Appendix S1. List of the species included in this study, including the values assigned to
biological traits. BL: Body length (mm); TW: Toe-webbing (1, full); DS: Dorsal skin
(1, smooth); MT: Metatarsal tubercle (1, absent); DT: Digital tips (1, expanded);
Reproductive type (1, terrestrial); LG: Larval guild.
Acris crepitans
Acris gryllus
Adelophryne adiastola
Adelotus brevis
Adelphobates castaneoticus
Adelphobates galactonotus
Adenomera bokermanni
Adenomera diptyx
Adenomera heyeri
Adenomera hylaedactyla
Adenomera marmorata
Agalychnis litodryas
Agalychnis moreletii
Allobates brunneus
Allobates femoralis
Allobates marchesianus
Allobates olfersioides
Allobates sumtuosus
Allobates trilineatus
Allophryne ruthveni
Alytes obstetricans
Ameerega bassleri
Ameerega hahneli
Ameerega parvula
Ameerega petersi
Ameerega picta
Ameerega trivittata
Anaxyrus americanus
Anaxyrus boreas
Anaxyrus houstonensis
Anaxyrus kelloggi
Anomaloglossus baeobatrachus
Anomaloglossus degranvillei
Ansonia minuta
Ansonia spinulifer
Aparasphenodon brunoi
Aparasphenodon venezolanus
Aplastodiscus albofrenatus
Aplastodiscus albosignatus
Aplastodiscus arildae
Aplastodiscus leucopygius
Aplastodiscus perviridis
Arcovomer passarellii
Ascaphus montanus
BL
38
29
14
44
23
38
25
20
23
23
23
76
64
18
32
16
17
16
18
21
55
40
19
23
26
23
42
85
108
70
36
18
21
23
40
69
58
40
52
42
45
46
20
50
TW
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
DS
0
0
1
0
1
1
0
0
0
0
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
0
0
0
0
1
1
0
0
1
0
1
1
1
1
1
1
0
MT
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
DT
0
0
0
0
1
1
0
0
0
0
0
1
1
0
0
0
0
0
1
1
0
1
1
1
1
1
1
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
0
0
RT
0
0
1
0
0
0
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
1
0
LG
LE
LE
ENDO
LE
PHY
LE
LE
LE
LE
ENDO
ENDO
SUS
SUS
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
LO
LO
LE
LE
LE
LE
LE
LE
LE
SUS
LO
Ascaphus truei
Assa darlingtoni
Atelopus cruciger
Atelopus elegans
Atelopus flavescens
Atelopus spumarius
Austrochaperina adelphe
Austrochaperina fryi
Austrochaperina gracilipes
Austrochaperina novaebritanniae
Austrochaperina pluvialis
Austrochaperina robusta
Babina adenopleura
Babina okinavana
Babina pleuraden
Barycholos pulcher
Batrachylodes elegans
Batrachylodes mediodiscus
Batrachylodes minutus
Batrachylodes montanus
Batrachylodes trossulus
Batrachylodes vertebralis
Batrachylodes wolfi
Bokermannohyla astartea
Bokermannohyla carvalhoi
Bokermannohyla circumdata
Bokermannohyla claresignata
Bombina variegata
Brachycephalus didactylus
Brachycephalus ephippium
Brachytarsophrys feae
Brachytarsophrys platyparietus
Buergeria buergeri
Buergeria japonica
Buergeria robusta
Bufo andrewsi
Bufo bankorensis
Bufo bufo
Bufo japonicus
Bufotes balearicus
Bufotes viridis
Calluella brooksii
Calluella yunnanensis
Callulops doriae
Callulops robustus
Ceratobatrachus guentheri
Ceratophrys aurita
Ceratophrys calcarata
Ceratophrys cornuta
Ceratophrys ornata
40
19
28
34
28
29
19
24
20
19
26
27
45
42
59
29
32
24
20
35
20
30
33
42
67
70
42
40
11
19
115
89
44
49
49
79
110
90
163
70
70
48
72
100
100
75
149
68
110
112
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
0
0
0
1
1
0
0
0
0
0
0
0
0
0
1
1
0
0
1
1
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
1
0
0
1
0
1
1
1
1
1
1
0
0
0
1
0
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
1
0
0
0
0
1
1
1
1
1
1
0
0
0
1
1
1
1
1
1
1
1
0
0
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
0
0
LO
ENDO
LO
LO
LO
LO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LE
LE
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LE
LE
LE
LE
ENDO
ENDO
LO
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
SUS
SUS
ENDO
ENDO
ENDO
MA
MA
MA
MA
Ceratophrys stolzmanni
Chaperina fusca
Chiasmocleis albopunctata
Chiasmocleis bassleri
Chiasmocleis hudsoni
Chiasmocleis panamensis
Chiasmocleis shudikarensis
Chiasmocleis ventrimaculata
Choerophryne proboscidea
Choerophryne rostellifer
Cochranella granulosa
Cochranella revocata
Colostethus panamansis
Colostethus pratti
Colostethus ruthveni
Cophixalus infacetus
Cophixalus neglectus
Cophixalus ornatus
Craugastor bransfordii
Craugastor chac
Craugastor crassidigitus
Craugastor fitzingeri
Craugastor gollmeri
Craugastor hobartsmithi
Craugastor laevissimus
Craugastor laticeps
Craugastor loki
Craugastor noblei
Craugastor occidentalis
