Chapter 3 Ecosystems: What Are They and How Do They Work? Chapter Overview Questions What is ecology? What basic processes keep us and other organisms alive? What are the major components of an ecosystem? What happens to energy in an ecosystem? What are soils and how are they formed? What happens to matter in an ecosystem? How do scientists study ecosystems? Core Case Study: Have You Thanked the Insects Today? Many plant species depend on insects for pollination. Insect can control other pest insects by eating them Figure 3-1 Core Case Study: Have You Thanked the Insects Today? …if all insects disappeared, humanity probably could not last more than a few months [E.O. Wilson, Biodiversity expert]. Insect’s role in nature is part of the larger biological community in which they live. Universe Galaxies Solar systems Biosphere Planets Earth Biosphere Ecosystems Ecosystems Communities Populations Realm of ecology Organisms Organ systems Communities Organs Tissues Cells Populations Protoplasm Molecules Atoms Organisms Subatomic Particles Fig. 3-2, p. 51 Organisms and Species Organisms, the different forms of life on earth, can be classified into different species based on certain characteristics. Figure 3-3 Case Study: Which Species Run the World? Multitudes of tiny microbes such as bacteria, protozoa, fungi, and yeast help keep us alive. Harmful microbes are the minority. Soil bacteria convert nitrogen gas to a usable form for plants. They help produce foods (bread, cheese, yogurt, beer, wine). 90% of all living mass. Helps purify water, provide oxygen, breakdown waste. Lives beneficially in your body (intestines, nose). Biosphere Atmosphere Membrane of air around the planet. Stratosphere Lower portion contains ozone to filter out most of the sun’s harmful UV radiation. Hydrosphere All the earth’s water: liquid, ice, water vapor Lithosphere The earth’s crust and upper mantle. What Sustains Life on Earth? Solar energy, the cycling of matter, and gravity sustain the earth’s life. Figure 3-7 Solar radiation Energy in = Energy out Reflected by atmosphere (34% ) UV radiation Absorbed by ozone Visible Light Absorbed by the earth Radiated by atmosphere as heat (66%) Lower Stratosphere (ozone layer) Troposphere Greenhouse effect Heat Heat radiated by the earth Fig. 3-8, p. 55 Oxygen (O2) Sun Producer Carbon dioxide (CO2) Secondary consumer Primary (fox) consumer (rabbit) Precipitation Falling leaves and twigs Producers Soil decomposers Water Fig. 3-10, p. 57 Factors That Limit Population Growth Availability of matter and energy resources can limit the number of organisms in a population. Figure 3-11 Abundance of organisms Upper limit of tolerance Few No organisms organisms Population size Lower limit of tolerance No Few organisms organisms Zone of intolerance Low Zone of physiological stress Optimum range Temperature Zone of physiological stress Zone of intolerance High Fig. 3-11, p. 58 Factors That Limit Population Growth The physical conditions of the environment can limit the distribution of a species. Figure 3-12 Producers: Basic Source of All Food Most producers capture sunlight to produce carbohydrates by photosynthesis: Photosynthesis: A Closer Look Chlorophyll molecules in the chloroplasts of plant cells absorb solar energy. This initiates a complex series of chemical reactions in which carbon dioxide and water are converted to sugars and oxygen. Figure 3-A Consumers: Eating and Recycling to Survive Consumers (heterotrophs) get their food by eating or breaking down all or parts of other organisms or their remains. Herbivores • Primary consumers that eat producers Carnivores • Primary consumers eat primary consumers • Third and higher level consumers: carnivores that eat carnivores. Omnivores • Feed on both plant and animals. Decomposers and Detrivores Decomposers: Recycle nutrients in ecosystems. Detrivores: Insects or other scavengers that feed on wastes or dead bodies. Figure 3-13 Aerobic and Anaerobic Respiration: Getting Energy for Survival Organisms break down carbohydrates and other organic compounds in their cells to obtain the energy they need. This is usually done through aerobic respiration. The opposite of photosynthesis Aerobic and Anaerobic Respiration: Getting Energy for Survival Anaerobic respiration