Invertebrates Importance • • • • • • 95-99% of all species Pollination Recycling Food in many cultures Food webs Maintenance of ecological communities E.O. Wilson “If human beings were not so impressed by size alone, they would consider an ant more wonderful than a rhinoceros.” Basics • Aquatic vs. Terrestrial – Arboreal, aerial • Anatomical Definitions – Hemolymph, exoskeleton, hydrostatic skeleton • Reproductive terms – Dioecious, hermaphroditic, parthenogenesis Ctenophora “Comb Jellies” • Marine waters • Carnivorous Diet • Water requirements dependent on species • Most are hermaphroditic Phylogeny Porifera “Sponges” • Primarily Marine; sessile • Water requirements dependent on species • Unique feeding system; tiny pores; water flow necessary • Reproduce by both sexual and asexual means Phylogeny Cnidaria Anemones Jellyfish •Radial symmetry,Hydrostatic skeleton, Dimorphic development •Can sting! Coral Anemones • Marine • Most capture animal prey; zooanthellae • Variable but specific temperature requirements dependent upon species • Asexual reproduction • Interspecific Aggression Jellyfish • • • • Marine Predators – have stingers! Sexual reproduction Lots of species variability Coral • • • • Marine Zooxanthellae; Most capture animal prey Sexual & Asexual Reproduction Can exhibit contact inhibition and interspecific aggression in the tank • Hermatypic vs Ahermatypic Phylogeny Echinoderms • • • • Marine Water quality important- toxicity problems Diet is largely dependent on species Asexual & sexual reproduction dependent on species • Water vascular system locomotion, gripping, feeding http://www.studentreader.com/files/purplestarfishonrocks.jpg Phylogeny Sea Squirts • Sessile, marine • Herbivorous diet • Sexual or Asexual reproduction • 90% of all urochordates http://www.lancashiremcs.org.uk/gallery/pics/sea-squirt.jpg Gastropods • • • • • Marine & freshwater Sessile and mobile Filter feeders Substrate Dioecious or hermaphroditic http://jrscience.wcp.muohio.edu/photos/SeagrassMolluscs.jpeg Cephalopods • • • • Marine Aquatic Varied diet Active hunters Dioecious http://www.aquariumofpacific.org/images/olc/nautilusj.baecke r_berlin_zoo_aquarium_pd600.jpg Phylogeny Pogonophora & Vestimentifera • Deep sea dwelling worms! • Often grouped together www.ucmp.berkeley.edu www.nsf.gov Phylogeny Annelids • • • • Aquatic and terrestrial Varied diet and feeding strategies Sexual reproduction; dioecious Biomedical & ecological importance http://www.education.umd.edu/blt/pic/Annelids.jpg Phylogeny Rotifers • Most freshwater but some marine and terrestrial • Omnivores • Parthenogenic or dioecious Phylogeny Flatworms (Turbellarians) • • • • Mostly Aquatic Varied diet Commensal and parasitic Varied reproduction http://www.dhadm.com/wp-content/uploads/2008/03/flatworm02.jpg Nematodes • • • • Aquatic & Marine Varied diet Mostly parasitic Dieocious or parthenogenic http://soils.usda.gov/sqi/concepts/soil_biology/images/large_todes_LR.jpg Phylogeny Tardigrades • Marine, freshwater & terrestrial • “Water bears” • Cryptobiosis http://www.uea.ac.uk/~b444219/images/TNWP_Echiniscus%20madonnae%20(SEM).jpg Phylogeny Onychophora • Tropical, terrestrial • Velvet worms or walking worms • Prey on smaller arthropods Phylogeny Crustaceans • • • • Terrestrial and Aquatic Diet dependent on species Dieocious Economic importance www.britannica.com/eb/art/print?id=104965 Phylogeny Insects • • • • • Predominantly terrestrial Varied Diet Reproduction Social Systems Economical & Ecological Importance Phylogeny Myriapods • Terrestrial • Diet dependent on species • Well defined environmental requirements • Dieocious; some parthenogenic Myriapods • • • • Millipedes Mostly herbivorous Two pairs per body segment Non-aggressive; slow Easy to handle • • • • Centipedes Mostly carnivorous One pair of legs per body segment Aggressive; fast! Extremely hard to handle; venomous http://cordially.narod.ru/album/insect/images/home-centipede.jpg http://www.garden-city.org/zoo/animalinfo/images/milipede_76pic.jpg Phylogeny Horseshoe Crabs • • • • • Marine, Aquatic Captive diet Various habitats Dieocious Limulus amebocyte lysate (LAL) extracted from hemolymph- Pharmaceutical use http://io.uwinnipeg.ca/~simmons/16cm05/1116/33-28-HorseshoeCrabs.jpg Phylogeny SPIDERS! Arachnids Scorpions • Terrestrial • Carnivorous • Environment: mostly desert • Sexual reproduction Spiders • • • • • Aquatic & terrestrial Carnivores Diverse Environments Sexual Reproduction Silk! Terrestrial Spiders • Arboreal vs. Non-arboreal Aquatic Spiders • Underwater “air bells” Silk • • • • • • Made of Amino Acids Very stable Very Strong Genetic Isolation Production Application