Craugastor opimus
Craugastor psephosypharus
Craugastor rhodopis
Craugastor rugulosus
Craugastor sabrinus
Craugastor sandersoni
Craugastor tabasarae
Craugastor talamancae
Crinia deserticola
Crinia parinsignifera
Crinia remota
Crinia signifera
Crinia tinnula
Crossodactylodes pintoi
Crossodactylus aeneus
Crossodactylus gaudichaudii
Crossodactylus schmidti
Cruziohyla calcarifer
Cruziohyla craspedopus
Ctenophryne geayi
Cycloramphus boraceiensis
68
20
27
20
18
25
25
23
23
23
29
24
27
24
27
16
33
27
23
26
31
35
37
16
32
35
32
48
28
69
36
29
50
44
50
34
30
18
22
16
25
18
17
23
32
28
81
57
34
37
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
1
1
0
1
1
0
1
1
1
0
0
0
1
1
0
0
0
1
0
0
1
0
0
0
1
1
1
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
1
1
1
1
0
0
0
0
0
1
0
0
1
1
0
0
1
1
0
1
0
0
1
0
0
1
1
0
1
0
0
0
0
0
1
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
1
1
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
1
MA
LE
SUS
SUS
SUS
SUS
SUS
SUS
ENDO
ENDO
LO
LO
LE
LE
LE
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LE
LE
LE
LE
PHY
LE
LE
LE
PHY
PHY
SUS
LO
Cycloramphus eleutherodactylus
Cycloramphus fuliginosus
Dendrobates auratus
Dendrobates leucomelas
Dendrobates tinctorius
Dendrophryniscus brevipollicatus
Dendrophryniscus minutus
Dendropsophus acreanus
Dendropsophus anceps
Dendropsophus berthalutzae
Dendropsophus bifurcus
Dendropsophus bipunctatus
Dendropsophus bokermanni
Dendropsophus branneri
Dendropsophus brevifrons
Dendropsophus decipiens
Dendropsophus ebraccatus
Dendropsophus elegans
Dendropsophus giesleri
Dendropsophus koechlini
Dendropsophus leali
Dendropsophus leucophyllatus
Dendropsophus luteoocellatus
Dendropsophus marmoratus
Dendropsophus melanargyreus
Dendropsophus meridianus
Dendropsophus microcephalus
Dendropsophus microps
Dendropsophus minusculus
Dendropsophus minutus
Dendropsophus miyatai
Dendropsophus nanus
Dendropsophus parviceps
Dendropsophus phlebodes
Dendropsophus pseudomeridianus
Dendropsophus rhodopeplus
Dendropsophus riveroi
Dendropsophus robertmertensi
Dendropsophus rossalleni
Dendropsophus rubicundulus
Dendropsophus sanborni
Dendropsophus sarayacuensis
Dendropsophus seniculus
Dendropsophus soaresi
Dendropsophus tintinnabulum
Dendropsophus triangulum
Dendropsophus tritaeniatus
Dendropsophus walfordi
Dermatonotus muelleri
Diasporus gularis
38
46
40
38
27
15
17
35
40
21
28
25
24
19
21
15
25
30
25
24
23
36
31
44
20
19
24
18
21
23
18
24
21
24
20
23
20
26
20
25
20
37
38
32
20
28
21
26
50
22
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
0
1
1
1
1
1
1
1
1
0
1
0
1
1
1
1
1
1
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
LO
LO
PHY
LE
PHY
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
MA
LE
LE
LE
LE
MA
LE
LE
LE
LE
LE
MA
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
SUS
ENDO
Diasporus quidditus
Discodeles bufoniformis
Discodeles guppyi
Discodeles ophistodon
Discodeles vogti
Duellmanohyla schmidtorum
Duttaphrynus melanostictus
Ecnomiohyla tuberculosa
Edalorhina nasuta
Edalorhina perezi
Elachistocleis bicolor
Elachistocleis helianneae
Elachistocleis surinamensis
Eleutherodactylus abbotti
Eleutherodactylus antillensis
Eleutherodactylus atkinsi
Eleutherodactylus audanti
Eleutherodactylus auriculatoides
Eleutherodactylus auriculatus
Eleutherodactylus barlagnei
Eleutherodactylus brittoni
Eleutherodactylus cochranae
Eleutherodactylus coqui
Eleutherodactylus cundalli
Eleutherodactylus cystignathoides
Eleutherodactylus eileenae
Eleutherodactylus eneidae
Eleutherodactylus flavescens
Eleutherodactylus gossei
Eleutherodactylus grabhami
Eleutherodactylus gryllus
Eleutherodactylus guttilatus
Eleutherodactylus haitianus
Eleutherodactylus inoptatus
Eleutherodactylus jamaicensis
Eleutherodactylus jasperi
Eleutherodactylus johnstonei
Eleutherodactylus juanariveroi
Eleutherodactylus junori
Eleutherodactylus karlschmidti
Eleutherodactylus lentus
Eleutherodactylus locustus
Eleutherodactylus marnockii
Eleutherodactylus martinicensis
Eleutherodactylus minutus
Eleutherodactylus montanus
Eleutherodactylus nitidus
Eleutherodactylus pantoni
Eleutherodactylus pinchoni
Eleutherodactylus pipilans
15
145
220
125
158
33
90
90
38
36
25
30
32
19
17
25
25
33
23
33
19
19
18
18
24
23
24
41
18
29
17
29
21
18
18
22
18
15
27
80
40
24
22
32
19
44
25
44
16
31
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
1
