Recent and otherwise important publications
Peer-review high-impact journals
1. Tarkhnishvili D., Murtskhvaladze M., Gavashelishvili A. 2013. Speciation in Caucasian lizards: climatic
dissimilarity of the habitats is more important than isolation time. Biological Journal of the Linnean Society (in
press).
2. Mumladze L., Tarkhnishvili D., Murtskhvaladze M. 2013. Systematics and evolutionary history of large
endemic snails from the Caucasus (Helix buchii, H. goderdziana). American Malacological Bulletin 31 (2) (in
press).
3. Tarkhnishvili, D. Gavashelishvili, A. Mumladze, L. 2012. Palaeoclimatic models help to understand current
distribution of Caucasian forest species. Biological Journal of the Linnean Society. 105, 231–248.
4. Murtskhvaladze, M. Gurielidze, Z. Kopaliani, N. Tarkhnishvili, D. 2012. Gene introgression between Gazella
subgutturosa and G. marica: limitations of maternal inheritance analysis for species identification with
conservation purposes. Acta Theriologica 57, 383-386
5. Pokryszko, B.M. Cameron, R.A.D. Mumladze, L. Tarkhnishvili, D. 2011. Forest snail faunas from Georgian
Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society 102,
239-250
6. Murtskhvaladze, M. Gavashelishvili, A. Tarkhnishvili, D. 2010. Geographic and genetic boundaries of brown
bear (Ursus arctos) population in the Caucasus. Molecular Ecology 19, 1829-1841.
7. Tarkhnishvili, D. Gavashelishvili, A. Avaliani, A. Murtskhvaladze, M. Mumladze, L. 2010. Unisexual rock lizard
might be outcompeting its bisexual progenitors in the Caucasus. Biological Journal of the Linnean Society 101,
447-460
8. Tarkhnishvili, D. Serbinova, I. Gavashelishvili, A. 2009. Modelling the range of Syrian spadefoot toad
(Pelobates syriacus) with combination of GIS-based approaches. Amphibia-Reptilia. 30, 401-412
9. SayIm, F. Baskale, E. Tarkhnishvili, D. Kaya, U. 2009. Some water chemistry parameters of breeding habitats
of the Caucasian salamander, Mertensiella caucasica in the Western Lesser Caucasus. Comptes Rendus
Biologies 332, 464-469.
10. Tarkhnishvili, D. Kaya, U. Gavashelishvili, A. Serbinova, I. 2008. Ecological divergence between two
evolutionary lineages of the Caucasian salamander: evidence from GIS analysis. The Herpetological Journal 18,
155-163.
11. Mumladze, L. Tarkhnishvili, D. Pokryszko, BM 2008. A new species of the genus Helix from the Lesser
Caucasus (SW Georgia). Journal of Conchology 39, 483-486.
12. Weddeling, K. Hachtel, M. Sander, U. Tarkhnishvili, D. 2004 Bias in estimation of newt population size: a
field study at five ponds using drift fences, pitfalls and funnel traps. The Herpetological Journal 14 1-8
13. Hille, A. Tarkhnishvili, D. Meinig, H. Hutterer, R. 2002. Morphometric, biochemical and molecular traits in
Caucasian wood mice (Apodemus/Sylvaemus), with remarks on species divergence. Acta theriologica 47, 389416.
14. Tarkhnishvili D, Hille A, Boehme W. 2001. Humid forest refugia, speciation and secondary introgression
between evolutionary lineages: differentiation in a Near Eastern brown frog, Rana macrocnemis. Biological
Journal of the Linnean Society 74 141-156.
15. Tarkhnishvili D, Thorpe R, Arntzen J.W. (2000) Pre-Pleistocene refugia and differentiation between
populations of the Caucasian salamander (Mertensiella caucasica). Molecular Phylogenetics and Evolution 14
414-422.
Books, book chapters
16. Tarkhnishvili, D. 2013 (in press). Rana macrocnemis (Boulenger, 1885) – Near Eastern Brown Frogs. In:
Grossenbacher, K. (ed): Handbuch der Reptilien und Amphibien Europas.
