and Hertz, I9SS).
67*
i
B rude rip 1S33
! s Lis I ;.il;i|i.igos
Isfe Jem s, 6 8« Typhisala clitrki (Keen und Cíiiiipíwil. 196-1),
Bajá < 'alitbmia , Sun Felipe; 69. Typhisala grandis (A.
\darns 1855 Ylevicu. Bahía fe Band otus I rt known magnilications
Or split. and a long, broad lateral cusp slightiv shorter thaii
central f-üsp. I .afer; 11 teeth sickle shaped, broad
T ype M aterial: RM^XH.1 197470, 3 sen type s here se­
lected as lectntvpe and p a r a lee totvpes (F ig u res 35—36),
<California“ i see Remarks ■.
O th e r M a teria l E x a m in ed :
M exico: P u n ta D o b l e ,
S a u (Carlos, S onora, 2 0 —22 n e in rubble, l s p e c . (R H ); 2
é d sp ec. 2(1-21 rn diving, i S D X H M 0U384): 1S -2I ire 4
s p e c . (GS : div ers. 2 0 - 2 1 m in r u b b l e , 2 dd s p e c ,
i S D X H M b o il Ballia San Carlos. Sonora, d r e d g e d 3 1 —7 5
t u . 12 spec. i C S ). 3 s p ec. (C JH ); d r e d g e d 61) n>, 2 .spec.
íC j íJ ) ; d r e d g e d 3 - 5 ru, 4 s p ec. ( 5 D X I I M 7 6 4 7 5 ); 64
spec. S B M N H 3 nii SK S a u Vntonio, G u ay m a s, S o nora, d r e d g e d 100 m. 2 d d spec. ( S D X i IM 8 0 7 6 4 ); M a n ­
z a n i l l o . C o lim a , 2 0 .spec. i S B M N H h in 6 m. ( S D X I Ï M
2 3 2 1 5 a*s qimdratu.% : C osta Rica Isla Ballena* Puhtaÿenas, offshore. 1 2 - 2 5 m. 4 spec, i RH); 4 spe<G ( RZ): Id a
Ballena. C a n to n d e Osa. Puntarenae, o ffsh o r e, b e t w e e n
12 and 2 5 m, 10 spec* (subariuli. dd) RH
8 sp ec.
f C | H ; 2 s p ec. i R Z 1: Panama: 10 s p e c
S RMXI 1 ) ; Isla
( i u b e r n u d o i u 1 s p e c . MXI E X ; e x t r e m e lo w tide. in
nii id tb sand n e a r rocks, I spec
sand, I spec
lili
RH
low tide. i
K
Isla Ctfoaco, dredged in 37 nu sajid
D iscussion:
Both ï'ifphtsopsiti aíid Ttjphisalä se e m to
be clo seh it feted* and il s e e m s rjuvstionable w h e t h e r
lioth ,.ire s e p a r a t e or not. However* several d if fe re n c e s
have b e e n d e l e c t e d so for: In Typhisopsis t h e varieal
spini' is Halo broad ly tria ngular a n d strongly inward b e n t
fF i g u r e s 56, 5 9 F c o m p a r e d to the longer, weakly tw is te d
s pine m Typhisala ( F i g u r e s 57, 60 and the a p ert ura l
varix of T yp h isa la is fimbriate and broad Iv e x p a n d e d
w h e r e a s the &p erin rai varix of Typhi.st.fysis is n a r r o w e r
and m o r e r o u n d e d with lieàvv raised spiral cofels Fhc
ijuestion is w h e t h e r or not sueli d i l l e r e n c e s are signifi­
cant e n o u g h lo w a r ra n t s e p a r a t e g e n e r a M a y b e D N A
woi'k w o u ld b e use Eul iu thi is part icular case.