or fermentation: Some decomposers get energy by breaking down glucose (or other organic compounds) in the absence of oxygen. The end products vary based on the chemical reaction: • • • • Methane gas Ethyl alcohol Acetic acid Hydrogen sulfide Two Secrets of Survival: Energy Flow and Matter Recycle An ecosystem survives by a combination of energy flow and matter recycling. Figure 3-14 BIODIVERSITY Figure 3-15 Biodiversity Loss and Species Extinction: Remember HIPPO H for habitat destruction and degradation I for invasive species P for pollution P for human population growth O for overexploitation Why Should We Care About Biodiversity? Biodiversity provides us with: Natural Resources (food water, wood, energy, and medicines) Natural Services (air and water purification, soil fertility, waste disposal, pest control) Aesthetic pleasure The Ecosystem Approach The Species Approach Goal Goal Protect populations of species in their natural habitats Protect species from premature extinction Strategy Preserve sufficient areas of habitats in different biomes and aquatic systems Strategies Tactics •Protect habitat areas through private purchase or government action •Eliminate or reduce populations of nonnative species from protected areas •Manage protected areas to sustain native species •Restore degraded ecosystems •Identify endangered species •Protect their critical habitats Tactics •Legally protect endangered species •Manage habitat •Propagate endangered species in captivity •Reintroduce species into suitable habitats Fig. 3-16, p. 63 ENERGY FLOW IN ECOSYSTEMS Food chains and webs show how eaters, the eaten, and the decomposed are connected to one another in an ecosystem. Figure 3-17 Food Webs Trophic levels are interconnected within a more complicated food web. Figure 3-18 Energy Flow in an Ecosystem: Losing Energy in Food Chains and Webs accordance with the 2nd law of thermodynamics, there is a decrease in the amount of energy available to each succeeding organism in a food chain or web. In Energy Flow in an Ecosystem: Losing Energy in Food Chains and Webs Ecological efficiency: percentage of useable energy transferred as biomass from one trophic level to the next. Figure 3-19 Productivity of Producers: The Rate Is Crucial Gross primary production (GPP) Rate at which an ecosystem’s producers convert solar energy into chemical energy as biomass. Figure 3-20 Net Primary Production (NPP) NPP = GPP – R Rate at which producers use photosynthesis to store energy minus the rate at which they use some of this energy through respiration (R). Figure 3-21 What are nature’s three most productive and three least productive systems? Figure 3-22 SOIL: A RENEWABLE RESOURCE Soil is a slowly renewed resource that provides most of the nutrients needed for plant growth and also helps purify water. Soil formation begins when bedrock is broken down by physical, chemical and biological processes called weathering. Mature soils, or soils that have developed over a long time are arranged in a series of horizontal layers called soil horizons. Oak tree Wood sorrel Lords and ladies Fern O horizon Leaf litter Dog violet Grasses and small shrubs Earthworm Millipede Honey fungus Mole Organic debris builds up Rock fragments Moss and lichen A horizon Topsoil B horizon Subsoil Bedrock Immature soil Regolith Young soil Pseudoscorpion C horizon Mite Parent material Nematode Root system Mature soil Red Earth Mite Springtail Actinomycetes Fungus Bacteria Fig. 3-23, p. 68 Layers in Mature Soils Infiltration: the downward movement of water through soil. Leaching: dissolving of minerals and organic matter in upper layers carrying them to lower layers. The soil type determines the degree of infiltration and leaching. Soil Profiles of the Principal Terrestrial Soil Types Figure 3-24 Some Soil Properties Soils vary in the size of the particles they contain, the amount of space between these particles, and how rapidly water flows through them. Figure 3-25 MATTER CYCLING IN ECOSYSTEMS Nutrient Cycles: Global Recycling Global Cycles recycle nutrients through the earth’s air, land, water, and living organisms. Nutrients are the elements and compounds that organisms need to live, grow, and reproduce. Biogeochemical cycles move these substances through air, water, soil, rock