Origin of Species Chapter 24 1 What you need to know! • The difference between microevolution and macroevolution. • The biological concept of species. • Prezygotic and postzygotic barriers that maintain reproductive isolation in natural populations. • How allopatric and sympatric speciation are similar and different. • How an autopolyploid or an allopolyploid chromosomal change can lead to sympatric speciation. • How punctuated equilibrium and gradualism describe two different tempos of speciation. 2 Species • A population (or group of populations) who can mate and produce fertile offspring • Speciation: One species transforms into another • A Population transforms over time as adaptations add up in its gene pool (microevolution) • Macroevolution: is major developmental changes spawning new branches of species – the evolution of the spinal cord: vertebrates – the evolution of feathers: birds 3 Adaptations • A genetic change that makes an organism better suited to their environment • Natural selection facilitates adaptation Special adaptations: • Mimicry: mimicking another organism that is potentially dangerous • Camouflage: blending in with the environment 4 Reproductive Isolation Prezygotic (prevent mating and/or hinder fertilization): 1. Habitat isolation: don’t share a habitat 2. Behavioral isolation: different mating practices 3. Temporal isolation: sex at different times of day or year 4. Mechanical isolation: not physically possible 5. Gametic isolation: sperm cannot fertilize 5 egg Reproductive Isolation Postzygotic (fertilized eggs don’t develop into fertile adults): 1. Reduced hybrid viability: genetic incompatibility terminates pregnancy 2. Reduced hybrid fertility: viable offspring that cannot reproduce (mule) 3. Hybrid breakdown: viable offspring that grow and reproduce but their 6 offspring are weak or sterile Speciation Allopatric Speciation • random mating becomes impossible due to geographic isolation (rivers, canyons, mountains) • New species evolve on the fringes of large ancestral populations Sympatric Speciation • Sudden mutation within a population leads to new species • Polyploidy plants mutate by altering chromosome number in a non-disjunction 7 error Adaptive Radiation • Colonization of new habitat – New habitats usually vary from the original habitat • Example Darwin’s Finches 8 Rate of Change • Species that change slowly are K (carrying capacity) – selected – elephants • Species that change quickly are r (growth rate) – selected – Bacteria • Gradualism: species descended from a common ancestor and gradually diverge • Punctuated equilibrium: period of apparent stasis followed by rapid change 9