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
1
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
ENDO
ENDO
ENDO
ENDO
ENDO
LO
LE
LO
LE
LE
SUS
SUS
SUS
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
Eleutherodactylus pituinus
Eleutherodactylus planirostris
Eleutherodactylus portoricensis
Eleutherodactylus richmondi
Eleutherodactylus riparius
Eleutherodactylus ruthae
Eleutherodactylus schmidti
Eleutherodactylus varians
Eleutherodactylus varleyi
Eleutherodactylus weinlandi
Eleutherodactylus wigthmanae
Engystomops coloradorum
Engystomops freibergi
Engystomops montubio
Engystomops petersi
Engystomops pustulatus
Epidalea calamita
Epipedobates boulengeri
Epipedobates machalilla
Euparkerella brasiliensis
Euparkerella cochranae
Eupemphix nattereri
Exerodonta smaragdina
Exerodonta sumichrasti
Fejervarya cancrivora
Flectonotus fissilis
Flectonotus fitzgeraldi
Flectonotus goeldii
Flectonotus ohausi
Flectonotus pygmaeus
Gastrophryne carolinensis
Gastrophryne elegans
Gastrophryne olivacea
Gastrophryne usta
Gastrophrynoides borneensis
Gastrotheca albolineata
Gastrotheca ernestoi
Gastrotheca microdiscus
Gastrotheca ovifera
Gastrotheca testudinea
Glandirana rugosa
Haddadus binotatus
Hamptophryne boliviana
Hemiphractus helioi
Hemiphractus proboscideus
Hemiphractus scutatus
Holoaden pholeter
Hoplobatrachus rugulosus
Huia masonii
Hyalinobatrachium colymbiphyllum
29
18
23
17
18
58
43
14
14
40
19
24
26
19
31
30
70
17
16
17
18
40
28
33
43
34
24
32
31
26
29
29
29
29
28
60
70
49
47
61
46
35
40
53
50
62
42
103
32
27
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
1
0
0
1
1
1
1
1
1
1
0
0
1
1
0
1
0
0
0
0
0
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
1
1
1
1
1
0
0
0
0
0
1
1
1
1
1
0
0
0
1
1
1
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
?
1
1
1
1
1
0
1
0
1
1
1
1
0
0
0
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
ENDO
LE
LE
LE
LE
PHY
PHY
PHY
PHY
PHY
SUS
SUS
SUS
SUS
?
ENDO
ENDO
ENDO
ENDO
ENDO
LE
ENDO
SUS
ENDO
ENDO
ENDO
ENDO
MA
LO
LO
Hyalinobatrachium fleischmanni
Hyalinobatrachium guairarepanense
Hyalinobatrachium taylori
Hyalinobatrachium valerioi
Hydrolaetare schmidti
Hyla andersonii
Hyla annectans
Hyla arborea
Hyla arenicolor
Hyla avivoca
Hyla chinensis
Hyla chrysoscelis
Hyla cinerea
Hyla femoralis
Hyla inframaculata
Hyla intermedia
Hyla japonica
Hyla meridionalis
Hyla versicolor
Hylarana arfaki
Hylarana baramica
Hylarana chalconota
Hylarana daemeli
Hylarana garritor
Hylarana glandulosa
Hylarana grandocula
Hylarana guentheri
Hylarana latouchii
Hylarana luzonensis
Hylarana nicobariensis
Hylarana papua
Hylarana picturata
Hylarana raniceps
Hylarana similis
Hylodes asper
Hylodes charadranaetes
Hylodes lateristrigatus
Hylodes nasus
Hylodes pipilans
Hylomantis lemur
Hylophorbus rufescens
Hyloscirtus colymba
Hyloxalus awa
Hyloxalus infraguttatus
Hyloxalus toachi
Hypodactylus nigrovittatus
Hypsiboas albomarginatus
Hypsiboas alemani
Hypsiboas caingua
Hypsiboas calcaratus
28
23
20
24
104
38
35
40
39
39
33
64
64
70
44
40
39
40
60
160
40
39
48
72
60
47
68
38
57
50
110
40
30
43
41
38
36
31
25
41
55
37
22
18
23
19
43
31
33
48
0
0
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
0
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
1
1
0
1
0
1
0
1
1
1
1
1
0
1
1
1
1
0
1
1
0
1
1
1
0
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
1
0
0
0
0
LO
LO
LO
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
LO
LE
LE
LE
ENDO
LE
LE
LE
LE
Hypsiboas cinerascens
Hypsiboas dentei
Hypsiboas faber
Hypsiboas fasciatus
Hypsiboas hobbsi
Hypsiboas lanciformis
Hypsiboas lundii
Hypsiboas microderma
Hypsiboas multifasciatus
Hypsiboas ornatissimus
Hypsiboas pardalis
Hypsiboas pellucens
Hypsiboas polytaenius
Hypsiboas prasinus
Hypsiboas pugnax
Hypsiboas pulchellus
Hypsiboas punctatus
Hypsiboas rosenbergi
Hypsiboas rufitelus
Hypsiboas semilineatus
Hypsiboas wavrini
Incilius alvarius
Incilius campbelli
Incilius canaliferus
Incilius coccifer
Incilius luetkenii
Incilius macrocristatus
Incilius marmoreus
Incilius nebulifer
Incilius occidentalis
Incilius tutelarius
Ingerophrynus biporcatus
Ingerophrynus divergens
Ingerophrynus quadriporcatus
Ischnocnema erythromera
Ischnocnema guentheri
Ischnocnema henselii
Ischnocnema juipoca
Ischnocnema lactea
Ischnocnema nasuta
Ischnocnema parva
Ischnocnema venancioi
Itapotihyla langsdorffii
Kalophrynus pleurostigma
Kaloula baleata
Kaloula picta