and living organisms. Condensation Rain clouds Transpiration Evaporation Transpiration Precipitation to land from plants Precipitation Runoff Surface runoff (rapid) Precipitation Evaporation from land Evaporation from ocean Precipitation to ocean Surface runoff (rapid) Infiltration and Percolation Groundwater movement (slow) Ocean storage Fig. 3-26, p. 72 Effects of Human Activities on Water Cycle We alter the water cycle by: Withdrawing large amounts of freshwater. Clearing vegetation and eroding soils. Polluting surface and underground water. Contributing to climate change. Fig. 3-27, pp. 72-73 Effects of Human Activities on Carbon Cycle We alter the carbon cycle by adding excess CO2 to the atmosphere through: Burning fossil fuels. Clearing vegetation faster than it is replaced. Figure 3-28 Gaseous nitrogen (N2) in atmosphere Food webs on land Nitrogen fixation Fertilizers Uptake by autotrophs Excretion, death, decomposition Ammonia, ammonium in soil Nitrogen-rich wastes, remains in soil Ammonification Loss by leaching Nitrification Uptake by Loss by autotrophs denitrification Nitrate in soil Nitrification Nitrite in soil Loss by leaching Fig. 3-29, p. 75 Effects of Human Activities on the Nitrogen Cycle We alter the nitrogen cycle by: Adding gases that contribute to acid rain. Adding nitrous oxide to the atmosphere through farming practices which can warm the atmosphere and deplete ozone. Contaminating ground water from nitrate ions in inorganic fertilizers. Releasing nitrogen into the troposphere through deforestation. Effects of Human Activities on the Nitrogen Cycle Human activities such as production of fertilizers now fix more nitrogen than all natural sources combined. Figure 3-30 mining excretion Fertilizer Guano agriculture uptake by uptake by weathering autotrophs autotrophs leaching, runoff Dissolved Land Marine Dissolved in Soil Water, Food Food in Ocean Lakes, Rivers Webs Webs Water death, death, decomposition decomposition weathering sedimentation settling out uplifting over geologic time Rocks Marine Sediments Fig. 3-31, p. 77 Effects of Human Activities on the Phosphorous Cycle We remove large amounts of phosphate from the earth to make fertilizer. We reduce phosphorous in tropical soils by clearing forests. We add excess phosphates to aquatic systems from runoff of animal wastes and fertilizers. Sulfur trioxide Water Acidic fog and precipitation Sulfuric acid Ammonia Oxygen Sulfur dioxide Ammonium sulfate Hydrogen sulfide Plants Dimethyl sulfide Volcano Industries Animals Ocean Sulfate salts Metallic sulfide deposits Decaying matter Sulfur Hydrogen sulfide Fig. 3-32, p. 78 Effects of Human Activities on the Sulfur Cycle We add sulfur dioxide to the atmosphere by: Burning coal and oil Refining sulfur containing petroleum. Convert sulfur-containing metallic ores into free metals such as copper, lead, and zinc releasing sulfur dioxide into the environment. Geographic Information Systems (GIS) A GIS organizes, stores, and analyzes complex data collected over broad geographic areas. Allows the simultaneous overlay of many layers of data. Figure 3-33 Critical nesting site locations USDA Forest Service USDA Private Forest Service owner 1 Private owner 2 Topography Forest Habitat type Wetland Lake Grassland Real world Fig. 3-33, p. 79 Systems Analysis Ecologists develop mathematical and other models to simulate the behavior of ecosystems. Figure 3-34 Systems Measurement Define objectives Identify and inventory variables Obtain baseline data on variables Data Analysis Make statistical analysis of relationships among variables Determine significant interactions System Modeling Objectives Construct mathematical model describing interactions among variables System Simulation System Optimization Run the model on a computer, with values entered for different Variables Evaluate best ways to achieve objectives Fig. 3-34, p. 80 Importance of Baseline Ecological Data We need baseline data on the world’s ecosystems so we can see how they are changing and develop effective strategies for preventing or slowing their degradation. Scientists have less than half of the basic ecological data needed to evaluate the status of ecosystems in the United Sates (Heinz Foundation 2002; Millennium Assessment 2005).