17. Tarkhnishvili, D. 2012. Evolutionary History, Habitats, Diversification, and Speciation in Caucasian Rock
Lizards. Advances in Zoology Research. Owen P. Jenkins, Ed. 2, 79-120
18. Tarkhnishvili, D. Kaya, U. 2009 Status and Conservation of the Caucasian Salamander (Mertensiella
caucasica) Zazanashvuli, N. and Mallon, D.(Eds.) Status and Protection of Globally Threatened Species in the
Caucasus 157-164
19. Sinsch, U., Schneider, H. & Tarkhnishvili, D. 2008. Bufo bufo Superspezies – Erdkröten-Artenkreis. In:
Grossenbacher, K. (ed): Handbuch der Reptilien und Amphibien Europas.
20. Hachtel, M. Weddeling, K. Schmidt, P. Sander, U. Tarkhnishvili, D. Böhme, W. 2006. Dynamik und Struktur
von Amphibienpopulationen in der Zivilisationslandschaft. Naturschutz und Biologische Vielfalt 30 420.
21. Zazanashvili, N. Sanadiradze, G. Bukhnikashvili, A. Kandaurov, A. Tarkhnishvili, D. 2004 Caucasus. In:
Hotspots Revisited: Earth’s biologically richest and most endangered terrestrial ecoregions, by R.A.
Mittermaier, P.G. Gil, M. Hoffmann, J. Pilgrim, T. Brooks, C. G. Mittermaier, J. Lamoreux, G. A. B. da Fonseca.
Pages 148-153. CEMEX/Agrupacion Sierra Madre, Mexico City.
22. Tarkhnishvili D., Gokhelashvili R. 1999. The Amphibians of the Caucasus. Sofia, Pensoft.
Other publications (last 10 years)
23. Chubinishvili, AT Gokhelashvili, RK Tarkhnishvili, DN 2011. Population ecology of the caucasian parsley frog
(Pelodytes caucasicus Boulenger) in the Borjomi Canyon. Russian Journal of Herpetology 2 2 79-86
24. Arntzen, J.W. Kuzmin, S. Jehle, R. Beebee, T. Tarkhnishvili, D. Ishchenko, V. Ananjeva, N. Orlov, N. Tuniyev,
B. Denoël, M. 2006 Triturus cristatus, 2006. www.redlist.org
25. Ortmann, D. Hachtel, M. Sander, U. Schmidt, P. Tarkhnishvili, D.N. Weddeling, K. Böhme, W. 2006 Capture
effectiveness of terrestrial drift fences and funnel traps for the Great Crested Newt, Triturus cristatus
Proceedings of the 13th Congress of the Societas Europaea Herpetologica. pp 103 105
26. Schmidt, P. Weddeling, K. Thomas, M. Rottscheidt, R. Tarkhnishvili, D.N. Hachtel, M. 2006 Dispersal of
Triturus alpestris and T. vulgaris in agricultural landscapes–comparing estimates from allozyme markers and
capture-mark-recapture analysis Proceedings of the 13th Congress of the Societas Europaea Herpetologica. pp
139 143
27. Murtskhvaladze M, Tarkhinshvili, D. 2006. Estimation of Brown Bear abundance and population structure
in Borjom-Kharagauli National Park with molecular-genetic methods. Proceedings of the Georgian Academy of
Sciences, Biological series B 4 43-50
28. Weddeling, K. Bosbach, G. Hachtel, M. Sander, U. Schmidt, P. Tarkhnishvili, D. 2005 Egg size versus clutch
size: variation and trade-offs in reproductive output of Rana dalmatina and R. temporaria in a pond near Bonn
(Germany) Herpetologia Petropolitana. Proc. of the 12th Ordinary General Meeting of the Societas Europaea
Herpetologica, August 12 16 2003
29. Hachtel, M. Schmidt, P. Sander, U. Tarkhnishvili, D. Weddeling, K. Böhme, W. 2005 Eleven years of
monitoring amphibian populations in an agricultural landscape near Bonn (Germany) Herpetologia
Petropolitana, Ananjeva N. and Tsinenko O.(eds.) 150-152
30. Tarkhnishvili, D. Kandaurov, A. Bukhnikashvili, A. 2002 Declines of amphibians and reptiles in Georgia
during the 20th century: virtual vs. actual problems Zeitschrift für Feldherpetologie 9 89-107