Kaloula verrucosa
Kurixalus eiffingeri
Kurixalus idiootocus
Lechriodus melanopyga
44
54
104
40
43
71
76
34
47
31
69
59
42
38
50
50
82
49
49
53
113
63
63
43
64
96
69
75
84
76
76
70
43
48
24
40
35
26
18
54
13
25
83
49
28
71
82
34
43
80
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
1
0
1
1
0
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
SUS
SUS
SUS
SUS
LE
LE
MA
Leptobrachella mjobergi
Leptobrachium hasseltii
Leptodactylus albilabris
Leptodactylus chaquensis
Leptodactylus diedrus
Leptodactylus discodactylus
Leptodactylus flavopictus
Leptodactylus fragilis
Leptodactylus furnarius
Leptodactylus fuscus
Leptodactylus gracilis
Leptodactylus knudseni
Leptodactylus labyrinthicus
Leptodactylus latinasus
Leptodactylus leptodactyloides
Leptodactylus lineatus
Leptodactylus lithonaetes
Leptodactylus longirostris
Leptodactylus melanonotus
Leptodactylus myersi
Leptodactylus mystaceus
Leptodactylus mystacinus
Leptodactylus natalensis
Leptodactylus paraensis
Leptodactylus pentadactylus
Leptodactylus peritoaktites
Leptodactylus plaumanni
Leptodactylus pustulatus
Leptodactylus rhodomerus
Leptodactylus rhodomystax
Leptodactylus rhodonotus
Leptodactylus riveroi
Leptodactylus savagei
Leptodactylus spixi
Leptodactylus stenodema
Leptodactylus syphax
Leptodactylus vastus
Leptolalax hamidi
Leptophryne borbonica
Leptophryne cruentata
Limnodynastes convexiusculus
Limnodynastes dumerilii
Limnodynastes lignarius
Limnodynastes salmini
Limnodynastes tasmaniensis
Limnodynastes terraereginae
Limnomedusa macroglossa
Limnonectes fujianensis
Limnonectes kuhlii
Limnonectes laticeps
20
49
85
71
34
35
120
36
35
47
47
135
35
35
48
47
71
52
45
109
65
65
34
139
180
146
36
50
134
134
72
63
177
41
90
83
125
30
30
30
62
70
62
65
42
76
65
54
70
37
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
0
0
1
0
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
Limnonectes leporinus
Limnonectes leytensis
Limnonectes macrocephalus
Limnonectes magnus
Limnonectes microdiscus
Limnonectes palavanensis
Liophryne schlaginhaufeni
Lithobates areolatus
Lithobates berlandieri
Lithobates blairi
Lithobates catesbeianus
Lithobates chiricahuensis
Lithobates clamitans
Lithobates forreri
Lithobates grylio
Lithobates juliani
Lithobates maculatus
Lithobates magnaocularis
Lithobates megapoda
Lithobates neovolcanicus
Lithobates palmipes
Lithobates palustris
Lithobates pipiens
Lithobates psilonota
Lithobates septentrionalis
Lithobates sevosus
Lithobates sylvaticus
Lithobates tarahumarae
Lithobates taylori
Lithobates virgatipes
Lithobates warszewitschii
Litoria alboguttata
Litoria aurea
Litoria australis
Litoria bicolor
Litoria brevipalmata
Litoria brevipes
Litoria caerulea
Litoria chloris
Litoria congenita
Litoria coplandi
Litoria cryptotis
Litoria dahlii
Litoria dayi
Litoria dentata
Litoria dorsalis
Litoria electrica
Litoria eucnemis
Litoria fallax
Litoria freycineti
100
30
120
17
47
30
35
90
62
62
152
95
89
152
131
70
75
117
117
71
120
57
110
65
76
84
83
67
88
52
52
67
43
79
27
43
45
77
62
35
36
44
63
42
40
24
38
48
26
39
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
1
0
1
0
0
0
0
0
1
0
1
1
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
1
1
0
1
1
1
1
0
0
1
1
1
1
1
1
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
LE
LE
LE
LE
LE
LE
ENDO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LO
LE
LE
LE
LE
LE
LE
Litoria genimaculata
Litoria gracilenta
Litoria impura
Litoria inermis
Litoria infrafrenata
Litoria jungguy
Litoria latopalmata
Litoria longipes
Litoria lutea
Litoria maculosa
Litoria manya
Litoria meiriana
Litoria microbelos
Litoria nannotis
Litoria nasuta
Litoria nigrofrenata
Litoria nigropunctata
Litoria novaehollandiae
Litoria nyakalensis
Litoria olongburensis
Litoria pallida
Litoria pearsoniana
Litoria peronii
Litoria personata
Litoria phyllochroa
Litoria platycephala
Litoria pygmaea
Litoria revelata
Litoria rheocola
Litoria rothii
Litoria rubella
Litoria thesaurensis
Litoria tornieri
Litoria tyleri
Litoria verrauxii
Litoria vocivincens
Litoria watjulumensis
Litoria wilcoxii
Litoria xanthomera
Macrogenioglottus alipioi
Mannophryne herminae
Mannophryne trinitatis
Mantephryne lateralis
Megaelosia goeldii
Megophrys montana
Megophrys nasuta
Megophrys stejnegeri
Melanophryniscus montevidensis
Meristogenys jerboa
Metaphrynella sundana
41
42
50
33
135
48
39
46
67
55
30
20
16
48
45
42
27
81
33
25
34