\t:k n ( m e r i x wífxts
W e are m o s t gratelul to M G. forry) H ar asew vch and
F a n i G r é c n h a l î I S W I for fhe loan ni Typhis Yjttarttjrîù
Pall. ¡¡902; to Kathie Wav and Fliil ( rabb ( B M N 1 L ha
digital images; to Virginie H er o s and D e l p h i n e Brabant
i MX I IX) tor h e lp in s e a r c h i n g l i t e r a t u r e a n d s p e c i m e n s
from foe collection oj MX1 I X a n d loi digital images, to
Paisley C a to a n d L a u ra Halverson ( S DN H M ) for loan o f
ila' two tvpe spec-ji iu -us n\ ifffd iiso j^is ù û w h k o ^ l u n d n f
to G e o r g e Mot/. N< ïvati ». ( 'al ii o t t i ia, lor digital imagi 'S of
the h o l o t m é ol Ttdipfphis <kirki at ( -AS.; to Kelvin Barwiek (Í tty of San Diego E M T S Lab ' tor digital im a ge s 4
th e I lolotvpe and p a r a t v p e ol Typhisopsis c a m f ik o ^ lu n dm:: to Daniel G e i g e r X B M M 1 ) for S E M p h o t o g r a p h s
ot t h e protocol ich s ol ila' f our species: to Patricia Sat le ghiau 3 R - M M I lor digi Fil im a ge s ni the c o m p l e t e speer
1
Pa ge 64
T H E N A U T I L U S , Vol. 120, No. 2
T a b l e 2 . < .'haracters ni E asteri i P a c ifie S p e c i e s o ï
Typhisopsis
ami
Typhisala.
< l u c i i li i' slah 's
i 'ijfiliixt >) J.vb
rifpttisdlft
( haraetei
1 Piotoeoneh
Last teleoconelí
2 7 5 w bite, conical
r o u n d e d whorls
p u s l u l t »so on lirsl w In jli
2 . 6 6 r o u n d e d w horls,
pusluli iso on liist vvhoi I
Modecitelv Ino;ul
Bn ui I
Elat I iroa< I not twisted at
base, sharp. in cu rv ed at
a c u te c m I
Flat .
T u b e s a d jacent to
c< |gt ■i il Ia e c i ‘I.ling v. iris
I 11hox jr :|ai s nt l<1 1' m( ling
e d g e o I v ari\ and
'Dj¡iii¡said clarki
2.5 crean i-colore« I
Vi »I 11Hl<M w IIf II Is.
nor n 1 pus Iuii »sí ■
2 .7 5 w h ite, s m o o th , cortical
rou n ded vv Itor Is pust i il< »st ■
on lirsl whorl
Bn ui I
w Ilori
1. Sii< Kilder sp in e
2
Ymd tmbi
position
3. Nurnbei ol spiral
conis i siphona)
canai inclín led 1
t
\vial ribs
I M l(
t\\ ist« l<I If »I l g ,
W i îakk twisted, long..
bn »ad al base curvei I
I »road al I »ase, In ait
at a c u te e n d
dorsallv
I u l u s at 11, icen t lo
ái lí I a[»pressed lí »
ap p n vw '»I to j m -v ions
preta d i n g varix
op press i 'd to
prex ions partiti« m
pai titii m
pi l 'ci al ujg partition
l i t - 1I broad
lo
21 22. narrow, b ro ad k
s | >lic<al o u s j p l k nial
Io narrow, broai Ik
spaced
cui ial
Bn iud, lopi like■n hiih\\■<I
\ arid -s, apertura I s arix
wi l h varica! flange
First ihn 'e te leoi, i nieli
Broad
w In n Is with .sharp
varit es. rem a in in g
w h o r ls broa«I. r o u n d e d
M eak k lv\ is led broad base
long, o p e n al i listal c u d
1 if »es upprovimulch midvvuv
between varices, appressed
to pn ei ding partition
1
12 I I. o n lv visiblc on sharp
e d g e ol axial ridges.
odicn-vi.se sin«it»th
rou n ded,
\n gnlali s s l u r p
a lu p ic a l portion
wi t h small thin
er ect lamellae:
h »mual I »\
t wo a p p r e s s e d lamellae:
.apertural varix exp a n d e d .
apertural varix wi th
In r», it ! v a m ul liui igi *
aperl'iira) varix w ith
fin ». it I v.m e a l flange,
j f hiikIK c u r v i n g i ut f»
.siphona! canai
a p p n »Minaick (I 25
In i« »re i iist.al e n d
5. \ | n a h mil \ a rica I
linage
Edge
n ' l l c e t ( ‘d :
W eald V
scjuanif »sí ■ s[lirai i •( a t Is
1l] IVIm is. m o r e si n nigh
rr
.
