LG 4 Outline Evolutionary Relationships and Classification Goals of Systematics Phylogenetic Trees – Taxonomy – The Linnaean System of Taxonomy Binomial Nomenclature – Taxonomic Hierarchy – Traits Used in Classification Morphological Characteristics – Homologies – Homoplasies (Analogies) – Ancestral and Derived Characters – Cladistics – Molecular Phylogenetics – AP Biology Unit III Evolution Learning Goal 4 Describe how evolutionary relationships are used to classify organisms. Goals of Systematics • Reconstruct phylogeny (evolutionary history) of a group of organisms. This history is illustrated in phylogenetic trees. These identify likely relationships among species. • Taxonomy, or the identification and naming of species is the second goal. Taxonomy also places organisms into a classification scheme. • Naming and Classifying Organisms • Began with the Swedish naturalist Carolus Linnaeus in the 1700’s. The Linnaean System of Taxonomy • He invented the system of binomial nomenclature. In this system, each species is assigned a two-part Latinized name. • The first part identifies a group of species with similar morphology, called a genus. • The second part is the species (specific) name. • Linnaeus also developed a taxonomic hierarchy to arrange large numbers of organisms into more inclusive groups. • A family is a group of genera (plural of genus) that closely resemble one another. • Similar families are group into orders. • Similar orders are grouped into classes. • Similar classes are grouped into phyla (singular phylum). • Each category of the taxonomic hierarchy is called a taxon. • Similar phyla are grouped into kingdoms. • All life on Earth is classified into three domains. Traits Used In Classification • Morphological Characteristics Visible, measurable traits that distinguish groups of organisms from one another. Reflect genetic differences. Are preserved in the fossil record. • Homologies Similarities that result from shared ancestry. • Homoplasies Phenotypic similarities that evolved independently in different lineages. Also know as analogies. Systematists exclude homoplasies from their analyses. • Ancestral and Derived Characters Ancestral characters are old forms of traits. Derived characters are newer forms of traits. All species exhibit a mix of both types of characters and they provide information about evolutionary relationships. Cladistics • A more recent approach to classification based solely on evolutionary relationships. • Cladists group together only species that share derived characters. • Phylogenetic trees produced by cladists are called cladograms. Molecular Phylogenetics • Mutations in some types of DNA appear to arise at a relatively constant rate. Differences in the DNA sequences of two species can serve as a molecular clock. Large differences imply divergence in the distant past. Small differences suggest a more recent common ancestor. • Since DNA provides a code for the production of proteins, these molecules can be analyzed to determine possible relationships between species. • When two species exhibit similar amino acid sequences for the same protein, systematists infer their genetic similarity and evolutionary relationship. Maximum Likelihood Programs • Statistical model that constructs numerous alternative phylogenetic trees from molecular data, and estimates how likely it is that each tree represents the true evolutionary history. • Sytematists then accept the phylogenetic tree that is most likely to be true until more data are available. LG 4 Vocab 1. 2. 3. 4. 5. 6. 7. 8. 9. 10. Systematics Taxonomic Hierarchy Homoplasies Ancestral Characters Derived Characters Cladistics Cladograms Molecular Phylogenetics Molecular Clocks Analogous Structures