29
53
29
32
64
30
28
32
48
37
27
36
48
36
28
38
43
42
105
22
21
55
97
92
74
49
23
82
23
0
1
0
0
1
1
0
0
0
0
0
1
0
1
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
1
1
1
1
1
1
1
0
1
0
1
1
1
1
0
1
1
1
0
0
1
0
1
1
1
0
1
1
1
1
0
1
1
1
1
0
1
1
0
1
1
1
1
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
0
0
1
0
0
0
0
1
0
0
1
0
1
1
0
1
1
1
1
0
1
1
1
1
1
1
0
1
0
1
0
0
1
0
1
0
0
0
1
1
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LO
LE
LE
LE
LE
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
LO
LE
LE
LE
LE
LO
PHY
Microhyla butleri
Microhyla fissipes
Microhyla okinavensis
Microhyla palmipes
Mixophyes coggeri
Mixophyes fasciolatus
Mixophyes iteratus
Mixophyes schevilli
Myersiella microps
Nanorana yunnanensis
Nelsonophryne aterrima
Notaden bennettii
Notaden melanoscaphus
Nyctixalus margaritifer
Nyctixalus spinosus
Nymphargus mariae
Occidozyga baluensis
Occidozyga laevis
Occidozyga sumatrana
Odontophrynus americanus
Odontophrynus cultripes
Odorrana andersonii
Odorrana grahami
Odorrana hosii
Odorrana supranarina
Odorrana swinhoana
Odorrana utsunomiyaorum
Oophaga sylvatica
Oreobates cruralis
Oreobates quixensis
Oreophryne biroi
Oreophryne brachypus
Oreophryne hypsiops
Osteocephalus buckleyi
Osteocephalus cabrerai
Osteocephalus elkejungingerae
Osteocephalus leprieurii
Osteocephalus oophagus
Osteocephalus planiceps
Osteocephalus taurinus
Osteocephalus yasuni
Osteopilus crucialis
Osteopilus marianae
Osteopilus ocellatus
Osteopilus pulchrilineatus
Osteopilus vastus
Osteopilus wilderi
Pachymedusa dacnicolor
Palmatorappia solomonis
Pelobates fuscus
22
19
26
18
93
100
63
78
40
88
54
63
48
35
37
25
25
42
30
46
70
74
69
68
76
60
48
38
30
48
21
23
26
64
46
22
48
47
59
73
56
100
40
50
32
109
26
27
27
65
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
1
1
1
0
0
1
1
1
1
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
0
1
0
0
1
1
1
1
1
1
0
0
1
0
0
1
1
1
1
1
0
1
1
1
0
0
0
0
0
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
1
1
1
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
0
0
0
0
0
0
0
1
0
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
SUS
SUS
SUS
SUS
LE
LE
LE
LE
ENDO
LE
LE
LE
LE
PHY
PHY
LO
MA
MA
MA
LE
LE
LE
LE
LE
LE
LE
LE
PHY
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LE
LE
LE
PHY
PHY
LE
LE
PHY
PHY
PHY
LE
LE
PHY
ENDO
ENDO
LE
Pelodytes punctatus
Pelophryne lighti
Pelophylax bergeri
Pelophylax esculentus
Pelophylax fukienensis
Pelophylax lessonae
Pelophylax nigromaculatus
Pelophylax perezi
Pelophylax ridibundus
Peltophryne fustiger
Peltophryne guentheri
Peltophryne gundlachi
Peltophryne lemur
Phasmahyla guttata
Pherohapsis menziensi
Philautus aurifasciatus
Philautus davidlabangi
Philautus pallidipes
Philautus petersi
Philautus surdus
Phrynomedusa marginata
Phyllomedusa bicolor
Phyllomedusa burmeisteri
Phyllomedusa camba
Phyllomedusa hypochondrialis
Phyllomedusa iheringii
Phyllomedusa palliata
Phyllomedusa rohdei
Phyllomedusa tarsius
Phyllomedusa tetraploidea
Phyllomedusa tomopterna
Phyllomedusa trinitatis
Phyllomedusa vaillantii
Physalaemus albonotatus
Physalaemus biligonigerus
Physalaemus centralis
Physalaemus cuvieri
Physalaemus ephippifer
Physalaemus fernandezae
Physalaemus fischeri
Physalaemus gracilis
Physalaemus henselii
Physalaemus maculiventris
Physalaemus marmoratus
Physalaemus olfersii
Physalaemus riograndensis
Physalaemus signifer
Phytotriades auratus
Pipa parva
Pipa pipa
40
15
80
80
66
80
81
80
120
65
65
38
36
35
25
26
20
25
26
31
31
115
72
70
46
75
44
45
90
69
48
80
59
32
32
40
27
40
20
33
35
19
23
42
23
20
25
29
37
110
0
0
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
0
1
1
1
1
1
1
1
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
LE
ENDO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
SUS
SUS
LE
SUS
SUS
LE
SUS
LE
SUS
SUS
LE
SUS
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
PHY
SUS
SUS
Pipa snethlageae
Platymantis acrochordus
Platymantis aculeodactylus
Platymantis admiraltiensis
Platymantis akarithymus
Platymantis boulengeri
Platymantis corrugatus
Platymantis gilliardi
Platymantis guentheri
Platymantis latro
Platymantis mimicus
Platymantis myersi
Platymantis neckeri
Platymantis nexipus
Platymantis papuensis
Platymantis schmidti
Platymantis solomonis
Platymantis weberi
Platyplectrum ornatum
Plectrohyla bistincta
Plectrohyla hartwegi
Plectrohyla lacertosa
Pleurodema bibroni