Brout I. r e c u n es I
1111 ihi jale on its i Ulloi
po i l i on ronndlv
Mj i i umnse i m oc ifa 'I
cu r\ m g
p i ii h o i i
canai
11 ilo
spi] n a ical
B roadk e x p a n d e d
recurved strough
Broa« I. it'fleeted as shallow
limi nia I« on lis
Ii 11 ibriale on il s <mti t
oul e i portion
St rough
const net ed
1 ■
I »i u l m »i i
d[ »eu spincli i s si rough
at a n le r iiu e n d
Sini mmai canai
M 'u lerateh broad long,
\ b n lí -rai i l\ I imaii
Ih o; a
1 apering. long
1, 11 m■ring
I Moan
breadth l e n g t h
B.I15/7.76 n u n
S (1.5/7 5 ” nini
9 11/9 .5 7 i......
1.91/7.62
2, Posit n m ol m e
snl m al Inu ■
B roadly ijverl.apping Lite
lii o;n |]\ ove d a p p in g t igkl
sii Ii
Broadlv i Act lap p in g
51 rongk ovni Inpj -mg ilo
righi s id e
lui rita«
I Jilist niei ive c hallo
i »II v luii ■. i Ii <pk
Cl i a l kv. o i l - w h i t e s im p le
vviili ori ’asi» Miul axial
si fiat1
( Ir Jkv oil c n Iluck.
lIx
righi side
i ne gi i l ar k p itied
i he i iglil side
»i l l-whil e. sim ple,
v\ ilk oecusit m a i axial
i
I dialIsx. If tin i »fl w lui» ■t<»
creani siinplt wi th
occasional axial striae
si riui ■
( Ijaraci i is oj adull s hel l s onlv
inous with ] >rr j[t Kanu 'hs lo façfv \ a 11 i .in i lieni. I T i n d i n e
Í 'liii's a n d D i a n a 11tit) rf m a o 1R 6 \ B 1 loi h e l p in s e n n i i
i ig li(, ■rat nri a n d a c c e s s tri d i c rot Ire Ejoi i ni IJISNR; to
Btel v Z a n d a l i Marsh llurbonr. A b a c o . Bahamas. f o r t h e
loan aial giti ul s p e c i m e n s ¡lini U\ Garni S k o g h m d lor
drmaling 1a j o inub r i a l ol Ihi new spet ios t o the I S _ \ \ I
and I .\ ( \1 as w ell as l e n d i n g c o m p a r a t i v e material ul
I h e Ione sj »een :s loi si i n k
u t e r vtuek
A b b o tt, R. T
<
it e d
197 1
\ m c n c a n HeiislïéMs, â î l d
Ed.
\ :m N o s -
tiEind R e i n h e i d N c w V o r k 6 (i3 pp.
\b b o li
!l I and 5 1' D u n c e 1952. I Ion i p t n t l n n 11 r»| Seu
s h e lls lv.L I iu ll o t i . Inc.. N e w York, ix i UII p p.
\b su lao
B 5
and I
\
»Ios S a n to s . 2 9 6 3
\ new
genus an d
s p e c i e s ol J v p l mi n e
M o llu s c a . G a s t r o p o d a M l i n e i d a e 1
h oí 11 oil nori he. ish i i i Biu/.i.I /.ootux.i 279: I—6.
H. I lonurt anil (
Ni. I lo rt/ . 2<M>6
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