Pleurodema brachyops
Polypedates colletti
Polypedates leucomystax
Polypedates macrotis
Polypedates megacephalus
Polypedates otilophus
Pristimantis achatinus
Pristimantis acuminatus
Pristimantis altamazonicus
Pristimantis bicumulus
Pristimantis carvalhoi
Pristimantis caryophyllaceus
Pristimantis chalceus
Pristimantis chiastonotus
Pristimantis conspicillatus
Pristimantis crenunguis
Pristimantis croceoinguinis
Pristimantis cruentus
Pristimantis diadematus
Pristimantis euphronides
Pristimantis eurydactylus
Pristimantis fenestratus
Pristimantis floridus
Pristimantis gaigei
Pristimantis imitatrix
Pristimantis labiosus
Pristimantis lacrimosus
92
27
27
36
24
66
38
34
30
33
40
58
62
43
46
46
54
50
37
68
64
48
40
44
50
57
60
72
80
35
23
24
35
17
24
27
40
25
49
18
28
27
27
24
35
18
30
20
51
26
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
1
1
1
1
1
1
1
1
1
0
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
0
1
0
1
1
1
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
0
0
0
0
1
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
SUS
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LO
LO
LO
LE
LE
LE
LE
LE
LE
LE
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
Pristimantis lanthanites
Pristimantis latidiscus
Pristimantis malkini
Pristimantis marmoratus
Pristimantis martiae
Pristimantis mendax
Pristimantis muricatus
Pristimantis ockendeni
Pristimantis ornatissimus
Pristimantis parvillus
Pristimantis peruvianus
Pristimantis rhabdolaemus
Pristimantis ridens
Pristimantis subsigillatus
Pristimantis taeniatus
Pristimantis tenebrionis
Pristimantis terraebolivaris
Pristimantis urichi
Pristimantis variabilis
Pristimantis ventrimarmoratus
Pristimantis vilarsi
Pristimantis walkeri
Pristimantis zeuctotylus
Proceratophrys appendiculata
Proceratophrys avelinoi
Proceratophrys bigibbosa
Proceratophrys boiei
Proceratophrys fryi
Proceratophrys melanopogon
Proceratophrys schirchi
Prostherapis dunni
Pseudacris brachyphona
Pseudacris brimleyi
Pseudacris cadaverina
Pseudacris clarkii
Pseudacris crucifer
Pseudacris feriarum
Pseudacris fouquettei
Pseudacris kalmi
Pseudacris maculata
Pseudacris nigrita
Pseudacris ocularis
Pseudacris ornata
Pseudacris regilla
Pseudacris streckeri
Pseudacris triseriata
Pseudis boliviana
Pseudis caraya
Pseudis laevis
Pseudis limellum
26
26
37
18
17
20
41
21
27
20
29
29
19
17
25
27
31
29
19
26
32
19
30
60
29
44
62
62
50
25
24
35
30
36
29
28
35
30
38
37
33
16
35
48
48
37
18
63
21
25
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
0
0
1
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
0
1
0
0
0
0
0
0
0
0
0
1
1
1
0
1
1
1
1
0
1
1
1
1
0
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
1
0
0
0
1
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
Pseudis minuta
Pseudis paradoxa
Pseudis platensis
Pseudobufo subasper
Pseudopaludicola boliviana
Pseudopaludicola llanera
Pseudopaludicola mystacalis
Pseudopaludicola pusilla
Pseudopaludicola riopiedadensis
Pseudopaludicola saltica
Pseudophryne bibronii
Pseudophryne coriacea
Pseudophryne major
Pseudophryne raveni
Quasipaa spinosa
Rana arvalis
Rana aurora
Rana boylii
Rana chaochiaoensis
Rana dalmatina
Rana draytonii
Rana italica
Rana japonica
Rana latastei
Rana luteiventris
Rana multidenticulata
Rana muscosa
Rana ornativentris
Rana pretiosa
Rana tagoi
Rana temporaria
Ranitomeya imitator
Ranitomeya minuta
Ranitomeya vanzolinii
Ranitomeya ventrimaculata
Relectivomer pearsei
Rhacophorus appendiculatus
Rhacophorus arboreus
Rhacophorus gadingensis
Rhacophorus margaritifer
Rhacophorus moltrechti
Rhacophorus nigropalmatus
Rhacophorus owstoni
Rhacophorus pardalis
Rhacophorus reinwardtii
Rhacophorus schlegelii
Rhacophorus taipeianus
Rhaebo caeruleostictus
Rhaebo glaberrimus
Rhaebo guttatus
51
65
55
94
17
20
15
20
21
14
30
24
29
30
80
60
80
70
56
60
120
60
63
60
60
35
50
60
50
58
70
22
15
17
17
42
37
60
24
44
42
100
45
55
53
43
45
81
80
97
1
1
1
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
0
1
1
1
0
1
0
1
0
1
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
0
0
0
1
1
1
1
1
1
1
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
PHY
PHY
PHY
PHY
SUS
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
Rhaebo haematiticus
Rhinella acutirostris
Rhinella beebei
Rhinella castaneotica
Rhinella ceratophrys
Rhinella dapsilis
Rhinella dorbignyi
Rhinella fernandezae
Rhinella granulosa
Rhinella hoogmoedi
Rhinella icterica
Rhinella jimi
Rhinella margaritifera
Rhinella marina
Rhinella ocellata
Rhinella ornata
Rhinella poeppigii
Rhinella proboscidea
Rhinella pygmaea
Rhinella roqueana
Rhinella schneideri
Rhinella sternosignata
Rhinophrynus dorsalis
Sachatamia albomaculata
Scaphiopus couchii
Scaphiopus holbrookii
Scaphiopus hurterii
Scarthyla vigilans
Scinax albicans
Scinax alter
Scinax argyreornatus
Scinax aromothyella
Scinax berthae
Scinax boesemani
Scinax boulengeri
Scinax cardosoi
Scinax chiquitanus
Scinax crospedospilus
Scinax cruentommus
Scinax cuspidatus
Scinax eurydice
Scinax flavoguttatus
Scinax funereus
Scinax fuscomarginatus
Scinax fuscovarius
Scinax garbei
Scinax granulatus
Scinax hayii
Scinax humilis
Scinax ictericus
62
47
51
37
65
77
64
57
49
52
115
168
47
150
44
75
63
54
38
72
210
41
75
29
70
72
62
16
44
32
23
32
25
33
49
21
36
38
27
23
40
26
37
20
43
42
40
36
34
32
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
0
0
1
1
1
1
1
0
1
0
1
1
1
0
1
1
0
0
1
1
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
1
1
1
0
1
1
1
1
0
1
0
1
0
0
0
1
0
0
1
1
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
LE
LE
LE
PHY
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
SUS
LO
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
Scinax littoreus
Scinax nasicus
Scinax nebulosus
Scinax parkeri
Scinax pedromedinae
Scinax perereca
Scinax perpusillus
Scinax quinquefasciatus
Scinax rostratus
Scinax ruber
Scinax similis
Scinax squalirostris
Scinax staufferi
Scinax sugillatus
Scinax trapicheiroi
Scinax trilineatus
Scinax v-signatus
Scinax wandae
Scinax x-signatus
Silverstoneia flotator
Silverstoneia nubicola
Smilisca baudinii
Smilisca cyanosticta
Smilisca fodiens
Smilisca phaeota
Smilisca sila
Spea bombifrons
Spea hammondii
Spea multiplicata
Sphaenorhynchus carneus
Sphaenorhynchus dorisae
Sphaenorhynchus lacteus
Sphaenorhynchus orophilus
Sphaenorhynchus planicola
Sphenophryne cornuta
Staurois natator
Stereocyclops parkeri
Strabomantis anomalus
Strabomantis biporcatus
Strabomantis necerus
Strabomantis sulcatus
Synapturanus mirandaribeiroi
Synapturanus salseri
Syncope antenori
Syncope tridactyla
Taudactylus acutirostris
Taudactylus rheophilus
Teratohyla midas
Teratohyla pulverata
Teratohyla spinosa
26
39
40
24
32
42
25
25
46
45
35
21
26
42
30
23
35
27
43
17
20
59
56
63
78
45
50
65
59
18
18
42
30
22
41
34
40
59
60
68
35
30
28
11
12
27
27
19
29
20
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
1
0
1
0
1
0
1
0
1
1
0
1
1
0
1
1
1
1
1
1
0
1
1
0
0
0
1
1
1
1
1
1
0
1
0
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
1
1
0
0
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
1
1
1
0
1
1
0
0
1
1
0
1
1
0
0
0
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
0
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
1
1
1
1
1
1
1
1
0
0
0
0
0
LE
LE
LE
LE
LE
LE
PHY
LE
LE
LE
LE
LE
LE
LE
LE
LE
PHY
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
ENDO
LO
SUS
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
ENDO
LO
LO
LO
LO
LO
Thoropa lutzi
Thoropa miliaris
Thoropa petropolitana
Tlalocohyla loquax
Tlalocohyla picta
Tlalocohyla smithii
Trachycephalus coriaceus
Trachycephalus hadroceps
Trachycephalus jordani
Trachycephalus mesophaeus
Trachycephalus nigromaculatus
Trachycephalus resinifictrix
Trachycephalus venulosus
Triprion petasatus
Uperoleia altissima
Uperoleia capitulata
Uperoleia fusca
Uperoleia inundata
Uperoleia laevigata
Uperoleia lithomoda
Uperoleia littlejohni
Uperoleia mimula
Uperoleia rugosa
Uperoleia trachyderma
Vitreorana eurygnatha
Vitreorana oyampiensis
Vitreorana uranoscopa
Xenohyla truncata
Xenophrys minor
Xenopus laevis
Xenorhina bidens
Xenorhina oxycephala
Xenorhina rostrata
Zachaenus parvulus
28
54
24
45
21
26
60
60
76
85
86
77
101
61
25
27
28
28
28
27
32
28
32
26
23
21
26
30
38
98
29
40
39
24
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
0
0
1
0
1
1
1
1
0
0
1
0
0
0
1
0
0
0
0
0
0
0
0
0
0
1
1
1
1
0
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
1
1
1
1
1
1
1
1
1
1
0
0
0
1
1
1
1
1
1
1
1
1
1
1
0
0
0
0
0
0
0
0
0
0
1
1
1
1
0
0
0
0
0
0
1
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
1
1
LO
LO
LO
LE
LE
LE
LE
PHY
LE
LE
LE
PHY
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LE
LO
LO
LO
LE
LE
SUS
ENDO
ENDO
ENDO
ENDO
Appendix S2. List of the species included in this study and the associate literature
sources.
References
Acris
crepitans
Acris
gryllus
Adelophry
ne
adiastola
Adelotus
brevis
Adelphobat
es
galactonot
us
Adenomera
bokermann
i
Adenomera
diptyx
Adenomera
heyeri
Adenomera
hylaedactyl
a
Adenomera
marmorata
Agalychnis
litodryas
Agalychnis
moreletii
Allobates
brunneus
Allobates
femoralis
Allobates
marchesia
nus
Allobates
olfersioide
s
Allobates
sumtuosus
Allobates
trilineatus
Allophryne
ruthveni
Alytes
obstetrican
s
Ameerega
bassleri
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Ameerega
hahneli
Ameerega
parvula
Ameerega
petersi
Ameerega
picta
Ameerega
trivittata
Anaxyrus
americanu
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Anaxyrus
boreas
Anaxyrus
houstonens
is
Anaxyrus
kelloggi
Anomalogl
ossus
baeobatrac
hus
Anomalogl
ossus
degranville
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Ansonia
minuta
Ansonia
spinulifer
Aparasphe
nodon
brunoi
Aparasphe
nodon
venezolanu
s
Aplastodis
cus
albofrenat
us
Aplastodis
cus
albosignat
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Aplastodis
cus arildae
Aplastodis
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Atelopus
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Atelopus
elegans
Atelopus
flavescens
Atelopus
spumarius
Austrochap
erina
adelphe
Austrochap
erina fryi
Austrochap
erina
gracilipes
Austrochap
erina
novaebrita
nniae
Austrochap
erina
pluvialis
Austrochap
erina
robusta
Babina
adenopleur
a
Babina
okinavana
Babina
pleuraden
Barycholos
pulcher
Batrachylo
des
elegans
Batrachylo
des
mediodiscu
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des
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mystacinus
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rhodonotus
Leptodacty
lus riveroi
Leptodacty
lus savagei
Leptodacty
lus spixi
Leptodacty
lus
stenodema
Leptodacty
lus syphax
Leptodacty
lus vastus
Leptolalax
hamidi
Leptophryn
e
borbonica
Leptophryn
e cruentata
Limnodyna
stes
convexiusc
ulus
Limnodyna
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stes
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Limnodyna
stes
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Limnodyna
stes
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Limnodyna
stes
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is
Limnodyna
stes
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Limnomed
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Limnonect
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Limnonect
es kuhlii
Limnonect
es laticeps
Limnonect
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leporinus
Limnonect
es leytensis
Limnonect
es
macroceph
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Limnonect
es magnus
Limnonect
es
microdiscu
s
Limnonect
es
palavanens
is
Liophryne
schlaginha
ufeni
Lithobates
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Lithobates
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Lithobates
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Lithobates
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Lithobates
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Lithobates
clamitans
Lithobates
forreri
Lithobates
grylio
Lithobates
juliani
Lithobates
maculatus
Lithobates
magnaocul
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Lithobates
megapoda
Lithobates
neovolcani
cus
Lithobates
palmipes
Lithobates
palustris
Lithobates
pipiens
Lithobates
psilonota
Lithobates
septentrion
alis
Lithobates
sevosus
Lithobates
sylvaticus
Lithobates
tarahumar
ae
Lithobates
taylori
Lithobates
virgatipes
Lithobates
warszewits
chii
Litoria
alboguttata
Litoria
aurea
Litoria
australis
Litoria
bicolor
Litoria
brevipalma
ta
Litoria
brevipes
Litoria
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caerulea
Litoria
chloris
Litoria
congenita
Litoria
coplandi
Litoria
cryptotis
Litoria
dahlii
Litoria
dayi
Litoria
dentata
Litoria
dorsalis
Litoria
electrica
Litoria
eucnemis
Litoria
fallax
Litoria
freycineti
Litoria
genimacul
ata
Litoria
gracilenta
Litoria
impura
Litoria
inermis
Litoria
infrafrenat
a
Litoria
jungguy
Litoria
latopalmat
a
Litoria
longipes
Litoria
lutea
Litoria
maculosa
Litoria
manya
Litoria
meiriana
Litoria
microbelos
Litoria
nannotis
Litoria
nasuta
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Litoria
nigrofrenat
a
Litoria
nigropunct
ata
Litoria
novaeholla
ndiae
Litoria
nyakalensi
s
Litoria
olongburen
sis
Litoria
pallida
Litoria
pearsonian
a
Litoria
peronii
Litoria
personata
Litoria
phyllochro
a
Litoria
platycepha
la
Litoria
pygmaea
Litoria
revelata
Litoria
rheocola
Litoria
rothii
Litoria
rubella
Litoria
thesaurensi
s
Litoria
tornieri
Litoria
tyleri
Litoria
verrauxii
Litoria
vocivincen
s
Litoria
watjulume
nsis
Litoria
wilcoxii
Litoria
xanthomer
a
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Synapturan
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Synapturan
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Syncope
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Taudactylu
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Appendix 3. Taxonomic trees and functional PCO ordination plots used in this study.
Taxonomic trees: A) L. catesbeianus; B) R. marina. The black arrows indicate the
position of L. catesbeianus and R. marina.C) PCO plots.
A)
B)
C)